Galula Formation

{{Infobox rockunit

| name = Galula Formation

| period = Late Cretaceous

| age = Aptian-Campanian
~{{fossilrange|120|75}}

| image =

| imagesize =

| caption =

| type = Geological formation

| prilithology = Sandstone

| otherlithology = Conglomerate, mudstone

| unitof = Red Sandstone Group

| subunits = Mtuka Member, Namba Member

| underlies = Unconformity with Nsungwe Formation

| overlies = Unconformity with Karoo Supergroup or basement

| thickness = {{convert|600|-|3000|m|ft|abbr=on}}

| area =

| map = {{Location map+ | Tanzania

| relief = 1

| width = 250

| float = center

| places =

{{Location map~ | Tanzania

| lat_deg = -7.1

| lon_deg = 31.2

| mark = Green pog.svg

| marksize = 12

}}

}}

| map_caption =

| location =

| coordinates = {{coord|7.1|S|31.2|E|display=inline,title}}

| paleocoordinates = {{coord|28.8|S|16.5|E|display=inline}}

| region = Rukwa Region, Songwe Region

| country = Tanzania

| extent = African Great Lakes, Rukwa Rift Basin

| namedfor = Galula

| namedby = Roberts et al.

| year_ts = 2010

| thickness_ts = {{convert|520|m|ft|abbr=on}}

| location_ts = Galula Coalfields

| coordinates_ts = {{coord|8|40|55.6|S|32|53|33.5|E|display=inline}}

}}

The Galula Formation is a geological formation located south of Lake Rukwa in Tanzania, part of the Red Sandstone Group of the Rukwa Rift Basin. Along with the unconformably overlying Oligocene Nsungwe Formation. It is divided into two members, the lower Mtuka Member and the upper Namba Member.{{Cite journal|last1=Roberts|first1=Eric M.|last2=O’Connor|first2=Patrick M.|last3=Stevens|first3=Nancy J.|last4=Gottfried|first4=Michael D.|last5=Jinnah|first5=Zubair A.|last6=Ngasala|first6=Sifael|last7=Choh|first7=Adeline M.|last8=Armstrong|first8=Richard A.|date=May 2010|title=Sedimentology and depositional environments of the Red Sandstone Group, Rukwa Rift Basin, southwestern Tanzania: New insight into Cretaceous and Paleogene terrestrial ecosystems and tectonics in sub-equatorial Africa|journal=Journal of African Earth Sciences|volume=57|issue=3|pages=179–212|doi=10.1016/j.jafrearsci.2009.09.002|bibcode=2010JAfES..57..179R |issn=1464-343X}}

The age of the deposit is poorly constrained, with the Mtuka Member likely being Aptian to Cenomanian in age, while the Namba Member being Cenomanian to Campanian in age based on Geomagnetic reversals.{{Cite journal|last1=Widlansky|first1=Sarah J.|last2=Clyde|first2=William C.|last3=O'Connor|first3=Patrick M.|last4=Roberts|first4=Eric M.|last5=Stevens|first5=Nancy J.|date=March 2018|title=Paleomagnetism of the Cretaceous Galula Formation and implications for vertebrate evolution|journal=Journal of African Earth Sciences|volume=139|pages=403–420|doi=10.1016/j.jafrearsci.2017.11.029|bibcode=2018JAfES.139..403W |issn=1464-343X|doi-access=free}} It is correlated with the Dinosaur Beds of Malawi.

The formation is fossiliferous, with Dinosaurs and Crocodyliformes being known from the formation.

Geology

= Lithology =

The Lithology of the formation is a sequence of red, pink, purple and occasionally white colored sandstones, conglomerates and mudstones. The Mtuka member is 160–180 m thick in the type section, and is typified by coarser sandstone, a higher frequency of conglomerates, higher proportions of extraformational clasts, thicker and a greater frequency of overbank siltstone and mudstone lenses and a higher proportion of paleosols. While the Namba member is between 340–360 m in the type section, and is less variable in facies, predominated by very fine-to medium-grained sandstones with less overbank mudstone and siltstone lenses.

