:Chipmunk

Chipmunks are classified as four genera: Tamias, of which the eastern chipmunk (T. striatus) is the only living member; Eutamias, of which the Siberian chipmunk (E. sibiricus) is the only living member; Nototamias, which consists of three extinct species, and Neotamias, which includes the 23 remaining, mostly western North American, species. These classifications were treated as subgenera due to the chipmunks' morphological similarities.{{cite journal |journal=Mammalia |year=2016 |volume=80 |issue=3 |pages=241–251 |title=Towards a uniform nomenclature for ground squirrels: the status of the Holarctic chipmunks |last1=Patterson|first1=Bruce D. |last2=Norris|first2=Ryan W. |doi=10.1515/mammalia-2015-0004 |s2cid=9955150 |url=https://www.webpages.uidaho.edu/~jacks/PattersonNorris.16.pdf |archive-url=https://web.archive.org/web/20190608182301/https://www.webpages.uidaho.edu/~jacks/PattersonNorris.16.pdf |archive-date=2019-06-08 |url-status=live |access-date=2019-06-08}} As a result, most taxonomies over the twentieth century have placed the chipmunks into a single genus. Joseph C. Moore reclassified chipmunks to form a subtribe Tamiina in a 1959 study, and this classification has been supported by studies of mitochondrial DNA.{{Cite journal |last=Moore |first=Joseph C. |date=1959-01-01 |title=Relationships among the living squirrels of the Sciurinae |journal=Bulletin of the American Museum of Natural History |publisher=American Museum of Natural History |volume=118}}{{cite web |author=Wilson, D. E. |author2=D. M. Reeder |year=2005 |title=Mammal Species of the World |url=http://nmnhgoph.si.edu/msw/ |archive-url=https://web.archive.org/web/20070623030727/http://nmnhgoph.si.edu/msw/ |archive-date=2007-06-23 |access-date=2007-06-27}}{{cite journal|last1=Piaggio|first1= A. J. |last2= Spicer|first2= G. S.|year= 2001|title=Molecular phylogeny of the chipmunks inferred from mitochondrial cytochrome b and cytochrome oxidase II gene sequences|journal=Molecular Phylogenetics and Evolution|volume= 20|issue= 3 |pages= 335–350|url=http://online.sfsu.edu/~gs/spicer/pages/spicerpdf/piaggio01.pdf |archive-url=https://web.archive.org/web/20120823153636/http://online.sfsu.edu/~gs/spicer/pages/spicerpdf/piaggio01.pdf |archive-date=2012-08-23 |url-status=live|doi=10.1006/mpev.2001.0975|pmid= 11527462 |bibcode= 2001MolPE..20..335P |citeseerx= 10.1.1.330.9046 }}{{cite journal|journal=Journal of Mammalian Evolution|volume= 7|issue=3|year=2000|title=Molecular Phylogeny of the Chipmunk Genus Tamias Based on the Mitochondrial Cytochrome Oxidase Subunit II Gene|first1=Antoinette J.|last1=Piaggio|first2=Greg S. |last2=Spicer|url=http://userwww.sfsu.edu/~gs/spicer/pages/spicerpdf/piaggio00.pdf |archive-url=https://web.archive.org/web/20110913140312/http://userwww.sfsu.edu/~gs/spicer/pages/spicerpdf/piaggio00.pdf |archive-date=2011-09-13 |url-status=live|doi=10.1023/a:1009484302799|pages=147–166|s2cid= 7623018}}{{cite journal|last1=Musser|first1= G. G.|last2=Durden|first2= L. A.|last3= Holden|first3= M. E.|last4= Light|first4= J. E.|year=2010|title=Systematic review of endemic Sulawesi squirrels (Rodentia, Sciuridae), with descriptions of new species of associated sucking lice (Insecta, Anoplura), and phylogenetic and zoogeographic assessments of sciurid lice|journal=Bulletin of the American Museum of Natural History|volume= 339|issue= 339|pages= 1–260|hdl= 2246/6067|doi= 10.1206/695.1|s2cid= 82712592|url= http://digitallibrary.amnh.org/bitstream/2246/6067/1/B339.pdf |archive-url=https://web.archive.org/web/20180430025925/http://digitallibrary.amnh.org/bitstream/2246/6067/1/B339.pdf |archive-date=2018-04-30 |url-status=live}}

