LYN

{{Short description|Mammalian protein found in Homo sapiens}}

{{other uses|Lyn (disambiguation)}}

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{{Infobox gene}}

Tyrosine-protein kinase Lyn is a protein that in humans is encoded by the LYN gene.{{cite journal | vauthors = Yamanashi Y, Fukushige S, Semba K, Sukegawa J, Miyajima N, Matsubara K, Yamamoto T, Toyoshima K | title = The yes-related cellular gene lyn encodes a possible tyrosine kinase similar to p56lck | journal = Molecular and Cellular Biology | volume = 7 | issue = 1 | pages = 237–243 | date = Jan 1987 | pmid = 3561390 | pmc = 365062 | doi = 10.1128/MCB.7.1.237 }}

Lyn is a member of the Src family of protein tyrosine kinases, which is mainly expressed in hematopoietic cells, in neural tissues liver, and adipose tissue. In various hematopoietic cells, Lyn has emerged as a key enzyme involved in the regulation of cell activation. In these cells, a small amount of LYN is associated with cell surface receptor proteins, including the B cell antigen receptor (BCR), CD40, or CD19. The abbreviation Lyn is derived from Lck/Yes novel tyrosine kinase, Lck and Yes also being members of the Src kinase family.

Function

Lyn has been described to have an inhibitory role in myeloid lineage proliferation. Following engagement of the B cell receptors, Lyn undergoes rapid phosphorylation and activation. This activation initiates a cascade of signaling events mediated by Lyn phosphorylation of tyrosine residues within the immunoreceptor tyrosine-based activation motifs (ITAMs) of receptor proteins. This cascade leads to the recruitment and activation of other kinases, including Syk, phospholipase Cγ2 (PLCγ2), and phosphatidyl inositol-3 kinase. These kinases generate activation signals critical for proliferation, Ca2+ mobilization, and cell differentiation.

Lyn also plays an essential role in transmitting inhibitory signals by phosphorylating tyrosine residues within the immunoreceptor tyrosine-based inhibitory motifs (ITIMs) of regulatory proteins such as CD22, PIR-B, and FCγRIIb1. ITIM phosphorylation subsequently recruits and activates phosphatases including SHIP-1 and SHP-1, leading to the attenuation of signaling pathways, downregulation of cell activation, and promotion of tolerance. In B cells, Lyn sets the threshold of signaling and maintains the balance between activation and inhibition, effectively functioning as a rheostat rather than a binary switch.

LYN is reported to be a key mediator of estrogen-dependent suppression of human osteoclast differentiation, survival, and function. It has also been implicated in the insulin signaling pathway, where activated Lyn phosphorylates insulin receptor substrate 1 (IRS1), promoting Glut-4 translocation to the membrane and enhancing glucose utilization. Insulin receptor activation has been shown to increase Lyn autophosphorylation, suggesting a feedback loop.

Lyn has been shown to protect against hepatocellular apoptosis and promote liver regeneration by preserving mitochondrial integrity.

In pulmonary function, Lyn activation in pulmonary epithelium has been linked to improved barrier integrity and reduced edema. Lyn activation in alveolar phagocytes enhances bacterial phagocytosis and reduces pulmonary infections. Furthermore, Lyn activation has been shown to reduce pulmonary mucus hypersecretion.

Clinical significance

= As a drug target =

HSP90 inhibitor NVP-BEP800 has been reported to affect Lyn kinase stability and inhibit the growth of B-cell acute lymphoblastic leukemias by interfering with NF-kappaB signaling.

The allosteric activator of Lyn kinase Tolimidone (MLR-1023) is currently under Phase 2a clinical investigation for Type II diabetes, with promising results reported from studies conducted by Melior Discovery.

The insulin secretagogue glimepiride (Amaryl®) activates Lyn in adipocytes by disrupting lipid rafts, potentially contributing to its extrapancreatic glycemic control effects. Tolimidone (MLR-1023), a small-molecule allosteric activator of Lyn kinase with an EC50 of 63 nM, is under Phase 2a investigation for Type II diabetes.

= Pathology =

Much of the current knowledge about Lyn has emerged from studies of genetically manipulated mice. Lyn deficient mice display a phenotype that includes splenomegaly, a dramatic increase in numbers of myeloid progenitors and monocyte/macrophage tumors. Biochemical analysis of cells from these mutants revealed that Lyn is essential in establishing ITIM-dependent inhibitory signaling and for activation of specific protein tyrosine phosphatases within myeloid cells.

