Chlamyphoridae
{{Short description|Family of armadillos}}
{{Automatic taxobox
| fossil_range = Middle Eocene to present
| image = SouthernThreeBandedArmadillo065b.jpg
| image_caption = Southern three-banded armadillo (Tolypeutes matacus)
| image2 = Doedicurus.gif
| image2_caption = Illustration of a skeleton of Doedicurus clavicaudatus
| taxon = Chlamyphoridae
| authority = Bonaparte, 1850
| type_genus = Chlamyphorus
| type_genus_authority = Harlan, 1825
| subdivision_ranks = Subfamilies
| subdivision = *Chlamyphorinae
| range_map = Chlamyphoridae Range map.jpg
}}
File:Chlamyphorus truncatus - Naturmuseum Senckenberg - DSC02081.JPG)]]
Chlamyphoridae is a family of cingulate mammals. While glyptodonts have traditionally been considered stem-group cingulates outside the group that contains modern armadillos, there had been speculation that the extant family Dasypodidae could be paraphyletic based on morphological evidence.Simpson, G. G. (1945). The principles of classification and a classification of mammals. Bull. Amer. Museum Nat. History., 85.{{cite journal | last1 = Grassé | first1 = P. P. | year = 1955 | title = Ordre des édentés | journal = Traité de zoologie | volume = 17 | issue = 2| pages = 1182–1246 }}Engelmann, G. F. (1985). The phylogeny of the Xenarthra. The evolution and ecology of armadillos, sloths, and vermilinguas. Smithsonian Institution Press, Washington, DC, 51-64.Wible, J. R. (2006). 6 The Phylogeny of Living and Extinct Armadillos (Mammalia, Xenarthra, Cingulata): A Craniodental Analysis. Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles: University of Chicago Press, Chicago, IL, 153-198. In 2016, an analysis of Doedicurus mtDNA found it was, in fact, nested within the modern armadillos as the sister group of a clade consisting of Chlamyphorinae and Tolypeutinae.{{cite journal|last1=Delsuc|first1= F.|last2= Gibb|first2=G. C.|last3=Kuch|first3=M. |last4=Billet|first4=G.|last5= Hautier|first5=L.|last6=Southon|first6= J.|last7= Rouillard|first7=J.-M.|last8=Fernicola|first8=J. C.|last9=Vizcaíno|first9=S. F.|last10=MacPhee|first10=R. D. E.|last11= Poinar|first11=H. N.|title=The phylogenetic affinities of the extinct glyptodonts|journal=Current Biology|volume= 26|issue=4|date= 2016-02-22|pages=R155–R156|doi=10.1016/j.cub.2016.01.039|url=https://hal.archives-ouvertes.fr/hal-01879335|pmid=26906483|doi-access=free|bibcode= 2016CBio...26.R155D|hdl=11336/49579|hdl-access=free}}{{cite journal |author1=Gillian C. Gibb |author2=Fabien L. Condamine |author3=Melanie Kuch |author4=Jacob Enk |author5=Nadia Moraes-Barros |author6=Mariella Superina |author7=Hendrik N. Poinar |author8=Frédéric Delsuc |year=2016 |title=Shotgun Mitogenomics Provides a Reference Phylogenetic Framework and Timescale for Living Xenarthrans |journal=Molecular Biology and Evolution |volume= 33|issue=3 |pages= 621–642|doi=10.1093/molbev/msv250 |pmid=26556496 |pmc=4760074}} For this reason, all extant armadillos but Dasypus were relocated to a new family. The scientific name comes from Ancient Greek {{lang|grc|χλαμύς}} ({{transliteration|grc|khlamús}}), "cloak", and {{lang|grc|φόρος}} ({{transliteration|grc|phóros}}), "bearing".{{cn|date=March 2025}}
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Classification
{{See also|List of cingulates}}
Below is a taxonomy of the extant species of armadillos in this family.
