Eastern sand darter
{{Short description|Species of fish}}
{{Speciesbox
| image = Ammocrypta_pellucida.jpg
| status = LC
| status_system = IUCN3.1
| taxon = Ammocrypta pellucida
| authority = (Putnam, 1863)
| synonyms =
- Pleurolepis pellucidus Putnam, 1863
- Etheostoma pellucidum (Putnam, 1863)
| synonyms_ref = {{Fishbase|Ammocrypta|pellucida|month=December|year=2019}}
}}
The eastern sand darter (Ammocrypta pellucida) is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. The eastern sand darter is a relatively small fish, most plentiful in the Mississippi and Ohio Rivers, as well as Lake Champlain and the Great Lakes. It prefers sandy-bottomed streams and sandy shoals in the lakes.{{cite journal | author = Daniels, R.A. | title = Habitat of the Eastern Sand Darter, Ammocrypta-Pellucida | journal = Journal of Freshwater Ecology | volume = 8 | pages = 462–479 | year =1993| issue = 4 | doi = 10.1080/02705060.1993.9664868 }} The eastern sand darter feeds on larvae of black flies and other small insects in the water. They also feed on zooplankton in small portions since their small mouth size limits their gape ability.{{cite journal | author1 = Drake, D.A.R. | author2 = M. Power | author3 = M.A. Koops | author4 = S.E. Doka | author5 = N.E. Mandrak | name-list-style = amp | title = Environmental Factors affecting growth of eastern sand darter (Ammocrypta pellucida) | journal = Canadian Journal of Zoology | volume = 86 | pages = 714–722 | year =2008| issue = 7 | doi = 10.1139/Z08-037 }} Its average body size is around 1.5-2 in long, and it breeds in the spring and summer in sandy-bottomed waters.{{cite journal | author1 = O'Brien, Shannon M. | author2 = Douglas E. Facey | name-list-style = amp | title = Habitat Use by the Eastern Sand Darter, Ammocrypta pellucida, in Two Lake Champlain Tributaries | journal = Canadian Field-Naturalist | volume = 122 | pages =239–246 | year = 2008| issue = 3 | doi = 10.22621/cfn.v122i3.606 | doi-access = free }}
Description
The average lifespan for the eastern sand darter is about 2–3 years, but if the siltation and pollution continue to worsen, the lifespan will gradually decrease.
A. pellucida reproduces in the sandy shoals at the bottom of the lakes and rivers it inhabits during the spring and summer when the water is at its warmest temperature. It reaches sexual maturity around age one and the males are able to mate once during the breeding season, while females mate twice, producing about 350 eggs with an average clutch size of 71 eggs. If siltation continues to worsen, many of the eggs will be smothered, resulting in reduced reproduction, further hindering the eastern sand darters' population count.
Distribution and habitat
The eastern sand darter can be found in many areas throughout the United States and into southern Canada. In addition to the Mississippi and Ohio Rivers, it can be found in great concentration in Lakes Champlain, Erie, Huron, Michigan and Ontario. They can also be found in the St. Lawrence River drainage in Canada along with the Lake Champlain drainage in Vermont south into New York, Pennsylvania, West Virginia, Kentucky, Illinois, and Indiana. Currently, the greatest concentration occurs in northern Alabama and eastern Tennessee.
Distribution has slowly decreased over the last few decades, and two key factors could be the cause. The eastern sand darter requires clean sand substrates; hence, siltation is a major factor in their decline. Siltation decreases the quality of habitat for both egg development and adult darters by decreasing the oxygen levels within the sand in which they bury themselves. Furthermore, silted habitats can cause adult darters to not fully burrow or decrease their time burrowed which then causes them to waste energy reserves. Channel or water flow alterations, nutrient enrichment, or any other habitat modification can completely change the amount and quality of the shifting sand bars which can then turn detrimental for the eastern sand darter.{{cite journal | author1 = Holm, E. | author2 = N.E. Mandrak | name-list-style = amp | title = The status of the Eastern sand darter,Ammocrypta Pellucida, in Canada | journal = Canadian Field-Naturalist | volume = 110 | pages = 462–469 | year =1996}} The increased construction of dams also has been a major factor contributing to the reduction of their habitats. Dams reduce river levels and flow, decrease oxygen levels in reservoir waters, and alter water temperatures, making it difficult for the eastern sand darter to reproduce at a comfortable temperature.{{cite journal | author = Daniels, R.A. | title = Significance in Burying in Ammocrypta-Pellucida. | journal = Copeia | volume = 79 | pages =92–94 | year = 1989}} Dam construction adds a whole new dimension to decreasing the reproduction and survival rate for A. pellucida.
Feeding
This fish seeks its prey from the sand along the bottom of the stream or lake. Their prey items usually range in size up to 3/16 inches, but prey selection varies depending on the age of the darter. Juvenile eastern sand darters consume small crustaceans, while adults prefer midge larvae, blackfly larvae, mayflies, and caddisflies. It has several species of fish predators, including rainbow trout, largemouth and smallmouth bass, and rock bass. However, since it spends extensive time buried in the sand and it has a translucent body, the eastern sand darter usually tends to be protected from predators. Minnows are the eastern sand darters' biggest competitor for food, but little conflict arises between the two since the darter occupies the lowest depths of the rivers and lakes while the minnows occupy the upper water column.{{cite journal | author = Simon, T.P. | title = Assessment of the range of the threatened darter, Ammocrypta pellucida, from the Maumee River basin, Indiana | journal = Proceedings of the Indiana Academy of Science | volume = 102 | pages = 139–145 | year =1993}}
Taxonomy
The Eastern sand darter was first formally described as Pleurolepsis pellucidus in 1863 by the American ichthyologist Frederic Ward Putnam (1839–1915) with the type locality given as the Black River at Elyria, Ohio.{{Cof record | spid = 9870 | title = Pleurolepsis pellucidus | access-date = 20 September 2020}} This species forms a clade with the scaly sand darter (A. vivax) and the Southern sand darter (A. meridiana).{{cite journal | author = James D. Williams | year = 1975 | title = Systematics of the Percid Fishes of the Subgenus Ammocrypta, Genus Ammocrypta, with Descriptions of Two New Species | url = https://web.b.ebscohost.com/abstract?direct=true&profile=ehost&scope=site&authtype=crawler&jrnl=01961039&AN=99556862&h=e%2b33biDSWbniAb9LSsGclt77ldYHRcoSCtrSYQOtmu%2flxarusuekSYro4v9aL98rx4P1OUaM1fSU4yGNm1ZtiQ%3d%3d&crl=c&resultNs=AdminWebAuth&resultLocal=ErrCrlNotAuth&crlhashurl=login.aspx%3fdirect%3dtrue%26profile%3dehost%26scope%3dsite%26authtype%3dcrawler%26jrnl%3d01961039%26AN%3d99556862 | journal = Bulletin of the Alabama Museum of Natural History | issue = 1 | pages = 1–56}}