Mice that expressed a hyperactive Lyn allele were tumor free and displayed no propensity toward hematological malignancy. These mice have reduced numbers of conventional B lymphocytes, down-regulated surface immunoglobulin M and costimulatory molecules, and elevated numbers of B1a B cells. With age these animals developed a glomerulonephritis phenotype associated with a 30% reduction in life expectancy.

Interactions

LYN has been shown to interact with:

{{div col|colwidth=20em}}

  • BCAR1,
  • CD117,
  • CD22,
  • Cdk1,
  • DOK1,
  • EPOR
  • GPVI,
  • INPP5D,
  • IRS1,
  • LCP2,
  • MUC1,
  • NEDD9,
  • PLCG2,{{cite journal | vauthors = Pleiman CM, Clark MR, Gauen LK, Winitz S, Coggeshall KM, Johnson GL, Shaw AS, Cambier JC | title = Mapping of sites on the Src family protein tyrosine kinases p55blk, p59fyn, and p56lyn which interact with the effector molecules phospholipase C-gamma 2, microtubule-associated protein kinase, GTPase-activating protein, and phosphatidylinositol 3-kinase | journal = Molecular and Cellular Biology | volume = 13 | issue = 9 | pages = 5877–5887 | date = Sep 1993 | pmid = 8395016 | pmc = 360336 | doi = 10.1128/MCB.13.9.5877 }}
  • PPP1R15A,
  • PTPRC,
  • Syk,
  • TRPV4,
  • UNC119,

{{Div col end}}

See also

References

{{reflist|35em|refs=

{{cite journal | vauthors = Pleiman CM, Clark MR, Gauen LK, Winitz S, Coggeshall KM, Johnson GL, Shaw AS, Cambier JC | title = Mapping of sites on the Src family protein tyrosine kinases p55blk, p59fyn, and p56lyn which interact with the effector molecules phospholipase C-gamma 2, microtubule-associated protein kinase, GTPase-activating protein, and phosphatidylinositol 3-kinase | journal = Molecular and Cellular Biology | volume = 13 | issue = 9 | pages = 5877–5887 | date = Sep 1993 | pmid = 8395016 | pmc = 360336 | doi = 10.1128/MCB.13.9.5877 }}

{{cite journal | vauthors = Anderwald C, Müller G, Koca G, Fürnsinn C, Waldhäusl W, Roden M | title = Short-term leptin-dependent inhibition of hepatic gluconeogenesis is mediated by insulin receptor substrate-2 | journal = Molecular Endocrinology | location = Baltimore, Md. | volume = 16 | issue = 7 | pages = 1612–1628 | date = Jul 2002 | pmid = 12089355 | doi = 10.1210/mend.16.7.0867 | doi-access = free }}

{{cite journal | vauthors = Baran CP, Tridandapani S, Helgason CD, Humphries RK, Krystal G, Marsh CB | title = The inositol 5'-phosphatase SHIP-1 and the Src kinase Lyn negatively regulate macrophage colony-stimulating factor-induced Akt activity | journal = The Journal of Biological Chemistry | volume = 278 | issue = 40 | pages = 38628–38636 | date = Oct 2003 | pmid = 12882960 | doi = 10.1074/jbc.M305021200 | doi-access = free }}

{{cite journal | vauthors = Brown VK, Ogle EW, Burkhardt AL, Rowley RB, Bolen JB, Justement LB | title = Multiple components of the B cell antigen receptor complex associate with the protein tyrosine phosphatase, CD45 | journal = The Journal of Biological Chemistry | volume = 269 | issue = 25 | pages = 17238–17244 | date = Jun 1994 | pmid = 7516335 | doi = 10.1016/S0021-9258(17)32545-0 | doi-access = free }}

{{cite journal | vauthors = Campbell MA, Sefton BM | title = Association between B-lymphocyte membrane immunoglobulin and multiple members of the Src family of protein tyrosine kinases | journal = Molecular and Cellular Biology | volume = 12 | issue = 5 | pages = 2315–2321 | date = May 1992 | pmid = 1569953 | pmc = 364403 | doi = 10.1128/MCB.12.5.2315 }}