Family Chlamyphoridae
- Subfamily Chlamyphorinae
- Genus Calyptophractus
- Greater fairy armadillo, Calyptophractus retusus
- Genus Chlamyphorus
- Pink fairy armadillo, Chlamyphorus truncatus
- Subfamily Euphractinae
- Genus Euphractus
- Six-banded armadillo, Euphractus sexcinctus
- Genus Zaedyus
- Pichi, Zaedyus pichiy
- Genus Chaetophractus
- Screaming hairy armadillo, Chaetophractus vellerosus
- Big hairy armadillo, Chaetophractus villosus
- Andean hairy armadillo, Chaetophractus nationi
- Subfamily Tolypeutinae
- Genus Cabassous
- Greater naked-tailed armadillo, Cabassous tatouay
- Chacoan naked-tailed armadillo, Cabassous chacoensis
- Northern naked-tailed armadillo, Cabassous centralis
- Southern naked-tailed armadillo, Cabassous unicinctus
- Genus Priodontes
- Giant armadillo, Priodontes maximus
- Genus Tolypeutes
- Southern three-banded armadillo, Tolypeutes matacus
- Brazilian three-banded armadillo, Tolypeutes tricinctus
Phylogeny
Chlamyphoridae, like Dasypodidae, is a basal clade within Cingulata, as shown below.
{{cladogram
|style=font-size:90%;line-height:100%;width:590px;
|width=590
|align=left
{{cite journal
|last1=Upham |first1=Nathan S.
|last2=Esselstyn |first2=Jacob A.
|last3=Jetz |first3=Walter
|date=2019
|title=Inferring the mammal tree: Species-level sets of phylogenies for questions in ecology, evolution and conservation
|journal=PLOS Biol
|volume=17 |issue=12 |page=e3000494
|doi=10.1371/journal.pbio.3000494 |doi-access=free
|pmid=31800571 |pmc=6892540
}}
{{cite journal
|last1=Gibb |first1=Gillian C. |last2=Condamine |first2=Fabien L.
|last3=Kuch |first3=Melanie |last4=Enk |first4=Jacob
|last5=Moraes-Barros |first5=Nadia |last6=Superina |first6=Mariella
|last7=Poinar |first7=Hendrik N. |last8=Delsuc |first8=Frédéric
|year=2015
|title=Shotgun mitogenomics provides a reference phylogenetic framework and timescale for living xenarthrans
|journal=Molecular Biology and Evolution
|volume=33 |issue=3 |pages=621–642
|doi=10.1093/molbev/msv250
|pmid=26556496 |pmc=4760074
}}
|1={{clade
|label1=Cingulata
|1={{clade
|label1=Chlamyphoridae
|1={{clade
|label1=Euphractinae
|1={{clade
|label1=
|1={{clade
|label1=Euphractus
|1={{clade
|label1=
}}
}}
|2={{clade
|label1=Zaedyus
|1={{clade
|label1=
|1=Z. pichiy
}}
|label2=Chaetophractus
|2={{clade
|label1=
|1=C. villosus
|2={{clade
|label1=
|1=C. nationi
}}
}}
}}
}}
|2={{clade
|label1=
|1={{clade
|label1={{extinct}}Glyptodontinae
|1={{clade
|label1=
|1={{clade
|label1={{extinct}}Doedicurus
|1={{extinct}}D. clavicaudatus
}}
|2={{extinct}}(31 other extinct genera)
}}
}}
|2={{clade
|label1=Chlamyphorinae
|1={{clade
|label1=
}}
|label2=Tolypeutinae
|2={{clade
|label1=
|1={{clade
|label1=Priodontes
|1=P. maximus
}}
|2={{clade
|label1=Tolypeutes
|1={{clade
|label1=
|2=T. matacus
}}
|label2=Cabassous
|2={{clade
|label1=
|1={{clade
|label1=
|1=C. tatouay
}}
|2={{clade
|label1=
|2={{clade
|label1=
|1=C. centralis
}}
}}
}}
}}
}}
}}
}}
}}
|label2=Dasypodidae
|2={{clade
|label1=
|1={{clade
|label1=
|1={{clade
|label1=Dasypus
|1=(7 living species)
}}
|2={{extinct}}(26 extinct genera)
}}
}}
}}
}}
}}
{{clear left}}
References
{{Commons}}
{{Wikispecies}}
{{Wikiquote}}
{{Reflist}}
{{Mammals}}
{{Cingulata}}
{{Taxonbar|from=Q22924208}}