Howesia

{{Short description|Extinct genus of reptiles}}

{{Automatic taxobox

| fossil_range = Middle Triassic, {{fossil_range|246}}

| image = Howesia browni.jpg

| image_caption =

| taxon = Howesia

| authority = Broom, 1905

| subdivision_ranks = Species

| subdivision =

}}

File:Howesia browni.png

Howesia is an extinct genus of basal rhynchosaur from early Middle Triassic (early Anisian stage) deposits of Eastern Cape, South Africa. It is known from the holotype SAM 5884, a partial skeleton with palate and partial lower jaws and from two paratypes, SAM 5885 and SAM 5886.{{cite journal |author=David M. Dilkes |year=1995 |title=The rhynchosaur Howesia browni from the Lower Triassic of South Africa |journal=Palaeontology |volume=38 |pages=665–685 }} It was found in the Burgersdorp Formation of the middle deposits of the Beaufort Group (Karoo Basin) and referred to Subzone B of the Cynognathus Assemblage Zone.{{Cite journal|author1=Richard J. Butler |author2=Martín D. Ezcurra |author3=Felipe C. Montefeltro |author4=Adun Samathi |author5=Gabriela Sobral |year=2015 |title=A new species of basal rhynchosaur (Diapsida: Archosauromorpha) from the early Middle Triassic of South Africa, and the early evolution of Rhynchosauria |journal=Zoological Journal of the Linnean Society |volume=174 |issue=3 |pages=571–588 |doi=10.1111/zoj.12246 |doi-access=free |hdl=11449/167867 |hdl-access=free }} It was first named by Robert Broom in 1905 and the type species is Howesia browni, named after Alfred Brown.{{cite journal |author=Robert Broom |year=1905 |title=Preliminary notice of some new fossil Reptiles collected by Mr. Alfred Brown at Aliwal North, S. Africa |journal=Records of the Albany Museum |volume=1 |issue=4 |pages=269–275 }}

Howesia in a cladogram based on Ezcurra et al. (2016):{{Cite journal|author1=Martin D. Ezcurra |author2=Felipe C. Montefeltro |author3=Richard J. Butler |year=2016 |url=http://pure-oai.bham.ac.uk/ws/files/25096585/fevo_03_00142.pdf| title= The Early Evolution of Rhynchosaurs |journal=Frontiers in Ecology and Evolution |volume=3 |pages=1–24|doi=10.3389/fevo.2015.00142|doi-access=free |archive-url=https://web.archive.org/web/20240220111216/https://pure-oai.bham.ac.uk/ws/files/25096585/fevo_03_00142.pdf |archive-date=2024-02-20 |url-status=live }}

{{clade| style=font-size:100%;line-height:85%

|label1=Rhynchosauria

|1={{clade

|1=Noteosuchus

|2=Mesosuchus

|3=Howesia

|4=Eohyosaurus

|label5=Rhynchosauridae

|5={{clade

|1=Rhynchosaurus

|2={{clade

|label1=Stenaulorhynchinae

|1={{clade

|1=Mariante rhynchosaur

|2=Stenaulorhynchus }}

|2={{clade

|1=Langeronyx

|2={{clade

|1=Bentonyx

|2={{clade

|1=Fodonyx

|label2=Hyperodapedontinae

|2={{clade

|1=Isalorhynchus

|2={{clade

|1=Teyumbaita

|2=Hyperodapedon }} }} }} }} }} }} }} }} }}

References