Bordered white
{{Short description|Species of moth}}
__NOTOC__
{{Speciesbox
| image = Bupalus pinaria male par Nemos.jpg
| image_caption = Upperwings of adult male
of a southern population
| taxon = Bupalus piniaria
| authority = (Linnaeus, 1758)
| synonyms = {{collapsible list|
- Bupalus piniarius (lapsus)
- Phalaena piniaria Linnaeus, 1758
}}}}
The bordered white or pine looper (Bupalus piniaria), is a moth of the family Geometridae. Among these, it belongs to tribe Bupalini of the subfamily Ennominae. B. piniaria is a common species throughout the western Palearctic region, the Near East and North Africa. However, its presence in certain regions – e.g. the northern Balkans – is doubtful.Skinner (1984), Chinery (2007): 180, FE (2009) East it is found to Siberia
and Amur Oblast.
Three subspecies are generally recognized, while two additional ones are doubtfully distinct:FE (2009)
- Bupalus piniaria bernieri de Lajonquiere, 1958
- Bupalus piniaria espagnolus Eitschberger & Steiniger, 1975
- Bupalus piniaria flavescens White, 1876 (usually included in piniaria)
(MHNT) Bupalus piniaria flavescens - Kosov Slovaquie - Male dorsal.jpg|MHNT Bupalus piniaria flavescens ♂
(MHNT) Bupalus piniaria flavescens - Kosov Slovaquie - Male ventral.jpg|MHNT Bupalus piniaria flavescens ♂ △
- Bupalus piniaria mughusaria Gumppenberg, 1887 (usually included in piniaria)
- Bupalus piniaria piniaria (Linnaeus, 1758)
In addition, many forms (e.g. kolleri) have also been named.Reissner (1942)Prout, L. B. (1912–16). Geometridae. In A. Seitz (ed.) The Macrolepidoptera of the World. The Palaearctic Geometridae, 4. 479 pp. Alfred Kernen, Stuttgart.[https://archive.org/details/macrolepidoptera1216seit pdf]
Description and ecology
{{Entomology glossary hatnote}}
This moth is an inhabitant of coniferous woodland. The adults fly in May and June,{{Cite web|url=http://www.habitas.org.uk/moths/species.asp?item=6046|title=Bordered White|last=Thompson, R. S. & Nelson, B.|date=August 2023|website=Habitas}} sometimes later (up to August or so) in the north of the range. Their wingspan is 34–40 mm. This is a variable species with strong sexual dimorphism, always conspicuous in the antennae which are combed in the males and plain in the females. Females, particularly when filled with ripe eggs, also have a plumper abdomen.Skinner (1984), Chinery (2007): 180–181Pasi Sihvonen and Peder Skou , 2015 In: Axel Hausmann (Hrsg.):, 2015 The Geometrid Moths of Europe. 1. Auflage. Volume 5: Ennominae I, Stenstrup 2015, 978-90-04-26573-8
(MHNT) Bupalus piniaria - Kosov Slovaquie - Male dorsal.jpg|Bupalus piniaria ♂
(MHNT) Bupalus piniaria - Kosov Slovaquie - Male ventral.jpg| ♂ △
(MHNT) Bupalus piniaria - Kosov Slovaquie - Female dorsal.jpg|Bupalus piniaria♀
(MHNT) Bupalus piniaria - Kosov Slovaquie - Female ventral.jpg| ♀ △
The male has upperwings with broad dark brown borders and spots and a background varying from white in the north to deep yellow in southern populations. The female is plainer, varying from yellow to brown on the upperwings, which have slightly darker crosswise stripes. In both sexes, the wingtips are darkest. The underwings are less dimorphic, orange-brown with darker tips on the forewings and marbled light brown with a whitish lengthwise stripe on the hindwings in both sexes. The male's underwings have a wider whitish hindwing stripe and darker forewing tips, while the females have a more contrasting hindwing pattern. All four wings are bordered by a short fringe of alternating sections of white and dark brown hairs. Bilateral gynandromorphs are easily recognized in this species.Reissner (1942), Skinner (1984), Chinery (2007): 180–181