{{cite journal | vauthors = Campbell KS | title = Signal transduction from the B cell antigen-receptor | journal = Current Opinion in Immunology | volume = 11 | issue = 3 | pages = 256–64 | date = June 1999 | pmid = 10375554 | doi = 10.1016/s0952-7915(99)80042-9 | url = }}

{{cite journal | vauthors = Cen O, Gorska MM, Stafford SJ, Sur S, Alam R | title = Identification of UNC119 as a novel activator of SRC-type tyrosine kinases | journal = The Journal of Biological Chemistry | volume = 278 | issue = 10 | pages = 8837–8845 | date = Mar 2003 | pmid = 12496276 | doi = 10.1074/jbc.M208261200 | doi-access = free }}

{{cite journal | vauthors = Chan VW, Lowell CA, DeFranco AL | title = Defective negative regulation of antigen receptor signaling in Lyn-deficient B lymphocytes | journal = Current Biology | volume = 8 | issue = 10 | pages = 545–553 | date = May 1998 | pmid = 9601638 | doi = 10.1016/S0960-9822(98)70223-4 | s2cid = 12195731 | doi-access = free | bibcode = 1998CBio....8..545C }}

{{cite journal | vauthors = Chin H, Arai A, Wakao H, Kamiyama R, Miyasaka N, Miura O | title = Lyn physically associates with the erythropoietin receptor and may play a role in activation of the Stat5 pathway | journal = Blood | volume = 91 | issue = 10 | pages = 3734–3745 | date = May 1998 | pmid = 9573010 | doi = 10.1182/blood.V91.10.3734 | doi-access = free }}

{{cite journal | vauthors = Corey SJ, Day RM | title = Shockingly: the loss of Lyn leads to leakiness | journal = Blood | volume = 122 | issue = 25 | pages = 4009–4010 | date = Dec 2013 | pmid = 24335031 | pmc = 3862277 | doi = 10.1182/blood-2013-10-533158 }}

{{cite journal | vauthors = Cornall RJ, Cyster JG, Hibbs ML, Dunn AR, Otipoby KL, Clark EA, Goodnow CC | title = Polygenic autoimmune traits: Lyn, CD22, and SHP-1 are limiting elements of a biochemical pathway regulating BCR signaling and selection | journal = Immunity | volume = 8 | issue = 4 | pages = 497–508 | date = Apr 1998 | pmid = 9586639 | doi = 10.1016/S1074-7613(00)80554-3 | doi-access = free }}

{{cite journal | vauthors = Durum SK, Aiello FB | title = Interleukin-7 induces MUC1 | journal = Cancer Biology & Therapy | volume = 2 | issue = 2 | pages = 194–195 | year = 2003 | pmid = 12750562 | doi = 10.4161/cbt.2.2.351 | doi-access = free }}

{{cite journal | vauthors = Gavali S, Gupta MK, Daswani B, Wani MR, Sirdeshmukh R, Khatkhatay MI | title = LYN, a key mediator in estrogen-dependent suppression of osteoclast differentiation, survival, and function | journal = Biochimica et Biophysica Acta. Molecular Basis of Disease | volume = 1865 | issue = 3 | pages = 547–557 | date = March 2019 | pmid = 30579930 | doi = 10.1016/j.bbadis.2018.12.016 | doi-access = free }}

{{cite journal | vauthors = Greer SF, Justement LB | title = CD45 regulates tyrosine phosphorylation of CD22 and its association with the protein tyrosine phosphatase SHP-1 | journal = Journal of Immunology | location = Baltimore, Md. | volume = 162 | issue = 9 | pages = 5278–5286 | date = May 1999 | pmid = 10228003 | doi = 10.4049/jimmunol.162.9.5278 | doi-access = free }}

{{cite journal | vauthors = Gringeri E, Carraro A, Tibaldi E, D'Amico FE, Mancon M, Toninello A, Pagano MA, Vio C, Cillo U, Brunati AM | title = Lyn-mediated mitochondrial tyrosine phosphorylation is required to preserve mitochondrial integrity in early liver regeneration | journal = The Biochemical Journal | volume = 425 | issue = 2 | pages = 401–412 | date = December 2009 | pmid = 19832701 | doi = 10.1042/BJ20090902 | url = https://hal.archives-ouvertes.fr/hal-00479212/file/PEER_stage2_10.1042%252FBJ20090902.pdf }}

