Hadrosauroidea

{{Automatic taxobox

| name = Hadrosauroids

| fossil_range = Early-Late Cretaceous, {{fossilrange|130|66|earliest=130|PS=}}

| image = Probactrosaurus v3.jpg

| image_caption = Probactrosaurus

| taxon = Hadrosauroidea

| authority = Cope, 1869

| type_species = {{extinct}}Hadrosaurus foulkii

| type_species_authority = Leidy, 1858

| subdivision_ranks = Subgroups

| subdivision = *{{extinct}}Altirhinus

{{extinct}}Batyrosaurus
{{extinct}}Bolong?
{{extinct}}Choyrodon
{{extinct}}Datonglong
{{extinct}}Equijubus
{{extinct}}Gongpoquansaurus
{{extinct}}Hypsibema?
{{extinct}}Jinzhousaurus?
{{extinct}}Koshisaurus
{{extinct}}Levnesovia
{{extinct}}Mantellisaurus?
{{extinct}}Ouranosaurus?
{{extinct}}Penelopognathus
{{extinct}}Portellsaurus
{{extinct}}Probactrosaurus
{{extinct}}Qianjiangsaurus
{{extinct}}Siamodon
{{extinct}}Sirindhorna
{{extinct}}Xuwulong
{{extinct}}Zuoyunlong
{{extinct}}Hadrosauromorpha

| synonyms = * Trachodontoidea Lydekker, 1888

Hadrosauria Huene, 1956

}}

Hadrosauroidea is a clade or superfamily of ornithischian dinosaurs that includes the "duck-billed" dinosaurs, or Hadrosauridae, and all dinosaurs more closely related to them than to Iguanodon. Their remains have been recovered in Asia, Europe, Africa and the Americas. Many primitive hadrosauroids, such as the Asian Probactrosaurus and Altirhinus, have traditionally been included in a paraphyletic (unnatural grouping) "Iguanodontidae". With cladistic analysis, the traditional Iguanodontidae has been largely disbanded, and probably includes only Iguanodon and perhaps its closest relatives.

Classification

Hadrosauroidea was given a formal phylogenetic definition in the PhyloCode by Daniel Madzia and colleagues in 2021 as "the largest clade containing Hadrosaurus foulkii, but not Iguanodon bernissartensis".{{cite journal |last1=Madzia |first1=D. |last2=Arbour |first2=V.M. |last3=Boyd |first3=C.A. |last4=Farke |first4=A.A. |last5=Cruzado-Caballero |first5=P. |last6=Evans |first6=D.C. |title=The phylogenetic nomenclature of ornithischian dinosaurs |journal=PeerJ |date=2021 |volume=9 |pages=e12362 |doi=10.7717/peerj.12362|doi-access=free |pmid=34966571 |pmc=8667728 }} The cladogram below follows an analysis by Andrew McDonald, 2012, and shows the position of Hadrosauroidea within Styracosterna:{{Cite journal | last1 = McDonald | first1 = A. T. | title = Phylogeny of basal iguanodonts (Dinosauria: Ornithischia): an update | doi = 10.1371/journal.pone.0036745 | journal = PLOS ONE | volume = 7 | issue = 5 | page = e36745 | year = 2012 | pmid = 22629328| pmc =3358318 | doi-access = free | bibcode = 2012PLoSO...736745M }}

