Khok Kruat Formation

{{short description|Geologic formation in Thailand}}

{{Infobox rockunit

| name = Khok Kruat Formation

| image =

| caption =

| type = Geological formation

| age = Aptian
~{{fossil range|120|113}}

| period = Aptian

| prilithology = Sandstone, siltstone

| otherlithology = Conglomerate, shale

| namedfor =

| namedby = Ward & Bunnag

| year_ts = 1964

| region = Indochina

| country = {{Flag|Thailand}}

| coordinates = {{coord|15.9|N|102.0|E|display=inline,title}}

| paleocoordinates = {{coord|16.3|N|105.3|E|display=inline}}

| unitof = Khorat Group

| subunits =

| underlies = Maha Sarakham Formation

| overlies = Phu Phan Formation

| thickness =

| extent = Khorat Plateau

| area =

| map = {{Location map+ | Thailand

| relief = 1

| width = 250

| float = center

| places =

{{Location map~ | Thailand

| lat_deg = 15.9

| lon_deg = 102.0

| mark = Green pog.svg

| marksize = 12

}}

}}

| map_caption =

}}

The Khok Kruat Formation ({{Langx|th|หมวดหินโคกกรวด}}) is a rock formation found in northeastern Thailand. It is the uppermost formation of the Khorat Group.{{cite book|author=Eric Buffetaut|title=Late Palaeozoic and Mesozoic Ecosystems in SE Asia|url=https://books.google.com/books?id=6wMfMgSmckkC&pg=PA146|year=2009|publisher=Geological Society of London|isbn=978-1-86239-275-5|page=146}} It is dated to the Aptian stage of the Early Cretaceous period, and is notable for its fossils of dinosaurs. It is equivalent to the Gres superieurs Formation of Laos.Weishampel, David B; et al. (2004). "Dinosaur distribution (Early Cretaceous, Asia)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 563-570. {{ISBN|0-520-24209-2}}. The group is a fluvial formation consisting primarily of red siltstones and sandstones.

Vertebrate paleofauna

{{Paleobiota-key-compact}}

The Khok Kruat Formation has yielded remains of sharks, fishes, turtles, crocodilians and pterosaurs. Phuwiangosaurus-like teeth, Siamosaurus teeth, and fragmentary postcranial remains of spinosaurids have also been recovered from it.{{Cite journal|author1=Eric Buffetaut |author2=Varavudh Suteethorn |author3=Jean Le Loeuff |author4=Sasidhorn Khansubha |author5=Haiyan Tong |author6=Kamonlak Wongko |year=2005 |title=New dinosaur species from the Upper Triassic Upper Maleri and Lower Dharmaram formations of central India |journal=International Conference on Geology, Geotechnology and Mineral Resources of Indochina (GEOINDO 2005) |pages=575–581 }} Fukuiraptor-like teeth have also been recovered from the formation.{{Cite journal|last1=Chokchaloemwong|first1=Duangsuda|last2=Azuma|first2=Yoichi|last3=Shibata|first3=Masateru|last4=Jintasakul|first4=Pratueng|date=2015|title=The Carcharodontosaurid Teeth from the Lower Cretaceous Khok Kruat Formation of Nakhon Ratchasima, Thailand|url=https://www.researchgate.net/publication/341040363|journal=2nd International Symposium on Asian Dinosaurs 2015 Bangkok}}

=Reptiles=

==Dinosaurs==

class="wikitable" align="center"
colspan="6" align="center" | Dinosaurs reported from the Khok Kruat Formation
GenusSpeciesLocationMaterialNotesImages
style="background:#FEF6E4;" |cf. Asianopodus

|style="background:#FEF6E4;" |cf. A. isp.

|style="background:#FEF6E4;" |"Huai Dam Chum site, Ban Lao Nat, Tha Uthen District, Nakhon Phanom Province".{{Cite journal |last1=Kozu |first1=Shohei |last2=Sardsud |first2=Apsorn |last3=Saesaengseerung |first3=Doungrutai |last4=Pothichaiya |first4=Cherdchan |last5=Agematsu |first5=Sachiko |last6=Sashida |first6=Katsuo |date=2017-11-01 |title=Dinosaur footprint assemblage from the Lower Cretaceous Khok Kruat Formation, Khorat Group, northeastern Thailand |journal=Geoscience Frontiers |language=en |volume=8 |issue=6 |pages=1479–1493 |doi=10.1016/j.gsf.2017.02.003 |bibcode=2017GeoFr...8.1479K |issn=1674-9871|doi-access=free |url=https://tsukuba.repo.nii.ac.jp/record/43785/files/GF_8-1479.pdf }}

|style="background:#FEF6E4;" |"At least 79 trackways composed of 341 consecutive tracks in total".