{{cite journal | vauthors = Grishin AV, Azhipa O, Semenov I, Corey SJ | title = Interaction between growth arrest-DNA damage protein 34 and Src kinase Lyn negatively regulates genotoxic apoptosis | journal = Proceedings of the National Academy of Sciences of the United States of America | volume = 98 | issue = 18 | pages = 10172–10177 | date = Aug 2001 | pmid = 11517336 | pmc = 56934 | doi = 10.1073/pnas.191130798 | bibcode = 2001PNAS...9810172G | doi-access = free }}

{{cite journal | vauthors = Gross BS, Lee JR, Clements JL, Turner M, Tybulewicz VL, Findell PR, Koretzky GA, Watson SP | title = Tyrosine phosphorylation of SLP-76 is downstream of Syk following stimulation of the collagen receptor in platelets | journal = The Journal of Biological Chemistry | volume = 274 | issue = 9 | pages = 5963–5971 | date = Feb 1999 | pmid = 10026222 | doi = 10.1074/jbc.274.9.5963 | doi-access = free }}

{{cite journal | vauthors = Guo B, Kato RM, Garcia-Lloret M, Wahl MI, Rawlings DJ | title = Engagement of the human pre-B cell receptor generates a lipid raft-dependent calcium signaling complex | journal = Immunity | volume = 13 | issue = 2 | pages = 243–253 | date = Aug 2000 | pmid = 10981967 | doi = 10.1016/s1074-7613(00)00024-8 | doi-access = free }}

{{cite journal | vauthors = Harder KW, Parsons LM, Armes J, Evans N, Kountouri N, Clark R, Quilici C, Grail D, Hodgson GS, Dunn AR, Hibbs ML | title = Gain- and loss-of-function Lyn mutant mice define a critical inhibitory role for Lyn in the myeloid lineage | journal = Immunity | volume = 15 | issue = 4 | pages = 603–615 | date = Oct 2001 | pmid = 11672542 | doi = 10.1016/S1074-7613(01)00208-4 | doi-access = free }}

{{cite journal | vauthors = Hibbs ML, Harder KW, Armes J, Kountouri N, Quilici C, Casagranda F, Dunn AR, Tarlinton DM | title = Sustained activation of Lyn tyrosine kinase in vivo leads to autoimmunity | journal = The Journal of Experimental Medicine | volume = 196 | issue = 12 | pages = 1593–1604 | date = Dec 2002 | pmid = 12486102 | pmc = 2196073 | doi = 10.1084/jem.20020515 }}

{{cite journal | vauthors = Jan J, Zhang G, Welch EJ, Liang Y, Fu J, Vogel SM, Lowell CA, Du X, Cheresh DA, Malik AB, Li Z | title = A critical role for Lyn kinase in strengthening endothelial integrity and barrier function | journal = Blood | volume = 122 | issue = 25 | pages = 4140–4149 | date = Dec 2013 | pmid = 24108461 | pmc = 3862279 | doi = 10.1182/blood-2013-03-491423 | doi-access = free }}

{{cite journal | vauthors = Kharbanda S, Yuan ZM, Rubin E, Weichselbaum R, Kufe D | title = Activation of Src-like p56/p53lyn tyrosine kinase by ionizing radiation | journal = The Journal of Biological Chemistry | volume = 269 | issue = 32 | pages = 20739–20743 | date = Aug 1994 | pmid = 8051175 | doi = 10.1016/S0021-9258(17)32054-9 | doi-access = free }}

{{cite journal | vauthors = Lee MK, Kim SG, Watkins E, Moon MK, Rhee SY, Frias JP, Chung CH, Lee SH, Block B, Cha BS, Park HK, Kim BJ, Greenway F | title = A Novel Non-PPARgamma Insulin Sensitizer: MLR-1023 Clinical Proof-of-concept in Type 2 Diabetes Mellitus | journal = Journal of Diabetes and Its Complications | volume = 34 | issue = 5 | pages = 107555 | date = May 2020 | pmid = 32019723 | doi = 10.1185/030079907x188152 | s2cid = 211036334 }}

{{cite journal | vauthors = Li X, Zhou X, Ye Y, Li Y, Li J, Privratsky B, Wu E, Gao H, Huang C, Wu M | title = Lyn regulates inflammatory responses in Klebsiella pneumoniae infection via the p38/NF-κB pathway | journal = European Journal of Immunology | volume = 44 | issue = 3 | pages = 763–773 | date = 2012 | pmid = 24338528 | pmc = 4103995 | doi = 10.1002/eji.201343972 }}