{{clade| style=font-size:100%;line-height:80%

|label1=Styracosterna

|1={{clade

|1=Uteodon aphanoecetes

|3={{clade

|1={{clade

|1=Hippodraco scutodens

|2=Theiophytalia kerri }}

|2={{clade

|1=Iguanacolossus fortis

|2={{clade

|1=Lanzhousaurus magnidens

|2={{clade

|1=Kukufeldia tilgatensis

|2={{clade

|1=Barilium dawsoni

|label2=Hadrosauriformes

|2={{clade

|1=Iguanodon bernissartensis

|label2=Hadrosauroidea

|2={{clade

|1=Mantellisaurus atherfieldensis

|2=Other hadrosauroids }} }} }} }} }} }} }} }} }}

The cladogram below follows an analysis by Wu Wenhao and Pascal Godefroit (2012):{{cite book |author1=Wu Wenhao |author2=Pascal Godefroit |name-list-style=amp |year=2012 |chapter=Anatomy and relationships of Bolong yixianensis, an Early Cretaceous iguanodontoid dinosaur from western Liaoning, China |editor=Godefroit, P. |title=Bernissart Dinosaurs and Early Cretaceous Terrestrial Ecosystems |url=https://archive.org/details/bernissartdinosa00gode |url-access=limited |publisher=Indiana University Press |pages=[https://archive.org/details/bernissartdinosa00gode/page/n309 293]–333}}

{{clade| style=font-size:90%;line-height:85%

|label1=Hadrosauriformes

|1={{clade

|label1=Iguanodontidae

|1={{clade

|1=Iguanodon

|2={{clade

|1=Ouranosaurus

|2=Mantellisaurus }} }}

|label2=Hadrosauroidea

|2={{clade

|1=Bolong

|2={{clade

|1=Equijubus

|2={{clade

|1=Jinzhousaurus

|2={{clade

|1=Altirhinus

|2={{clade

|1=Batyrosaurus

|2={{clade

|1=Probactrosaurus

|2={{clade

|1=Eolambia

|2={{clade

|1=Protohadros

|2={{clade

|1=Bactrosaurus

|2=Levnesovia

|3={{clade

|1=Shuangmiaosaurus

|2={{clade

|1=Tethyshadros

|2={{clade

|1=Telmatosaurus

|2=Hadrosauridae }} }} }} }} }} }} }} }} }} }} }} }} }} }}

Cladogram after Prieto-Marquez and Norell (2010):{{cite journal |author=Albert Prieto-Marquez |author2=Mark A. Norell |name-list-style=amp |year=2010 |title=Anatomy and relationships of Gilmoreosaurus mongoliensis (Dinosauria: Hadrosauroidea) from the Late Cretaceous of Central Asia |journal=American Museum Novitates |issue=3694 |pages=1–52 |url=https://digitallibrary.amnh.org/items/ad60ee42-08a4-47ce-9f49-e62ba5b8ea64 |doi=10.1206/3694.2 |s2cid=56372891 }}

{{clade| style=font-size:90%;line-height:85%

|label1=Hadrosauroidea

|1={{clade

|1=Jinzhousaurus

|2=Penelopognathus }}

|2={{clade

|1=Equijubus

|2={{clade

|1=Probactrosaurus

|2={{clade

|1={{clade

|1=Eolambia

|2=Protohadros }}

|2={{clade

|1=Tanius

|2={{clade

|1=Bactrosaurus

|2={{clade

|1=Gilmoreosaurus

|2={{clade

|1=Telmatosaurus

|2={{clade

|1=Shuangmiaosaurus

|2=Nanyangosaurus

|3=Lophorhothon

|4=Hadrosauridae }} }} }} }} }} }} }} }} }} }}

A phylogenetic analysis performed by Ramírez-Velasco et al. (2012) found a large polytomy between all hadrosauroids that are more derived than Probactrosaurus but less derived than Hadrosauridae. The exclusion of Claosaurus, Jeyawati, Levnesovia, Nanyangosaurus, Shuangmiaosaurus and Telmatosaurus from the analysis resulted in a more resolved topology:{{Cite journal|author1=Angel Alejandro Ramírez-Velasco |author2=Mouloud Benammi |author3=Albert Prieto-Márquez |author4=Jesús Alvarado Ortega |author5=René Hernández-Rivera |name-list-style=amp |year=2012 |title=Huehuecanauhtlus tiquichensis, a new hadrosauroid dinosaur (Ornithischia: Ornithopoda) from the Santonian (Late Cretaceous) of Michoacán, Mexico |journal=Canadian Journal of Earth Sciences |volume=49 |issue=2 |pages=379–395 |doi=10.1139/e11-062 |bibcode=2012CaJES..49..379R |url=http://www.nrcresearchpress.com/doi/abs/10.1139/e11-062 }}