Archaeaspis#Key to the species
{{Short description | Extinct redlichiid trilobites from the Cambrian.}}
{{Italic title}}
{{Taxobox
| name = Archaeaspis
| fossil_range = {{Geological range|521|515|Late Atdabanian}}
| image = Archaeaspis hupei recon.jpg
| image_caption = Reconstruction of A. hupei
| regnum = Animalia
| phylum = Arthropoda
| classis = Trilobita
| ordo = Redlichiida
| subordo = Olenellina
| superfamilia = Fallotaspidoidea
| familia = Archaeaspididae
| genus = Archaeaspis
| genus_authority = Repina in Khomentovskii and Repina, 1965{{cite journal|last=Lieberman|first=B.S.|title=Phylogenetic analysis of some basal early Cambrian Trilobites, the biogeographic origins of the Eutrilobites, and the timing of the Cambrian radiation|year=2002|journal=Journal of Paleontology|volume=76|number=4|pages=692–708|url=http://paleo.ku.edu/geo/faculty/BSL/JPTrilos2002.pdf|doi=10.1666/0022-3360(2002)076<0692:paosbe>2.0.co;2|access-date=2012-08-14|archive-url=https://web.archive.org/web/20110812151230/http://www.paleo.ku.edu/geo/faculty/BSL/JPTrilos2002.pdf|archive-date=2011-08-12|url-status=dead}} non Archaeaspis Ivantsov, 2001{{cite book|title= The Rise of Animals: Evolution and diversification of the Kingdom Animalia|publisher= The Johns Hopkins University Press|location= Baltimore|isbn=978-0-8018-8679-9 |url= https://books.google.com/books?id=KsFFIrJ8IxEC&pg=PA261 |year= 2007}} = Archaeaspinus Ivantsov, 2007
| subdivision_ranks = Species
| subdivision =
- A. hupei Repina in Khomentovskii and Repina, 1965 (Type)
- A. nelsoni Lieberman, 2002
- A. macropleuron Lieberman, 2002
}}
Archaeaspis is an extinct genus of redlichiid trilobites. It lived during the late Atdabanian stage,{{cite journal|last=Sepkoski |first=Jack |title=A compendium of fossil marine animal genera (Trilobita entry) |journal=Bulletins of American Paleontology |volume=364 |pages=1–560 |year=2002 |url=http://strata.ummp.lsa.umich.edu/jack/showgenera.php?taxon=307&rank=class |accessdate=2008-01-12 |url-status=dead |archiveurl=https://web.archive.org/web/20060905162524/http://strata.ummp.lsa.umich.edu/jack/showgenera.php?taxon=307&rank=class |archivedate=2006-09-05 }} which lasted from 521 to 514 million years ago during the early part of the Cambrian Period. The first specimens were first found in Siberia, where they were originally thought to have been endemic. Later, though, more specimen were found in the Inyo Mountains in Southern California.{{Cite web |url=http://www.westernta.com/Species/A.macropleuron.htm |title=Archaeaspis macropleuron Lieberman, 2002 |access-date=2008-04-29 |archive-url=https://web.archive.org/web/20080516064608/http://www.westernta.com/Species/A.macropleuron.htm |archive-date=2008-05-16 |url-status=dead }}
An Ediacarian proarticulatan was given the same name, Archaeaspis Ivantsov, 2001. This is however a junior homonym. The name Archaeaspis Repina in Khomentovskii and Repina, 1965, for the trilobite has priority. The new valid name for the proarticulate is Archaeaspinus Ivantsov, 2007.{{Cite journal|last=Ivantsov|first=A. Yu|title=Small Vendian transversely Articulated fossils|url=https://www.academia.edu/2352394|journal=Paleontological Journal|year=2007 |language=en|volume=41|issue=2|pages=113–122|doi=10.1134/S0031030107020013 |bibcode=2007PalJ...41..113I |s2cid=86636748 |issn=0031-0301}}
Etymology
The genus name is the combination of the Ancient Greek ἀρχαῖος (archaīos) meaning "ancient" and ἀσπίς (aspis) meaning "shield". The names of the species have the following derivations.
- A. hupei is named in honor of Pierre Hupé, a distinguished French paleontologist.
- A. nelsoni in named in honor of Clem Nelson, emeritus professor, who worked at the University of California at Los Angeles, and at the White Mountain Research Station. He collected the material the specimens the description of this species is based upon.
- A. macropleuron is the combination of the Greek words μακρος (makros) meaning "large" and πλευρών (pleuron) meaning "rib", named for the macropleural spine of the 3rd thorax segment in this species.
Distribution
- A. hupei is found on the Siberian platform.{{citation|last1=Palmer|first1=A.R.|last2=Repina|first2=L.N.|year=1993|title=Through a Glass Darkly: Taxonomy, Phylogeny and Biostratigraphy of the Olenellina|journal=The University of Kansas Paleontological Contributions|volume=3|pages=1–35|url=http://kuscholarworks.ku.edu/dspace/handle/1808/3760}}
- A. nelsoni was collected in the Lower Cambrian of California, USA (probably Nevadella-zone), Inyo County; on a small hill just south of the east-west road leading to Silver Canyon, Montenegro Member of the Campito Formation, Blanco Mountains; and on a south facing dip slope on the North side of a small east-west canyon on the east side of the road, Campito Formation, near Bishop.
- A. macropleuron occurs in the Lower Cambrian of California, USA (probably Nevadella-zone), Inyo County; on a small hill just south of the east-west road leading to Silver Canyon, Montenegro Member of the Campito Formation, Blanco Mountains; Campito Formation near Waucoba Springs.
Description
As with most early trilobites, Archaeaspis has an almost flat exoskeleton that is only thinly calcified, and has crescent-shaped eye ridges. As part of the Olenellina suborder, Archaeaspis lack dorsal sutures. The superfamily Fallotaspidoidea, to which Archaeaspis belongs can be distinguished from all other Olenellina by features of the cephalon and in particular the glabella. The glabella tapers forward. The frontal lobe of the glabella (because it is counted from the back, it is numbered L4) is as long as the most backward lobe (L0), less than in the other Olenellina. The eye ridges (or ocular lobes) contact, but do not merge with, the entire frontal margin of the glabella.{{cite journal |last1= Lieberman |first1=B. S. |year=1998 |title=Cladistic analysis of the Early Cambrian olenelloid trilobites |journal=Journal of Paleontology |volume=72 |pages=59–78 |url=http://jpaleontol.geoscienceworld.org/content/72/1/59.short|doi=10.1017/S0022336000024021 |bibcode=1998JPal...72...59L |s2cid=86307339 |url-access=subscription }} In Archaeaspis the frontal lobe of the glabella (L4) does not contact the anterior border furrow, but is connected with it by a ridge at midline (called plectrum). There is an obvious ridge that crosses the area between the ocular lobe and the glabella backward and slightly outward at approximately 10° (called interocular ridge).
Species key
References
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Category:Cambrian trilobites of North America