{{cite journal | vauthors = Li X, He S, Zhou X, Ye Y, Tan S, Zhang S, Li R, Yu M, Jundt MC, Hidebrand A, Wang Y, Li G, Huang C, Wu M | title = Lyn delivers bacteria to lysosomes for eradication through TLR2-initiated autophagy related phagocytosis | journal = PLOS Pathogens | volume = 12 | issue = 1 | pages = e1005363 | date = Jan 2016 | pmid = 26735693 | pmc = 4703367 | doi = 10.1371/journal.ppat.1005363 | doi-access = free }}

{{cite journal | vauthors = Liang X, Wisniewski D, Strife A, Clarkson B, Resh MD | title = Phosphatidylinositol 3-kinase and Src family kinases are required for phosphorylation and membrane recruitment of Dok-1 in c-Kit signaling | journal = The Journal of Biological Chemistry | volume = 277 | issue = 16 | pages = 13732–13738 | date = Apr 2002 | pmid = 11825908 | doi = 10.1074/jbc.M200277200 | doi-access = free }}

{{cite journal | vauthors = Linnekin D, DeBerry CS, Mou S | title = Lyn associates with the juxtamembrane region of c-Kit and is activated by stem cell factor in hematopoietic cell lines and normal progenitor cells | journal = The Journal of Biological Chemistry | volume = 272 | issue = 43 | pages = 27450–27455 | date = Oct 1997 | pmid = 9341198 | doi = 10.1074/jbc.272.43.27450 | doi-access = free }}

{{cite journal | vauthors = Lowell CA | title = Src-family kinases: rheostats of immune cell signaling | journal = Molecular Immunology | volume = 41 | issue = 6–7 | pages = 631–643 | date = Jul 2004 | pmid = 15220000 | doi = 10.1016/j.molimm.2004.04.010 }}

{{cite journal | vauthors = Maeda A, Scharenberg AM, Tsukada S, Bolen JB, Kinet JP, Kurosaki T | title = Paired immunoglobulin-like receptor B (PIR-B) inhibits BCR-induced activation of Syk and Btk by SHP-1 | journal = Oncogene | volume = 18 | issue = 14 | pages = 2291–2297 | date = Apr 1999 | pmid = 10327049 | doi = 10.1038/sj.onc.1202552 | doi-access = free }}

{{cite journal | vauthors = Manié SN, Beck AR, Astier A, Law SF, Canty T, Hirai H, Druker BJ, Avraham H, Haghayeghi N, Sattler M, Salgia R, Griffin JD, Golemis EA, Freedman AS | title = Involvement of p130(Cas) and p105(HEF1), a novel Cas-like docking protein, in a cytoskeleton-dependent signaling pathway initiated by ligation of integrin or antigen receptor on human B cells | journal = The Journal of Biological Chemistry | volume = 272 | issue = 7 | pages = 4230–4236 | date = Feb 1997 | pmid = 9020138 | doi = 10.1074/jbc.272.7.4230 | doi-access = free | hdl = 20.500.12613/9177 | hdl-access = free }}

{{cite web | title = Melior Pharmaceuticals Initiates Phase 2 Study with MLR-1023 for Type 2 Diabetes | date = 3 March 2015 | url = http://www.businesswire.com/news/home/20150303005277/en/Melior-Pharmaceuticals-Initiates-Phase-2-Study-MLR-1023#.VVy7eGCKKkJ | publisher = Business Wire | access-date = March 3, 2015 }}

{{Cite web | title = Melior Pharmaceuticals Announces Positive Phase 2A Results in Type 2 Diabetes Study | date = 13 June 2016 | url = http://www.businesswire.com/news/home/20160613005028/en/Melior-Pharmaceuticals-Announces-Positive-Phase-2A-Results | website = www.businesswire.com | language = en | access-date = 2017-08-04 }}

{{Cite web | title = Melior Discovery website press releases | url = http://www.meliordiscovery.com/media_press.html | access-date = 2018-06-11 | archive-date = 2018-05-17 | archive-url = https://web.archive.org/web/20180517032422/http://www.meliordiscovery.com/media_press.html | url-status = dead }}

