Carangiformes
{{Short description|Order of fishes}}{{Automatic taxobox
| fossil_range = {{fossil range|Late Paleocene|present}}
| image = Trevally Nick Hobgood.jpg
| image_caption = Bluefin trevally (Caranx melampygus)
| image2 = Remora remora 2.jpg
| image2_caption = Remora remora
| taxon = Carangiformes
| authority = Jordan, 1923{{cite web | url = http://taxonomicon.taxonomy.nl/TaxonTree.aspx?id=201902&src=0 | title = Taxon: Order Carangiformes Jordan, 1923 (fish) | access-date = 15 November 2019 | website = Taxonomicon}}
| type_species = Caranx praeustus
Anonymous [Bennett], 1830
}}
Carangiformes is a large, diverse order of ray-finned fishes within the clade Percomorpha. It is part of a sister clade to the Ovalentaria, alongside its sister group, the Anabantaria (including Anabantiformes and Synbranchiformes). The Carangiformes have been long regarded as a monotypic order with only the family Carangidae within it by some authorities, and the other current families within the order have been previously classified as part of the wider order Perciformes. The 5th edition of Fishes of the World classify six families within Carangiformes,{{cite book |title=Fishes of the World |edition=5th |author1=J. S. Nelson |author2=T. C. Grande |author3=M. V. H. Wilson |year=2016 |pages=380–383 |publisher=Wiley |isbn=978-1-118-34233-6 |url=https://sites.google.com/site/fotw5th/ |access-date=2019-11-15 |archive-date=2019-04-08 |archive-url=https://web.archive.org/web/20190408194051/https://sites.google.com/site/fotw5th/ |url-status=dead }} with more recent authorities expanding the order to include up to 30 families, based on phylogenetic evidence.{{Cite journal|last1=Girard|first1=Matthew G.|last2=Davis|first2=Matthew P.|last3=Smith|first3=W. Leo|date=2020-05-08|title=The Phylogeny of Carangiform Fishes: Morphological and Genomic Investigations of a New Fish Clade|journal=Copeia|volume=108|issue=2|pages=265|doi=10.1643/CI-19-320|issn=0045-8511|doi-access=free}}
The earliest known carangiforms are two fossil species of Mene, Mene purydi from Peru and Mene phosphatica from Tunisia, both of which are known from the Late Paleocene.{{Cite journal |last1=Friedman |first1=Matt |last2=V. Andrews |first2=James |last3=Saad |first3=Hadeel |last4=El-Sayed |first4=Sanaa |date=2023-06-16 |title=The Cretaceous–Paleogene transition in spiny-rayed fishes: surveying "Patterson's Gap" in the acanthomorph skeletal record André Dumont medalist lecture 2018 |url=https://popups.uliege.be/1374-8505/index.php?id=7048 |journal=Geologica Belgica |language=en |doi=10.20341/gb.2023.002 |issn=1374-8505|doi-access=free }}
Taxonomy
This order has often been either subsumed within Perciformes or used exclusively to refer to families classified within the suborder Carangoidei. However, more recent studies using genetic data have found such a placement to be paraphyletic, and have incorporated many more groups into it, including the highly unusual flatfishes.