|style="background:#FEF6E4;" |Theropod tracks.

|style="background:#FEF6E4;" |

style="background:#FEF6E4;" |Ichnogen. et. sp. indet.

|style="background:#FEF6E4;" |

|style="background:#FEF6E4;" |"Huai Dam Chum site, Ban Lao Nat, Tha Uthen District, Nakhon Phanom Province".

|style="background:#FEF6E4;" |"Two trackways: T23 composed of 6 consecutive tracks and T35 composed of 3 discontinuous tracks".

|style="background:#FEF6E4;" |Flattened possible theropod tracks.

|style="background:#FEF6E4;" |

Ornithomimosauria{{cite journal|author1=Chokchaloemwong, D.|author2=Hattori, S.|author3=Yukawa, H.|author4=Shibata, M.|author5=Naksri, W.|year=2024|title=A new ornithomimosaurian material from the Khok Kruat Formation, Lower Cretaceous of Thailand|journal=Memoir of the Fukui Prefectural Dinosaur Museum|volume=23|pages=1–7|url=https://www.dinosaur.pref.fukui.jp/archive/memoir/memoir023-001.pdf}}

| Indeterminate

| Ban Krok Duean Ha

| "NRRU-F01020052, distal part of the right metatarsal III"

| An ornithomimosaurian

|

Psittacosaurus

| P. sattayarakiBuffetaut, Eric & Suteethorn, Varavudh. (1992). A new species of the ornithischian dinosaur Psittacosaurus from the Early Cretaceous of Thailand. Palaeontology 35: 801-812.

| Ban Dong Bang Noi

| "Well preserved left dentary and an incomplete maxilla."

| A ceratopsian

|

Ratchasimasaurus

| R. suranareae

|Nakhon Ratchasima province.{{cite journal |first1=Masateru |last1=Shibata |first2=Pratueng |last2=Jintasakul |first3=Yoichi |last3=Azuma |year=2011 |title=A New Iguanodontian Dinosaur from the Lower Cretaceous Khok Kruat Formation, Nakhon Ratchasima in Northeastern Thailand |journal=Acta Geologica Sinica |language=en |volume=85 |issue=5 |pages=969–976 |doi=10.1111/j.1755-6724.2011.00230.x}}

|

| An iguanodontian ornithopod

|File:Ratchasimasaurus suranareae 02.jpg

Siamodon

| S. nimngami{{cite journal |first1=Eric |last1=Buffetaut |first2=Varavudh |last2=Suteethorn |year=2011 |title=A new iguanodontian dinosaur from the Khok Kruat Formation (Early Cretaceous, Aptian) of northeastern Thailand |journal=Annales de Paléontologie |volume=97 |issue=1–2 |pages=51–62 |doi=10.1016/j.annpal.2011.08.001 |bibcode=2011AnPal..97...51B }}

| Ban Saphan Hin

| PRC-4, a well-preserved left maxilla, PRC-5, an isolated maxillary tooth and PRC-6, a braincase.

| An iguanodontian ornithopod

|File:Left Siamodon nimngami maxillary labial view.jpg

Sirindhorna

| S. khoratensis

|Muaeng Nakhon Ratchasima District, Nakhon Ratchasima Province.{{Cite journal |last1=Shibata |first1=Masateru |last2=Jintasakul |first2=Pratueng |last3=Azuma |first3=Yoichi |last4=You |first4=Hai-Lu |date=2015-12-30 |title=A New Basal Hadrosauroid Dinosaur from the Lower Cretaceous Khok Kruat Formation in Nakhon Ratchasima Province, Northeastern Thailand |journal=PLOS ONE |volume=10 |issue=12 |pages=e0145904 |doi=10.1371/journal.pone.0145904 |issn=1932-6203 |pmc=4696827 |pmid=26716981|bibcode=2015PLoSO..1045904S |doi-access=free }}