{{cite journal | vauthors = Mshaik R, Simonet J, Georgievski A, Jamal L, Bechoua S, Ballerini P, Bellaye PS, Mlamla Z, Pais de Barros JP, Geissler A, Francin PJ, Girodon F, Garrido C, Quéré R | title = HSP90 inhibitor NVP-BEP800 affects stability of SRC kinases and growth of T-cell and B-cell acute lymphoblastic leukemia | journal = Blood Cancer Journal | volume = 11 | issue = 3 | pages = 61 | date = March 2021 | pmid = 33737511 | pmc = 7973815 | doi = 10.1038/s41408-021-00450-2 | doi-access = free }}

{{cite journal | vauthors = Müller G, Wied S, Frick W | title = Cross talk of pp125(FAK) and pp59(Lyn) non-receptor tyrosine kinases to insulin-mimetic signaling in adipocytes | journal = Molecular and Cellular Biology | volume = 20 | issue = 13 | pages = 4708–4723 | date = Jul 2000 | pmid = 10848597 | pmc = 85892 | doi = 10.1128/mcb.20.13.4708-4723.2000 }}

{{cite journal | vauthors = Müller G, Wied S, Frick W | title = Cross talk of pp125(FAK) and pp59(Lyn) non-receptor tyrosine kinases to insulin-mimetic signaling in adipocytes | journal = Molecular and Cellular Biology | volume = 20 | issue = 13 | pages = 4708–4723 | date = Jul 2000 | pmid = 10848597 | pmc = 85892 | doi = 10.1128/mcb.20.13.4708-4723.2000 }}

{{cite journal | vauthors = Müller G | title = The molecular mechanism of the insulin-mimetic/sensitizing activity of the antidiabetic sulfonylurea drug Amaryl | journal = Molecular Medicine | location = Cambridge, Mass. | volume = 6 | issue = 11 | pages = 907–933 | date = Nov 2000 | pmid = 11147570 | pmc = 1949923 | doi = 10.1007/BF03401827 }}

{{cite journal | vauthors = Müller G, Schulz A, Wied S, Frick W | title = Regulation of lipid raft proteins by glimepiride- and insulin-induced glycosylphosphatidylinositol-specific phospholipase C in rat adipocytes | journal = Biochemical Pharmacology | volume = 69 | issue = 5 | pages = 761–780 | date = Mar 2005 | pmid = 15710354 | doi = 10.1016/j.bcp.2004.11.014 }}

{{cite journal | vauthors = Nishizumi H, Horikawa K, Mlinaric-Rascan I, Yamamoto T | title = A double-edged kinase Lyn: a positive and negative regulator for antigen receptor-mediated signals | journal = The Journal of Experimental Medicine | volume = 187 | issue = 8 | pages = 1343–1348 | date = Apr 1998 | pmid = 9547345 | pmc = 2212230 | doi = 10.1084/jem.187.8.1343 }}

{{cite journal | vauthors = Ochman AR, Lipinski CA, Handler JA, Reaume AG, Saporito MS | title = The Lyn kinase activator MLR-123 is a novel insulin receptor potentiator that elicits a rapid-onset and durable improvement in glucose homeostasis in animal models of type 2 diabetes | journal = The Journal of Pharmacology and Experimental Therapeutics | volume = 342 | issue = 1 | pages = 23–32 | date = 2012 | pmid = 22431203 | doi = 10.1124/jpet.112.192187 | s2cid = 7288053 | url = http://jpet.aspetjournals.org/content/suppl/2012/03/19/jpet.112.192187 | url-access = subscription }}{{Dead link|date=March 2024 |bot=InternetArchiveBot |fix-attempted=yes }}

{{cite journal | vauthors = Pathan NI, Geahlen RL, Harrison ML | title = The protein-tyrosine kinase Lck associates with and is phosphorylated by Cdc2 | journal = The Journal of Biological Chemistry | volume = 271 | issue = 44 | pages = 27517–27523 | date = Nov 1996 | pmid = 8910336 | doi = 10.1074/jbc.271.44.27517 | doi-access = free }}