Vertebrate paleofauna

= Fish =

! Genus !! Species

!! Stratigraphic position !! Abundance !! Notes !! Images
class="wikitable" align="center"

! colspan="7" align="center" | Fish of the Galula Formation

GenusSpecies Locality
Lupaceradotus

| L. useviaensis

| RRBP 2004-06-30

|

|

| Lungfish

|

Ceratodontidae

| Indeterminate

|

| Mtuka Member

|

| Lungfish

|

=Crocodyliformes=

! Genus !! Species

!! Stratigraphic position !! Abundance !! Notes !! Images
class="wikitable" align="center"

! colspan="7" align="center" | Crocodyliformes of the Galula Formation

GenusSpecies Locality
Pakasuchus{{Cite journal|last1=O’Connor|first1=Patrick M.|last2=Sertich|first2=Joseph J. W.|last3=Stevens|first3=Nancy J.|last4=Roberts|first4=Eric M.|last5=Gottfried|first5=Michael D.|last6=Hieronymus|first6=Tobin L.|last7=Jinnah|first7=Zubair A.|last8=Ridgely|first8=Ryan|last9=Ngasala|first9=Sifa E.|date=August 2010|title=The evolution of mammal-like crocodyliforms in the Cretaceous Period of Gondwana|journal=Nature|volume=466|issue=7307|pages=748–751|doi=10.1038/nature09061|pmid=20686573|bibcode=2010Natur.466..748O |s2cid=4405106 |issn=0028-0836}}

| P. kapilimai

| RRBP 2007-04

| Namba Member

| [Two] skulls and an articulated skeleton.

| A notosuchian crocodyliform with mammal-like teeth.

| File:Pakasuchus.jpg

Rukwasuchus{{Cite journal|last1=Sertich|first1=Joseph J. W.|last2=O’Connor|first2=Patrick M.|date=2014-04-16|title=A new crocodyliform from the middle Cretaceous Galula Formation, southwestern Tanzania|journal=Journal of Vertebrate Paleontology|volume=34|issue=3|pages=576–596|doi=10.1080/02724634.2013.819808|bibcode=2014JVPal..34..576S |s2cid=16644660 |issn=0272-4634}}

| R. yajabalijekundu

| Namba 2

| Middle Namba member

| A partial skull and isolated teeth.

| A notosuchian crocodyliform, and the only known sub-Saharan peirosaurid from Africa.

|

Notosuchia

| Indeterminate

|

| Namba Member

|

|

|

=Dinosaurs=

! Genus !! Species

!! Stratigraphic position !! Abundance !! Notes !! Images
class="wikitable" align="center"

! colspan="7" align="center" | Dinosaurs of the Galula Formation

GenusSpecies Locality
Mnyamawamtuka{{cite journal|first1=Eric|last1=Gorscak|first2=Patrick M.|last2=O’Connor|year=2019|title=A new African titanosaurian sauropod dinosaur from the middle Cretaceous Galula Formation (Mtuka Member), Rukwa Rift Basin, southwestern Tanzania|journal=PLoS ONE|issue=14|volume=2|pages=e0211412|doi=10.1371/journal.pone.0211412|pmc=6374010|pmid=30759122|bibcode=2019PLoSO..1411412G |doi-access=free }}

| M. moyowamkia

|

| Mtuka Member

| An anterior cervical vertebral neural arch, cervical vertebral centra, partial dorsal vertebrae, sacral neural arch, partial sacral centra, sacral ribs, caudal vertebral neural arches, centra, chevrons, numerous dorsal rib fragments, scapula, sternal plate, humeri, partial ulna, metacarpal I, metacarpal III, partial ischium, partial pubis, partial femora, tibiae, fibula, metatarsal I, metatarsal II, metatarsal III, metatarsal IV, metatarsal V, pedal phalanges, ungual, and numerous unidentifiable fragments.

| A basal lithostrotian titanosaur with heart-shaped caudal vertebrae.

|

Rukwatitan{{Cite journal|last1=Gorscak|first1=Eric|last2=O'Connor|first2=Patrick M.|last3=Stevens|first3=Nancy J.|last4=Roberts|first4=Eric M.|date=2014-07-29|title=The basal titanosaurianRukwatitan bisepultus(Dinosauria, Sauropoda) from the middle Cretaceous Galula Formation, Rukwa Rift Basin, southwestern Tanzania|journal=Journal of Vertebrate Paleontology|volume=34|issue=5|pages=1133–1154|doi=10.1080/02724634.2014.845568|bibcode=2014JVPal..34.1133G |s2cid=677002 |issn=0272-4634}}

| R. bisepultus

| Locality RRBP 2007-02 (Namba 2)

| Middle Namba member

| Posterior cervical vertebrae, partial anterior dorsal vertebral neural arch, anterior caudal vertebrae, middle caudal vertebrae, chevrons, multiple partial dorsal ribs, distal scapula, partial coracoids, humerus, partial ulna, ilium, and proximal pubis.