The common name originally may have been spelled "chitmunk", from the native Odawa (Ottawa) word jidmoonh, meaning "red squirrel" (cf. Ojibwe {{lang|oj|ᐊᒋᑕᒨ}} ajidamoo).{{cite web|url=http://etymonline.com/index.php?term=chipmunk |title=Online Etymology Dictionary |publisher=Etymonline.com |access-date=7 December 2012}}Nichols, John D. and Earl Nyholm (1995). A Concise Dictionary of Minnesota Ojibwe. Minneapolis: University of Minnesota Press. The earliest form cited in the Oxford English Dictionary is "chipmonk", from 1842. Other early forms include "chipmuck" and "chipminck", and in the 1830s they were also referred to as "chip squirrels", probably in reference to the sound they make. In the mid-19th century, John James Audubon and his sons included a lithograph of the chipmunk in their Viviparous Quadrupeds of North America, calling it the "chipping squirrel [or] hackee".{{Cite book | last1 = Audubon | first1 = John James | last2 = Bachman | first2 = John | title = Imperial Collection of Audubon Mammals | publisher = Bonanza Books, a division of Crown Publishing Group | year = 1967 | location = New York | page = 52}} Chipmunks have also been referred to as "ground squirrels"{{Cite EB1911|wstitle=Ground-squirrel|volume=12|page=626}} (although the name "ground squirrel" may refer to other squirrels, such as those of the genus Spermophilus).{{cite journal | author1=Kryštufek, B. | author2=B. Vohralík | year=2013 | title=Taxonomic revision of the Palaearctic rodents (Rodentia). Part 2. Sciuridae: Urocitellus, Marmota and Sciurotamias | journal=Lynx, N. S. (Praha) | volume=44 | pages=22 }}

File:Tamia striatus eating.jpg

Chipmunks have an omnivorous diet primarily consisting of seeds, nuts and other fruits, and buds. They also commonly eat grass, shoots, and many other forms of plant matter, as well as fungi, insects and other arthropods, small frogs, worms, and bird eggs. They will also occasionally eat newly hatched baby birds.{{Unbulleted list citebundle|{{cite web|url=http://www.nhptv.org/natureworks/chipmunk.htm |title=Eastern Chipmunk - Tamias striatus - NatureWorks |publisher=Nhptv.org |access-date=2012-12-07}}|{{cite iucn |author=Cassola, F. |year=2016 |errata=2017 |title=Neotamias minimus |page=e.T42572A115190804 |doi=10.2305/IUCN.UK.2016-3.RLTS.T42572A22267269.en}}|{{cite iucn |author=Tsytsulina, K. |author2=Formozov, N. |author3=Shar, S. |author4=Lkhagvasuren, D. |author5=Sheftel, B. |year=2016 |errata=2017 |title=Eutamias sibiricus |page=e.T21360A115161465 |doi=10.2305/IUCN.UK.2016-3.RLTS.T21360A22268598.en}}|{{cite iucn |author=Cassola, F. |year=2016 |errata=2017 |title=Neotamias townsendii |page=e.T42584A115191888 |doi=10.2305/IUCN.UK.2016-3.RLTS.T42584A22268822.en}}}} Around humans, chipmunks can eat cultivated grains and vegetables, and other plants from farms and gardens, so they are sometimes considered pests.{{cite web |url=http://www.massaudubon.org/Nature_Connection/wildlife/index.php?subject=Mammals&id=22 |title=Chipmunks | Living With Wildlife |publisher=Mass Audubon |access-date=2012-12-07 |archive-url=https://web.archive.org/web/20121218031247/http://www.massaudubon.org/Nature_Connection/wildlife/index.php?subject=Mammals&id=22 |archive-date=2012-12-18 |url-status=dead }} Chipmunks mostly forage on the ground, but they climb trees to obtain nuts such as hazelnuts and acorns.{{cite web|url=http://www.animalcorner.co.uk/wildlife/chipmunk.html |title=Chipmunk at Animal Corner |publisher=Animalcorner.co.uk |date=2004-01-01 |access-date=2012-12-07}} At the beginning of autumn, many species of chipmunk begin to stockpile nonperishable foods for winter. They mostly cache their foods in a larder in their burrows and remain in their nests until spring, unlike some other species which make multiple small caches of food. Cheek pouches allow chipmunks to carry food items to their burrows for either storage or consumption.{{cite web|url=http://www.wvdnr.gov/Wildlife/Magazine/Archive/04Fall/Eastern_Chipmunk.shtm |title=West Virginia Wildlife Magazine: Wildlife Diversity Notebook. Eastern chipmunk |publisher=Wvdnr.gov |access-date=2012-12-07}}