{{cite journal | vauthors = Poe JC, Fujimoto M, Jansen PJ, Miller AS, Tedder TF | title = CD22 forms a quaternary complex with SHIP, Grb2, and Shc. A pathway for regulation of B lymphocyte antigen receptor-induced calcium flux | journal = The Journal of Biological Chemistry | volume = 275 | issue = 23 | pages = 17420–17427 | date = Jun 2000 | pmid = 10748054 | doi = 10.1074/jbc.M001892200 | doi-access = free }}

{{cite journal | vauthors = Qiu W, Cobb RR, Scholz W | title = Inhibition of p130cas tyrosine phosphorylation by calyculin A | journal = Journal of Leukocyte Biology | volume = 63 | issue = 5 | pages = 631–635 | date = May 1998 | pmid = 9581808 | doi = 10.1002/jlb.63.5.631 | s2cid = 11177730 | doi-access = free }}

{{cite journal | vauthors = Ren CL, Morio T, Fu SM, Geha RS | title = Signal transduction via CD40 involves activation of lyn kinase and phosphatidylinositol-3-kinase, and phosphorylation of phospholipase C gamma 2 | journal = The Journal of Experimental Medicine | volume = 179 | issue = 2 | pages = 673–680 | date = Feb 1994 | pmid = 7507510 | pmc = 2191357 | doi = 10.1084/jem.179.2.673 }}

{{cite journal | vauthors = Saijo K, Schmedt C, Su IH, Karasuyama H, Lowell CA, Reth M, Adachi T, Patke A, Santana A, Tarakhovsky A | title = Essential role of Src-family protein tyrosine kinases in NF-kappaB activation during B cell development | journal = Nature Immunology | volume = 4 | issue = 3 | pages = 274–279 | date = Mar 2003 | pmid = 12563261 | doi = 10.1038/ni893 | s2cid = 32559368 }}

{{cite journal | vauthors = Saporito MS, Ochman AR, Lipinski CA, Handler JA, Reaume AG | title = MLR-1023 is potent and selective allosteric activator of Lyn kinase in vitro that improves glucose tolerance in vivo | journal = The Journal of Pharmacology and Experimental Therapeutics | volume = 342 | issue = 1 | pages = 15–22 | date = 2011 | pmid = 22473614 | doi = 10.1124/jpet.112.192096 | s2cid = 26419896 | url = http://jpet.aspetjournals.org/content/342/1/15 | url-access = subscription }}

{{cite journal | vauthors = Sidorenko SP, Law CL, Chandran KA, Clark EA | title = Human spleen tyrosine kinase p72Syk associates with the Src-family kinase p53/56Lyn and a 120-kDa phosphoprotein | journal = Proceedings of the National Academy of Sciences of the United States of America | volume = 92 | issue = 2 | pages = 359–363 | date = Jan 1995 | pmid = 7831290 | pmc = 42739 | doi = 10.1073/pnas.92.2.359 | bibcode = 1995PNAS...92..359S | doi-access = free }}

{{cite journal | vauthors = Smith KG, Tarlinton DM, Doody GM, Hibbs ML, Fearon DT | title = Inhibition of the B cell by CD22: a requirement for Lyn | journal = The Journal of Experimental Medicine | volume = 187 | issue = 5 | pages = 807–811 | date = Mar 1998 | pmid = 9480991 | pmc = 2212179 | doi = 10.1084/jem.187.5.807 }}

{{cite journal | vauthors = Suzuki-Inoue K, Tulasne D, Shen Y, Bori-Sanz T, Inoue O, Jung SM, Moroi M, Andrews RK, Berndt MC, Watson SP | title = Association of Fyn and Lyn with the proline-rich domain of glycoprotein VI regulates intracellular signaling | journal = The Journal of Biological Chemistry | volume = 277 | issue = 24 | pages = 21561–21566 | date = Jun 2002 | pmid = 11943772 | doi = 10.1074/jbc.M201012200 | doi-access = free }}

{{cite journal | vauthors = Umemori H, Wanaka A, Kato H, Takeuchi M, Tohyama M, Yamamoto T | title = Specific expressions of Fyn and Lyn, lymphocyte antigen receptor-associated tyrosine kinases, in the central nervous system | journal = Brain Research. Molecular Brain Research | volume = 16 | issue = 3–4 | pages = 303–310 | date = Dec 1992 | pmid = 1337939 | doi = 10.1016/0169-328X(92)90239-8 }}