The caterpillar (also known as a larva) can be a serious pest in conifer plantations.Skinner (1984), Chinery (2007): 180, and see references in Savela (2001) They are green with pale lines and usually feed on various species of pine (Pinus), especially Scots pine (P. sylvestris) and European black pine (P. nigra). It has also been recorded feeding on Douglas-fir (Pseudotsuga menziesii), larch (Larix) and spruce (Picea, e.g. Norway spruce P. abies). [http://www.nhm.ac.uk/research-curation/research/projects/hostplants/search/index.dsml HOSTS database, National History Museum, London]. Retrieved 2024Larvae have four to six instars and pupate in the soil where they overwinter.{{cite book |last1=Leverton |first1=Roy |title=Enjoying Moths |date=2001 |publisher=T & A D Poyser Ltd |location=London |isbn=0-85661-124-7 |pages=16–17}}
File:Bupalus piniaria femelle par Nemos.jpg|Upperwings of southern adult female
File:Bupalus.piniaria.7489.jpg|Underwings of adult male from Dresden (Germany)
File:Bupalus.piniaria.7513.jpg|Underwings of adult female from Dresden (Germany)
File:Bupalus_piniaria_ova.jpg|Eggs
File:Männivaksiku röövik.jpg|Early instar larvae
File:Bupalus piniarius 1 beentree.jpg|Caterpillar
Taxonomy
It is (under its original scientific name Phalaena piniaria) the type species of its genus Bupalus, as well as the junior objective synonyms Catograpta, Chleuastes and Phaophyga, and the preoccupied Bupala. Via its genus, it is also the type of the Bupalini.Pitkin & Jenkins (2004) Bupalus was raised by the English zoologist William Elford Leach, in 1815 and is the name of a 6th-century BC Greek sculptor. The specific name piniaria refers to Pinus the pine tree genus and food-plant.{{cite book |last1=Emmet |first1=A Maitland |author1-link=A. Maitland Emmet |editor1-link= |title=The Scientific Names of the British Lepidoptera. Their History and Meaning |date=1991 |publisher=Harley Books |location=Colchester |isbn=0-946589-28-3 |page=183}}
Footnotes
{{Reflist}}
References
- {{aut|Chinery, Michael}} (2007): Domino Guide to the Insects of Britain and Western Europe (Revised ed.). A. & C. Black, London. {{ISBN|978-0-7136-7239-8}}
- {{aut|Pitkin, Brian & Jenkins, Paul}} (2004): [http://www.nhm.ac.uk/jdsml/research-curation/research/projects/butmoth/index.dsml Butterflies and Moths of the World, Generic Names and their Type-species] – [http://www.nhm.ac.uk/jdsml/research-curation/research/projects/butmoth/GenusDetails.dsml?NUMBER=4421.0 Bupalus]. Version of 5 November 2004. Retrieved 13 May 2010.
- {{aut|Reissner, Hans}} (1942): Ein geteilter Zwitter von Bupalus piniarius L. ["A bilateral gynandromorph of the Bordered White"]. Zeitschrift des Wiener Entomologen-Vereins 27: 287-288 [in German]. [http://www.biologiezentrum.at/pdf_frei_remote/ZOEV_27_0287-0288.pdf PDF fulltext]
- {{aut|Savela, Markku}} (2001): Markku Savela's Lepidoptera and some other life forms – [http://www.nic.funet.fi/pub/sci/bio/life/insecta/lepidoptera/ditrysia/geometroidea/geometridae/ennominae/bupalus/index.html#piniarius Bupalus piniarius] [sic]. Version of 1 October 2001. Retrieved 13 May 2010.
- {{aut|Skinner, Bernard}} (1984): Colour Identification Guide to Moths of the British Isles.
External links
{{Commons category|Bupalus piniaria}}
- [https://www.ukmoths.org.uk/species/bupalus-piniaria/ Bordered white at UKMoths]
- [https://web.archive.org/web/20060824204227/http://kimmos.freeshell.org/lnel/g4/Buppinia.htm Bordered white at kimmos.freeshell.org]
- [https://projects.biodiversity.be/lepidoptera/species/5656/ Lepidoptera of Belgium]
- [https://treesforlife.org.uk/into-the-forest/habitats-and-ecology/ecology/herbivory/ Trees for Life]
- [http://www.vlindernet.nl/vlindersoort.php?vlinderid=383 Vlindernet.nl] {{in lang|nl}}
{{Taxonbar|from=Q24342}}