File:Chimps2.theora.ogv]]

File:Chipmunk-burrow (exits).jpg

Eastern chipmunks, the largest of the chipmunks,{{Cite web|title=National Geographic|website=National Geographic Society |date=11 April 2010 |url=https://www.nationalgeographic.com/animals/mammals/facts/chipmunks|url-status=dead|archive-url=https://web.archive.org/web/20210301043528/https://www.nationalgeographic.com/animals/mammals/facts/chipmunks |archive-date=2021-03-01 }} mate in early spring and again in early summer, producing litters of four or five young twice each year.{{cite book | last = Hazard | first = Evan B. | title = The Mammals of Minnesota | publisher = University of Minnesota Press | year = 1982 | pages = 52–54 | url = https://books.google.com/books?id=sjoQK1bedB0C&q=eastern+chipmunk+mate&pg=PA53 | isbn = 978-0-8166-0952-9}} Western chipmunks breed only once a year. The young emerge from the burrow after about six weeks and strike out on their own within the next two weeks.{{cite book | last = Schwartz | first = Charles Walsh |author2=Elizabeth Reeder Schwartz |author3=Jerry J. Conley | title = The Wild Mammals of Missouri | publisher = University of Missouri Press | year = 2001 | pages = 135–140 | url = https://books.google.com/books?id=uEWl0ZM6DfUC&q=eastern+chipmunk+young&pg=PA140 | isbn = 978-0-8262-1359-4}}

These small mammals fulfill several important functions in forest ecosystems. Their activities harvesting and hoarding tree seeds play a crucial role in seedling establishment. They consume many different kinds of fungi, including those involved in symbiotic mycorrhizal associations with trees, and are a vector for dispersal of the spores of subterranean sporocarps (truffles) in some regions.{{Cite journal |last1=Pyare |first1=Sanjay |last2=Longland |first2=William S. |date=2001 |title=Patterns of Ectomycorrhizal-Fungi Consumption by Small Mammals in Remnant Old-Growth Forests of the Sierra Nevada |url=https://www.jstor.org/stable/1383605 |journal=Journal of Mammalogy |volume=82 |issue=3 |pages=681–689 |doi=10.1644/1545-1542(2001)082<0681:POEFCB>2.0.CO;2 |jstor=1383605 |issn=0022-2372}}

Chipmunks construct extensive burrows which can be more than {{convert|3.5|m|ft|abbr=on}} in length with several well-concealed entrances.{{cite web | url=http://www.esf.edu/aec/adks/mammals/chipmunk.htm | title=Eastern Chipmunk | publisher=Adirondack Ecological Center | work=Adirondack Mammals | date=1988 | access-date=2015-09-19 | author=Saunders, D. A.}} The sleeping quarters are kept clear of shells, and feces are stored in refuse tunnels.{{cite book | url=https://books.google.com/books?id=D9fWfWLmMcIC&q=chipmunk+refuse+tunnel&pg=PA250 | title=Field Guide to the Natural World of New York City | publisher=Johns Hopkins University Press | isbn=9780801886829 | date=October 2007 | access-date=2015-09-19 | author=Leslie Day}}

The eastern chipmunk hibernates in the winter, while western chipmunks do not, relying on the stores in their burrows.{{cite book|last1=Kays|first1=R. W.|first2=Don E.|last2=Wilson|year=2009|publisher=Princeton University Press|title=Mammals of North America|edition=2nd|isbn=978-0-691-14092-6|page=72}}

Chipmunks play an important role as prey for various predatory mammals and birds but are also opportunistic predators themselves, particularly with regard to bird eggs and nestlings, as in the case of eastern chipmunks and mountain bluebirds (Siala currucoides).{{cite web|last1=Sullivan|first1=Janet|title=WILDLIFE SPECIES: Sialia currucoides|url=https://www.fs.fed.us/database/feis/animals/bird/sicu/all.html|website=Fire Effects Information System, [Online]|publisher=U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory|access-date=23 January 2016}}

