Coccinelloidea

{{Automatic taxobox

| oldest_fossil = Early Cretaceous

| image = 7-Spotted-Ladybug-Coccinella-septempunctata-sq1.jpg

| image_caption = Seven-Spotted Ladybug
(Coccinella septempunctata)

| taxon = Coccinelloidea

| authority = Latreille, 1807

| display_parents = 2

| subdivision_ranks = Families

| subdivision = see text

}}

Coccinelloidea is a superfamily of beetles in the order Coleoptera, formerly included in the superfamily Cucujoidea. There are more than 10,000 species in Coccinelloidea, including more than 6000 in the lady beetle family Coccinellidae.

Morphology

Adult Coccinelloidea have a reduced tarsal formula (each tarsus with 4 or 3 segments), hind coxae separated by more than 1/3 their width, the intercoxal process of abdominal ventrite 1 usually broadly rounded or truncate, hindwings with reduced anal veins and lacking a closed radial cell, the adeagus resting on its side when retracted and the phallobase usually reduced.

Larval Coccinelloidea have a unisetose pretarsal claw, spiracles that are usually annular, and the sensory appendage of the second antennal segment usually as long as the third segment.

Families

The family constituency of this lineage has changed considerably over time, from as few as four or five recognized families in 1970{{cite book |author1=Donald Borror |author2=Richard White |title=A field guide to the insects of America north of Mexico |url=https://archive.org/details/fieldguidetoinse00borr |url-access=registration |year=1970 |publisher=Houghton Mifflin |isbn=978-0-395-07436-7}} to 15 in 2015.{{cite journal |last1=Robertson |first1=James A. |last2=Ślipiński |first2=Adam |last3=Moulton |first3=Matthew |last4=Shockley |first4=Floyd W. |last5=Giorgi |first5=Adriano |last6=Lord |first6=Nathan P. |last7=McKenna |first7=Duane D. |last8=Tomaszewska |first8=Wioletta |last9=Forrester |first9=Juanita |last10=Miller |first10=Kelly B. |last11=Whiting |first11=Michael F. |last12=McHugh |first12=Joseph V. |display-authors=3 |title=Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia) |year=2015 |journal=Systematic Entomology |volume=40 |issue=4 |pages=745–778 |doi=10.1111/syen.12138 |s2cid=55206626 }}

{{clade

|label1=Cucujiformia

|1={{clade

|1=Lymexyloidea File:Lymexylon navale (adult).jpg

|2=Tenebrionoidea File:Nephodinus metallescens (Kuester, 1846) (10113622594).png

}}

|label2=Cleroidea

|2=File:Thaneroclerus buqueti (Lefebvre, 1835).png

}}

|2={{clade

|1={{clade

|1=Chrysomeloidea File:Chrysomela vigintipunctata (Scopoli, 1763).jpg