This classification is from Eschmeyer's Catalog of Fishes Classification.{{Cite web |last= |first= |last2= |title=Eschmeyer's Catalog of Fishes Classification - California Academy of Sciences |url=https://www.calacademy.org/scientists/catalog-of-fishes-classification/ |access-date=2024-12-27 |website=www.calacademy.org |language=en}}
- Order Carangiformes
- Suborder Centropomoidei
- Family Latidae {{small|Jordan 1888}} (giant perches)
- Family Centropomidae {{small|Poey 1967}} (snooks)
- Family Lactariidae {{small|Boulenger 1904}} (false trevallies)
- Family Sphyraenidae {{small|Rafinesque 1815}} (barracudas)
- Suborder Pleuronectoidei
- Family Polynemidae {{small|Rafinesque 1815}} (threadfins or tassel-fishes)
- Family Psettodidae {{small|Regan 1910}} (spiny turbots)
- Family Citharidae {{small|de Buen 1935}} (largescale flounders)
- Family Scophthalmidae {{small|Chabanaud 1933}} (turbots)
- Family Cyclopsettidae {{small|Campbell et al. 2019}} (sand whiffs or large-tooth flounders)
- Family Bothidae {{small|Smitt 1892}} (lefteye flounders)
- Family Paralichthyidae {{small|Regan 1910}} (sand flounders)
- Family Pleuronectidae {{small|Rafinesque 1815}} (righteye flounders)
- Subfamily Atheresthinae {{small|Vinnikov, Thomson & Munroe 2018}}
- Subfamily Pleuronichthyinae {{small|Vinnikov, Thomson & Munroe 2018}}
- Subfamily Microstominae {{small|Cooper & Chapleau 1998}} (smallmouth flounders)
- Subfamily Hippoglossinae {{small|Gill 1864}} (halibuts)
- Subfamily Pleuronectinae {{small|Rafinesque 1815}} (true flounders)
- Family Paralichthodidae {{small|Regan 1920}} (peppered flounders)
- Family Oncopteridae {{small|Jordan & Goss 1889}} (remo flounders)
- Family Rhombosoleidae {{small|Regan 1910}} (South Pacific flounders)
- Family Achiropsettidae {{small|Heemstra 1990}} (southern flounders or armless flounders)
- Family Achiridae {{small|Rafinesque 1815}} (American soles)
- Family Samaridae {{small|Jordan & Goss 1889}} (crested flounders)
- Family Poecilopsettidae {{small|Norman 1934}} (bigeye flounders)
- Family Soleidae {{small|Bonaparte 1833}} (soles)
- Family Cynoglossidae {{small|Jordan 1888}} (tonguefishes)
- Subfamily Symphurinae {{small|Ochiai 1963}} (straightsnout tongue soles)
- Subfamily Cynoglossinae {{small|Jordan 1888}} (hookedsnout tongue soles)
- Suborder Toxotoidei
- Family Leptobramidae {{small|Ogilby 1913}} (beachsalmons)
- Family Toxotidae {{small|Bleeker 1859}} (archerfishes)
- Suborder Nematistioidei
- Family Nematistiidae {{small|Gill 1862}} (roosterfishes)
- Suborder Menoidei
- Family Menidae {{small|Fitzinger 1873}} (moonfishes)
- Family Xiphiidae {{small|Rafinesque 1815}} (swordfishes)
- Family Istiophoridae {{small|Rafinesque 1815}} (billfishes and marlins)
- Suborder Carangoidei
- Family Carangidae {{small|Rafinesque 1815}} (jacks or jack mackerels)
- Subfamily Naucratinae {{small|Bleeker 1859}} (amberjacks)
- Subfamily Caranginae {{small|Rafinesque 1815}} (trevallies and kingfishes)
- Subfamily Scomberoidinae {{small|Jordan & Gilbert 1883}} (leatherjackets and queenfishes)
- Subfamily Trachinotinae {{small|Gill 1861}} (pompanos)
- Family Echeneidae {{small|Rafinesque 1810}} (remoras and sharksuckers)
- Family Rachycentridae {{small|Gill 1896}} (cobias)
- Family Coryphaenidae {{small|Rafinesque 1815}} (dolphinfishes)
The Coryphaenidae, Rachycentridae, and Echeneidae have been suggested to comprise a monophyletic grouping, which has been recovered as a sister clade to the Carangidae. A basal member of this clade is thought to be Ductoridae from the early Eocene.{{Cite journal |last1=Friedman |first1=Matt |last2=Johanson |first2=Zerina |last3=Harrington |first3=Richard C. |last4=Near |first4=Thomas J. |last5=Graham |first5=Mark R. |date=2013-09-07 |title=An early fossil remora (Echeneoidea) reveals the evolutionary assembly of the adhesion disc |journal=Proceedings of the Royal Society B: Biological Sciences |volume=280 |issue=1766 |pages=20131200 |doi=10.1098/rspb.2013.1200 |pmc=3730593 |pmid=23864599}}
= Traditional Classification =
In past classifications such as Fishes of the World 5, Carangiformes were restricted to these families. This placement is now known to be paraphyletic:
- Nematistiidae Gill{{cite journal | author1 = Richard van der Laan | author2 = William N. Eschmeyer | author3 = Ronald Fricke | name-list-style = amp |year=2014 | title = Family-group names of Recent fishes | url = https://biotaxa.org/Zootaxa/article/view/zootaxa.3882.1.1/10480 | journal = Zootaxa | volume = 3882 | issue =2 | pages = 001–230}} (roosterfish)
- Coryphaenidae Rafinesque, 1815 (dolphinfish)
- Rachycentridae Gill 1896 (cobia)
- Echeneidae Rafinesque, 1815 (remoras)
- Carangidae Rafinesque, 1815 (jacks)
- Menidae Fitzinger, 1873 (moonfishes)
