Anaeromonadea

{{Short description|Class of excavate protists}}

{{cs1 config|name-list-style=vanc}}

{{Taxobox

| image = Monocermonoides melolanthae.jpg

| image_upright = 0.4

| image_caption = Monocercomonoides melolanthae

| domain = Eukaryota

| unranked_regnum = Excavata

| phylum = Metamonada

| subphylum = Anaeromonada

| subphylum_authority = Cavalier-Smith, 1996/7

| classis = Anaeromonadea

| classis_authority = Cavalier-Smith, 1996/7, emend. 1999

| subdivision_ranks = Orders

| subdivision = *Paratrimastigidae

• Paratrimastix
• Trimastigida
• Trimastix
• Oxymonadida

| synonyms = * Oxymonadea Grassé 1952

• Pyrsonympha
• Axostylea

}}

Anaeromonadea, also known as Preaxostyla,{{cite journal |display-authors=etal |vauthors=Hampl V, Hug L, Leigh JW |date=March 2009 |title=Phylogenomic analyses support the monophyly of Excavata and resolve relationships among eukaryotic "supergroups" |journal=Proc. Natl. Acad. Sci. U.S.A. |volume=106 |issue=10 |pages=3859–64 |bibcode=2009PNAS..106.3859H |doi=10.1073/pnas.0807880106 |pmc=2656170 |pmid=19237557 |doi-access=free}}{{Citation |last=Hampl |first=Vladimir |title=Preaxostyla |date=2016 |work=Handbook of the Protists |pages=1–36 |editor-last=Archibald |editor-first=John M. |url=https://link.springer.com/10.1007/978-3-319-32669-6_8-1 |access-date=2024-04-15 |place=Cham |publisher=Springer International Publishing |language=en |doi=10.1007/978-3-319-32669-6_8-1 |isbn=978-3-319-32669-6 |editor2-last=Simpson |editor2-first=Alastair G.B. |editor3-last=Slamovits |editor3-first=Claudio H. |editor4-last=Margulis |editor4-first=Lynn}} is a class of excavate protists, comprising the oxymonads, Trimastix,{{cite journal |vauthors =Cavalier-Smith T |title=The excavate protozoan phyla Metamonada Grassé emend. (Anaeromonadea, Parabasalia, Carpediemonas, Eopharyngia) and Loukozoa emend. (Jakobea, Malawimonas): their evolutionary affinities and new higher taxa |journal=Int. J. Syst. Evol. Microbiol. |volume=53 |issue=Pt 6 |pages=1741–58 |date=November 2003 |pmid=14657102 |doi= 10.1099/ijs.0.02548-0|url=http://ijs.sgmjournals.org/cgi/pmidlookup?view=long&pmid=14657102|doi-access=free }} and Paratrimastix. This group is studied as a model system for reductive evolution of mitochondria,{{Cite web |title=Genomics of Eukaryotes and Lateral Gene Transfer {{!}} Biocev |url=https://www.biocev.eu/en/research/cellular-biology-and-virology.4/genomics-of-eukaryotes-and-lateral-gene-transfer.40 |access-date=2024-04-15 |website=www.biocev.eu |language=en}} because it includes both organisms with anaerobic mitochondrion-like organelles (Trimastix{{Cite journal |last1=Novák |first1=Lukáš V. F. |last2=Treitli |first2=Sebastian C. |last3=Pyrih |first3=Jan |last4=Hałakuc |first4=Paweł |last5=Pipaliya |first5=Shweta V. |last6=Vacek |first6=Vojtěch |last7=Brzoň |first7=Ondřej |last8=Soukal |first8=Petr |last9=Eme |first9=Laura |last10=Dacks |first10=Joel B. |last11=Karnkowska |first11=Anna |last12=Eliáš |first12=Marek |last13=Hampl |first13=Vladimír |date=2023-12-07 |editor-last=Dutcher |editor-first=Susan K. |title=Genomics of Preaxostyla Flagellates Illuminates the Path Towards the Loss of Mitochondria |journal=PLOS Genetics |language=en |volume=19 |issue=12 |pages=e1011050 |doi=10.1371/journal.pgen.1011050 |doi-access=free |issn=1553-7404 |pmc=10703272 |pmid=38060519}} and Paratrimastix{{Cite journal |last1=Hampl |first1=Vladimir |last2=Silberman |first2=Jeffrey D. |last3=Stechmann |first3=Alexandra |last4=Diaz-Triviño |first4=Sara |last5=Johnson |first5=Patricia J. |last6=Roger |first6=Andrew J. |date=2008-01-02 |editor-last=Redfield |editor-first=Rosemary |title=Genetic Evidence for a Mitochondriate Ancestry in the 'Amitochondriate' Flagellate Trimastix pyriformis |journal=PLOS ONE |language=en |volume=3 |issue=1 |pages=e1383 |doi=10.1371/journal.pone.0001383 |doi-access=free |issn=1932-6203 |pmc=2148110 |pmid=18167542|bibcode=2008PLoSO...3.1383H }}), and those that have completely lost their mitochondria (oxymonads Monocercomonoides,{{Cite journal |last1=Karnkowska |first1=Anna |last2=Vacek |first2=Vojtěch |last3=Zubáčová |first3=Zuzana |last4=Treitli |first4=Sebastian C. |last5=Petrželková |first5=Romana |last6=Eme |first6=Laura |last7=Novák |first7=Lukáš |last8=Žárský |first8=Vojtěch |last9=Barlow |first9=Lael D. |last10=Herman |first10=Emily K. |last11=Soukal |first11=Petr |last12=Hroudová |first12=Miluše |last13=Doležal |first13=Pavel |last14=Stairs |first14=Courtney W. |last15=Roger |first15=Andrew J. |date=May 2016 |title=A Eukaryote without a Mitochondrial Organelle |url=https://linkinghub.elsevier.com/retrieve/pii/S0960982216302639 |journal=Current Biology |language=en |volume=26 |issue=10 |pages=1274–1284 |doi=10.1016/j.cub.2016.03.053|pmid=27185558 |bibcode=2016CBio...26.1274K |doi-access=free }} Streblomastix, and Blattamonas).