| A basal lithostrotian titanosaur.

|

Shingopana{{Cite journal|last1=Gorscak|first1=Eric|last2=O'Connor|first2=Patrick M.|last3=Roberts|first3=Eric M.|last4=Stevens|first4=Nancy J.|date=2017-07-04|title=The second titanosaurian (Dinosauria: Sauropoda) from the middle Cretaceous Galula Formation, southwestern Tanzania, with remarks on African titanosaurian diversity|journal=Journal of Vertebrate Paleontology|volume=37|issue=4|pages=e1343250|doi=10.1080/02724634.2017.1343250|bibcode=2017JVPal..37E3250G |s2cid=90885040 |issn=0272-4634|url=https://figshare.com/articles/dataset/The_second_titanosaurian_Dinosauria_Sauropoda_from_the_middle_Cretaceous_Galula_Formation_southwestern_Tanzania_with_remarks_on_African_titanosaurian_diversity/5340808 |url-access=subscription}}

| S. songwensis

| TZ-07

| Namba Member

| An angular, partial anterior cervical vertebra, partial middle-to-posterior cervical vertebrae, partial cervical ribs, partial dorsal ribs, humerus, pubis, and many incomplete and/or unidentifiable fragments pertaining to the axial skeleton.

| A lithostrotian titanosaur.

|

Theropoda{{Cite journal|last1=O’Connor|first1=Patrick M.|last2=Gottfried|first2=Michael D.|last3=Stevens|first3=Nancy J.|last4=Roberts|first4=Eric M.|last5=Ngasala|first5=Sifa|last6=Kapilima|first6=Saidi|last7=Chami|first7=Remigius|date=March 2006|title=A new vertebrate fauna from the Cretaceous Red Sandstone Group, Rukwa Rift Basin, Southwestern Tanzania|journal=Journal of African Earth Sciences|volume=44|issue=3|pages=277–288|doi=10.1016/j.jafrearsci.2005.11.022|bibcode=2006JAfES..44..277O |issn=1464-343X}}

| Indeterminate

|

| Mtuka and Namba Members

| A pair of articulated proximal caudal vertebrae, and 11 teeth.

| An indeterminate theropod.

|

=Turtles=

! Genus !! Species

!! Stratigraphic position !! Abundance !! Notes !! Images
class="wikitable" align="center"

! colspan="7" align="center" | Turtles of the Galula Formation

GenusSpecies Locality
Testudines

| Indeterminate

|

| Mtuka and Namba Members.

| Plastron and carapace fragments.

|

|

= Mammals =

! Genus !! Species

!! Stratigraphic position !! Abundance !! Notes !! Images
class="wikitable" align="center"

! colspan="7" align="center" | Mammals of the Galula Formation

GenusSpecies Locality
Galulatherium{{Cite journal|last1=OConnor|first1=Patrick|last2=Krause|first2=David|last3=Stevens|first3=Nancy|last4=MacPhee|first4=Ross|last5=Groenke|first5=Joseph|last6=Kalthoff|first6=Daniela|date=2019|title=A new mammal from the Upper Cretaceous (Turonian–Campanian) Galula Formation, southwestern Tanzania|journal=Acta Palaeontologica Polonica|volume=64|doi=10.4202/app.00568.2018|doi-access=free}}

| G. jenkinsi

| TZ-07

| Namba Member

| Left dentary

| Probable Gondwanathere{{Cite journal|last1=KRAUSE|first1=DAVID W.|last2=GOTTFRIED|first2=MICHAEL D.|last3=O’CONNOR|first3=PATRICK M|last4=ROBERTS|first4=ERIC M.|date=2003|title=A Cretaceous mammal from Tanzania|url=https://www.app.pan.pl/archive/published/app48/app48-321.pdf|journal=Acta Palaeontologica Polonica|volume=48|issue=3|pages=321–330}}

| File:Galulatherium_photograph_lingual.jpg

References