Chipmunks typically live about three years, although some have been observed living to nine years in captivity.{{cite web |url=http://www.essortment.com/information-chipmunks-56048.html |title=Information on Chipmunks |publisher=Essortment.com |date=1986-05-16 |access-date=2012-12-07 |archive-date=2012-12-22 |archive-url=https://web.archive.org/web/20121222102625/http://www.essortment.com/information-chipmunks-56048.html |url-status=dead }}

Chipmunks are diurnal. In captivity, they are said to sleep for an average of about 15 hours a day. It is thought that mammals which can sleep in hiding, such as rodents and bats, tend to sleep longer than those that must remain on alert.Jennifer S. Holland "40 Winks?" (July 2011). National Geographic Vol. 220, No. 1.

Genus Eutamias

• Siberian chipmunk, Eutamias sibiricus

Genus Tamias[https://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=12401131 Tamias], Mammal Species of the World, 3rd ed.

• Eastern chipmunk, Tamias striatus
• Tamias aristus †

Genus Neotamias

{{Div col}}

• Allen's chipmunk, Neotamias senex
• Alpine chipmunk, Neotamias alpinus
• Buller's chipmunk, Neotamias bulleri
• California chipmunk, Neotamias obscurus
• Cliff chipmunk, Neotamias dorsalis
• Colorado chipmunk, Neotamias quadrivittatus
• Durango chipmunk, Neotamias durangae
• Gray-collared chipmunk, Neotamias cinereicollis
• Gray-footed chipmunk, Neotamias canipes
• Hopi chipmunk, Neotamias rufus
• Least chipmunk, Neotamias minimus
• Lodgepole chipmunk, Neotamias speciosus
• Long-eared chipmunk, Neotamias quadrimaculatus
• Merriam's chipmunk, Neotamias merriami
• Palmer's chipmunk, Neotamias palmeri
• Panamint chipmunk, Neotamias panamintinus
• Red-tailed chipmunk, Neotamias ruficaudus
• Siskiyou chipmunk, Neotamias siskiyou
• Sonoma chipmunk, Neotamias sonomae
• Townsend's chipmunk, Neotamias townsendii
• Uinta chipmunk, Neotamias umbrinus
• Yellow-cheeked chipmunk, Neotamias ochrogenys
• Yellow-pine chipmunk, Neotamias amoenus

{{Div col end}}

Genus Nototamias †{{Cite book |last1=Fostowicz-Frelik |first1=Lucja A. |url=https://books.google.com/books?id=GU3iEAAAQBAJ&pg=PA27 |title=Biotic changes in terrestrial environments around the Eocene–Oligocene transition |last2=Li |first2=Qian |last3=Métais |first3=Grégoire |date=2023-11-09 |publisher=Frontiers Media SA |isbn=978-2-8325-3864-7 |language=en}}

• Nototamias ateles †
• Nototamias hulberti †
• Nototamias quadratus †

• Alvin and the Chipmunks, an animated virtual band
• Chip 'n' Dale, cartoon Disney chipmunks

{{Reflist}}

• {{Cite journal |last1=Baack |first1=Jessica K. |last2=Switzer |first2=Paul V. |date=December 2003 |title=Alarm Calls Affect Foraging Behavior in Eastern Chipmunks (Tamias Striatus, Rodentia: Sciuridae) |url=https://onlinelibrary.wiley.com/doi/abs/10.1046/j.1439-0310.2000.00619.x |url-access=subscription |journal=Ethology |volume=106 |issue=12 |pages=1057–1066 |doi=10.1046/j.1439-0310.2000.00619.x |access-date=}} Derived from [https://thekeep.eiu.edu/theses/1641/ Baack's master's thesis].
• {{Cite book |last=Gordon |first=Kenneth Llewellyn |year=1943 |title=The Natural History and Behavior of the Western Chipmunk and the Mantled Ground Squirrel |url= |url-access= |series=Oregon State College Monographs. Studies in zoology, no. 5 |location=Corvallis, Or. |publisher=Oregon State College |pages= |lccn=44053700 |oclc=752445896 |access-date=}}

• {{webarchive |url=https://web.archive.org/web/20080424111229/http://www.britishcolumbia.com/Wildlife/wildlife/landmammals/cw/cw_chipmunk.html |date=April 24, 2008 |title=Wildlife and Nature – Chipmunk }}

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Category:Extant Miocene first appearances

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