|

| An iguanodontian ornithopod

|File:Sirindhorna skull and head.PNG

Siamosaurus

| S. sp.{{Cite journal|last1=Milner|first1=Angela|last2=Buffetaut|first2=Eric|last3=Suteethorn|first3=Varavudh|date=2007|title=A tall-spined spinosaurid theropod from Thailand and the biogeography of spinosaurs|url=http://vertpaleo.org/Annual-Meeting/Future-Past-Meetings/MeetingPdfs/JVP_07.aspx|journal=Journal of Vertebrate Paleontology|volume=27 (supplement 3)|pages=118A|access-date=2020-06-08|archive-date=2019-09-23|archive-url=https://web.archive.org/web/20190923012213/http://vertpaleo.org/Annual-Meeting/Future-Past-Meetings/MeetingPdfs/JVP_07.aspx|url-status=dead}}

|

| Teeth

| A spinosaurid

|File:Siamosaurus suteethorni by PaleoGeek.png

Siamraptor

| S. suwati

|Muaeng Nakhon Ratchasima District, Nakhon Ratchasima Province.{{Cite journal |last1=Chokchaloemwong |first1=Duangsuda |last2=Hattori |first2=Soki |last3=Cuesta |first3=Elena |last4=Jintasakul |first4=Pratueng |last5=Shibata |first5=Masateru |last6=Azuma |first6=Yoichi |date=2019-10-09 |title=A new carcharodontosaurian theropod (Dinosauria: Saurischia) from the Lower Cretaceous of Thailand |journal=PLOS ONE |language=en |volume=14 |issue=10 |pages=e0222489 |doi=10.1371/journal.pone.0222489 |issn=1932-6203 |pmc=6784982 |pmid=31596853|bibcode=2019PLoSO..1422489C |doi-access=free }}

| "A composite cranial and postcranial skeleton comprising premaxilla, maxilla, jugal, surangular, prearticular, articular, vertebrae, manual ungual, ischium, tibia, and pedal phalanx".

| A carcharodontosaur

|File:Siamraptor reconstruction 2019 (Mario Lanzas).jpg

==Pseudosuchians==

class="wikitable" align="center"
colspan="6" align="center" | Pseudosuchians reported from the Khok Kruat Formation
GenusSpeciesLocation

!Material

NotesImages
Atoposauridae indet.

|Indeterminate

|Lam Pao Reservoir, Ban Noi, Sahatsakhan district, Kalasin province.{{Cite journal |last1=Lauprasert |first1=Komsorn |last2=Laojumpon |first2=Chalida |last3=Saenphala |first3=Wanitchaphat |last4=Cuny |first4=Gilles |last5=Thirakhupt |first5=Kumthorn |last6=Suteethorn |first6=Varavudh |date=2010-06-12 |title=Atoposaurid crocodyliforms from the Khorat Group of Thailand: first record of Theriosuchus from Southeast Asia |url=https://www.researchgate.net/publication/215486888 |journal=Paläontologische Zeitschrift |volume=85 |issue=1 |pages=37–47 |doi=10.1007/s12542-010-0071-z |s2cid=128623617 |issn=0031-0220}}

|An isolated lanceolate tooth (PRCMR 282).

|May be attributable to Theriosuchus.

|

Khoratosuchus

|K. jintasakuli

| Ban Saphan Hin, Nakhon Ratchasima province.{{Cite journal |last1=Lauprasert |first1=K. |last2=Cuny |first2=G. |last3=Thirakhupt |first3=K. |last4=Suteethorn |first4=V. |date=May 2009 |title=Khoratosuchus jintasakuli gen. et sp. nov., an advanced neosuchian crocodyliform from the Early Cretaceous (Aptian-Albian) of NE Thailand |url=https://www.researchgate.net/publication/215486865 |journal=Geological Society, London, Special Publications |volume=315 |issue=1 |pages=175–187 |doi=10.1144/sp315.13 |bibcode=2009GSLSP.315..175L |s2cid=131412579 |issn=0305-8719}}

|A partial skull (NNRU-A 1803).

|An advanced neosuchian.

|

==Turtles==

class="wikitable" align="center"
colspan="6" align="center" | Turtles reported from the Khok Kruat Formation
GenusSpeciesLocation

!Material

NotesImages
Kizylkumemys

|K. khoratensis

| Ban Saphan Hin and Ban Khok Kruat localities, Nakhon Ratchasima province.{{Cite journal |last1=Tong |first1=Haiyan |last2=Claude |first2=Julien |last3=Suteethorn |first3=Varavudh |last4=Naksri |first4=Wilailuck |last5=Buffetaut |first5=Eric |date=May 2009 |title=Turtle assemblages of the Khorat Group (Late Jurassic-Early Cretaceous) of NE Thailand and their palaeobiogeographical significance |url=https://www.researchgate.net/publication/240675681 |journal=Geological Society, London, Special Publications |volume=315 |issue=1 |pages=141–152 |doi=10.1144/sp315.11 |bibcode=2009GSLSP.315..141T |s2cid=130709324 |issn=0305-8719}}

|Shell material.

