Actinodon

Actinodon was named in 1866 by French paleontologist Jean Albert Gaudry based on a holotype skull that was collected by Charles Frossard near Muse in the Autun Basin (early Permian) of France.{{Cite journal |last=Gaudry |first=Jean A. |date=1866 |title=Sur le reptile découvert par M. Frossard, à la partie supériere du terrain houiller de Muse, près Autun (Saône-et-Loire) |url=https://www.biodiversitylibrary.org/item/23701#page/347/mode/1up |journal=Comptes Rendus Hebdomadaires des Séances de l'Académie des Sciences de Paris |volume=63 |pages=341–344 |via=Biodiversity Heritage Library}} The status and relationship of the taxon was long problematic because the holotype was thought to be lost, until it was rediscovered in the collections of the Museum d'Histoire Naturelle in 1996.{{Cite journal |last=Werneburg |first=Ralf |date=1997 |title=Der Eryopidae Onchiodon (Amphibia) aus dem Rotliegend des Beckens von Autun (Frankreich) |url=https://www.researchgate.net/publication/284048038 |journal=Freiberger Forschungsheft |volume=466 |pages=167–181 |via=ResearchGate}} In the intervening 130 years, a variety of specimens were described by other workers, some of which were attributed to other species or only to the genus level.{{Cite journal |last=Gaudry |first=Jean A. |date=1883 |title=Enchainements du monde animal |journal=Fossiles Primaires |volume=25 |pages=264–272}}{{Cite journal |last=Thevenin |first=Armand |date=1910 |title=Les plus anciens quadrupèdes de France |journal=Annales de Paléontologie |volume=5 |page=65}}{{Cite journal |last1=Dutuit |first1=Jean-Michel |last2=Heyler |first2=Daniel |date=1994 |title=Rachitomes, Lépospondyles et Reptiles due Stephanien (Carbonifere superieur) du basin de Montceau-les-Mines (Massif central, France) |journal=Mémoires de la Section des Science |volume=12 |pages=249–266}} Werneburg & Steyer (1999) were the most recent to redescribe material of this taxon,{{Cite journal |last1=Werneburg |first1=Ralf |last2=Steyer |first2=J.-Sébastien |date=1999 |title=Redescription of the holotypeof Actinodon frossardi (Amphibia, Temnospondyli) from the Lower Permian of the Autun basin (France) |url=https://linkinghub.elsevier.com/retrieve/pii/S0016699599800096 |journal=Geobios |language=en |volume=32 |issue=4 |pages=599–607 |doi=10.1016/S0016-6995(99)80009-6|bibcode=1999Geobi..32..599W }} and they referred it to the eryopoid Onchiodon as a valid species, while Schoch & Milner (2000) argued that it might be a species of the stereospondylomorph Sclerocephalus,{{Cite book |last1=Schoch |first1=Rainer R. |url=https://www.worldcat.org/oclc/580976 |title=Handbuch der Paläoherpetologie Part 3B |last2=Milner |first2=Andrew R. |date=2000 |publisher=Verlag Dr. Friedrich Pfeil |isbn=978-3-931516-26-0 |location=Stuttgart |pages=1–220 |oclc=580976}} but phylogenetic analyses have not recovered A. frossardi in a clade with the type species of either genus (O. labyrinthicus, S. haeuseri),{{Cite journal |last1=SCHOCH |first1=RAINER R. |last2=WITZMANN |first2=FLORIAN |date=2009 |title=Osteology and relationships of the temnospondyl genusSclerocephalus |journal=Zoological Journal of the Linnean Society |volume=157 |issue=1 |pages=135–168 |doi=10.1111/j.1096-3642.2009.00535.x |issn=0024-4082|doi-access=free }}{{Cite journal |last=Schoch |first=Rainer R. |date=2021-09-30 |title=The life cycle in late Paleozoic eryopid temnospondyls: developmental variation, plasticity and phylogeny |journal=Fossil Record |volume=24 |issue=2 |pages=295–319 |doi=10.5194/fr-24-295-2021 |s2cid=244200015 |issn=2193-0074|doi-access=free }} and Schoch & Milner (2014) maintained it separate from Onchiodon.{{Cite book |last1=Schoch |first1=Rainer R. |url=https://www.worldcat.org/oclc/580976 |title=Handbuch der Paläoherpetologie Part 3A2. Temnospondyli |last2=Milner |first2=Andrew R. |date=2014 |publisher=Verlag Dr. Friedrich Pfeil |isbn=978-3-931516-26-0 |location=Stuttgart |pages=1–150 |oclc=580976}}

Other species of Actinodon remain synonymized with other species: A. brevis and Euchirosaurus rochei with A. frossardi; and A. germanicus with Cheliderpeton vranyi.{{Cite journal |last1=Werneburg |first1=Ralf |last2=Steyer |first2=J. Sébastien |date=2002 |title=Revision ofCheliderpeton vranyi Fritsch, 1877 (Amphibia, Temnospondyli) from the Lower Permian of Bohemia (Czech Republic) |url=http://link.springer.com/10.1007/BF02988193 |journal=Paläontologische Zeitschrift |language=en |volume=76 |issue=1 |pages=149–162 |doi=10.1007/BF02988193 |s2cid=129307253 |issn=0031-0220}} Several other species have been previously placed in Actinodon after being named in other genera, but have since been restored to their original genera or placed elsewhere: Glanochthon latirostris and Lysipterygium risinense.{{Cite journal |last=Boy |first=Jürgen A. |date=1993 |title=Über einige Vertreter der Eryopoidea (Amphibia: Temnospondyli) aus dem europäischen Rotliegend (?höchstes Karbon - Perm) 4. Cheliderpeton latirostre |url=http://link.springer.com/10.1007/BF02985874 |journal=Paläontologische Zeitschrift |language=de |volume=67 |issue=1–2 |pages=123–143 |doi=10.1007/BF02985874 |s2cid=126947653 |issn=0031-0220}} In the present concept, A. frossardi is the only species within the genus, and material of this taxon is only known from the early Permian of France.

Actinodon has a similar skull profile to stereospondylomorphs like Sclerocephalus, but like other eryopoids, it has a proportionately shorter and wider posterior skull table; a longer prefrontal with a pointed anterior end; and a sutured basicranial articulation. Within eryopoids, the anatomy is indeed very similar to Onchiodon, and the continued separation of these genera by Schoch & Milner (2014) is based on a disagreement over the interpretation of two features purportedly shared between them, a wide choana and palatal fangs only on the palatine and ectopterygoid, that Werneburg & Steyer (1999) identified. Schoch & Milner argued that the choana is actually slit-like, and that the size and number of teeth is more variable. Some of this also stems from perceptions over the monophyly of Onchiodon, of which there are several valid species. Material of Actinodon tends to be smaller than that of many other eryopoids, which may account for some of the more proportional differences.

Below is the phylogeny of Eryopidae from Schoch (2021):

{{Clade|{{clade

|1=Actinodon frossardi

|2={{clade

|1=Osteophorus roemeri

|2={{clade

|label1=

|1=Glaukerpeton avinoffi

|label2=

|2={{clade

|1={{clade

|1=Onchiodon labyrinthicus

|2=Onchiodon thuringiensis

}}

|label2=

|2={{clade

|1=Clamorosaurus nocturnus

|label2=

|2={{clade

|1=Eryops sp. Moran

|label2=

|2={{clade

|1=Eryops anatinus

|2=Eryops megacephalus

}} }} }} }} }} }} }}|label1=Eryopidae}}

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{{Temnospondyli|E.}}

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Category:Eryopidae

Category:Permian temnospondyls of Europe