| A carettochelyid.

|

Shachemys

|S. sp.

|Ban Saphan Hin, Nakhon Ratchasima province.

|Shell material.

|An adocid, remains too fragmentary for specific assignment.

|

=Fish=

==Bony fish==

class="wikitable" align="center"
colspan="6" align="center" | Bony fish reported from the Khok Kruat Formation
GenusSpeciesLocation

!Material

NotesImages
Khoratamia

|K. phattharajani

|Ban Krok Duean Ha, Nakhon Ratchasima province.{{Cite journal |last1=Deesri |first1=Uthumporn |last2=Naksri |first2=Wilailuck |last3=Jintasakul |first3=Pratueng |last4=Noda |first4=Yoshikazu |last5=Yukawa |first5=Hirokazu |last6=Hossny |first6=Tamara El |last7=Cavin |first7=Lionel |date=April 2023 |title=A New Sinamiin Fish (Actinopterygii) from the Early Cretaceous of Thailand: Implications on the Evolutionary History of the Amiid Lineage |journal=Diversity |language=en |volume=15 |issue=4 |pages=491 |doi=10.3390/d15040491 |issn=1424-2818 |doi-access=free }}

|Holotype (NRRU-F01020023), paratype (NRRU-F01020024) and a patch of scales (NRRU-F01020025).

|An amiid.

|File:Holotype of Khoratamia phattharajani.png

Lanxangichthys

|L. sp.

| Khok Pha Suam, Ubon Ratchathani province.{{Cite journal |last1=Cavin |first1=Lionel |last2=Deesri |first2=Uthumporn |last3=Veran |first3=Monette |last4=Khentavong |first4=Bounsou |last5=Jintasakul |first5=Pratueng |last6=Chanthasit |first6=Phornphen |last7=Allain |first7=Ronan |date=2019-03-04 |title=A new Lepisosteiformes (Actinopterygii: Ginglymodi) from the Early Cretaceous of Laos and Thailand, SE Asia |url=https://figshare.com/articles/journal_contribution/A_new_Lepisosteiformes_Actinopterygii_Ginglymodi_from_the_Early_Cretaceous_of_Laos_and_Thailand_SE_Asia/5872695/1/files/10438053.pdf |journal=Journal of Systematic Palaeontology |volume=17 |issue=5 |pages=393–407 |doi=10.1080/14772019.2018.1426060 |bibcode=2019JSPal..17..393C |s2cid=90886053 |issn=1477-2019}}

|

| A lepisosteiform.

|

==Cartilaginous fish==

class="wikitable" align="center"
colspan="6" align="center" | Cartilaginous fish reported from the Khok Kruat Formation
GenusSpeciesLocation

!Material

NotesImages
Acrorhizodus

|A. khoratensis

|Khok Pha Suam locality, Ubon Ratchathani province.{{Cite journal |last1=Cappetta |first1=Henri |last2=Buffetaut |first2=Eric |last3=Cuny |first3=Gilles |last4=Suteethorn |first4=Varavudh |date=May 2006 |title=A New Elasmobranch Assemblage From The Lower Cretaceous of Thailand |journal=Palaeontology |language=en |volume=49 |issue=3 |pages=547–555 |doi=10.1111/j.1475-4983.2006.00555.x |bibcode=2006Palgy..49..547C |s2cid=129380407 |issn=0031-0239|doi-access=free }}

|5 teeth, most of them incomplete.

|A hybodontiform.

|

Heteroptychodus

|H. aff. steinmanni

|Khok Pha Suam locality, Ubon Ratchathani province.

|20 teeth, most incomplete.

|A ptychodontid.

|

Hybodus?

|H.? sp.

|Khok Pha Suam locality, Ubon Ratchathani province.

|22 incomplete teeth.

|Possibly represents a new genus, but the available material is too fragmentary to warrant erecting a new taxon.

|

Thaiodus

|T. ruchae

| Khok Pha Suam locality, Ubon Ratchathani province.

|One incomplete tooth.

| A hybodontid.

|

See also

References