{{cite journal | vauthors = Wang X, Li Y, Wang X, Zhang Y, Liu Z, Zhong N, Wu M, Li G | title = Lyn regulates mucus secretion and MUC5AC via the STAT6 signaling pathway during allergic airway inflammation | journal = Scientific Reports | volume = 7 | pages = 42675 | date = Feb 2016 | pmid = 28205598 | pmc = 5312001 | doi = 10.1038/srep42675 | doi-access = free }}

{{cite journal | vauthors = Xu H, Zhao H, Tian W, Yoshida K, Roullet JB, Cohen DM | title = Regulation of a transient receptor potential (TRP) channel by tyrosine phosphorylation. SRC family kinase-dependent tyrosine phosphorylation of TRPV4 on TYR-253 mediates its response to hypotonic stress | journal = The Journal of Biological Chemistry | volume = 278 | issue = 13 | pages = 11520–11527 | date = Mar 2003 | pmid = 12538589 | doi = 10.1074/jbc.M211061200 | doi-access = free }}

{{cite journal | vauthors = Xu Y, Harder KW, Huntington ND, Hibbs ML, Tarlinton DM | title = Lyn tyrosine kinase: accentuating the positive and the negative | journal = Immunity | volume = 22 | issue = 1 | pages = 9–18 | date = Jan 2005 | pmid = 15664155 | doi = 10.1016/j.immuni.2004.12.004 | doi-access = free }}

{{cite journal | vauthors = Yamada E, Pessin JE, Kurland IJ, Schwartz GJ, Bastie CC | title = Fyn-dependent regulation of energy expenditure and body weight is mediated by tyrosine phosphorylation of LKB1 | journal = Cell Metabolism | volume = 11 | issue = 2 | pages = 113–124 | date = Feb 2010 | pmid = 20142099 | pmc = 2830006 | doi = 10.1016/j.cmet.2009.12.010 }}

{{cite journal | vauthors = Yamamoto T, Yamanashi Y, Toyoshima K | title = Association of Src-family kinase Lyn with B-cell antigen receptor | journal = Immunological Reviews | volume = 132 | pages = 187–206 | date = Apr 1993 | pmid = 8349296 | doi = 10.1111/j.1600-065X.1993.tb00843.x | s2cid = 10782326 }}

{{cite journal | vauthors = Yamanashi Y, Fukushige S, Semba K, Sukegawa J, Miyajima N, Matsubara K, Yamamoto T, Toyoshima K | title = The yes-related cellular gene lyn encodes a possible tyrosine kinase similar to p56lck | journal = Molecular and Cellular Biology | volume = 7 | issue = 1 | pages = 237–243 | date = Jan 1987 | pmid = 3561390 | pmc = 365062 | doi = 10.1128/MCB.7.1.237 }}

{{cite journal | vauthors = Yamanashi Y, Mori S, Yoshida M, Kishimoto T, Inoue K, Yamamoto T, Toyoshima K | title = Selective expression of a protein-tyrosine kinase, p56lyn, in hematopoietic cells and association with production of human T-cell lymphotropic virus type I | journal = Proceedings of the National Academy of Sciences of the United States of America | volume = 86 | issue = 17 | pages = 6538–6542 | date = Sep 1989 | pmid = 2505253 | pmc = 297879 | doi = 10.1073/pnas.86.17.6538 | bibcode = 1989PNAS...86.6538Y | doi-access = free }}

{{cite journal | vauthors = Yamanashi Y, Fukui Y, Wongsasant B, Kinoshita Y, Ichimori Y, Toyoshima K, Yamamoto T | title = Activation of Src-like protein-tyrosine kinase Lyn and its association with phosphatidylinositol 3-kinase upon B-cell antigen receptor-mediated signaling | journal = Proceedings of the National Academy of Sciences of the United States of America | volume = 89 | issue = 3 | pages = 1118–1122 | date = Feb 1992 | pmid = 1371009 | pmc = 48397 | doi = 10.1073/pnas.89.3.1118 | bibcode = 1992PNAS...89.1118Y | doi-access = free }}

{{cite journal | vauthors = van Dijk TB, van Den Akker E, Amelsvoort MP, Mano H, Löwenberg B, von Lindern M | title = Stem cell factor induces phosphatidylinositol 3'-kinase-dependent Lyn/Tec/Dok-1 complex formation in hematopoietic cells | journal = Blood | volume = 96 | issue = 10 | pages = 3406–3413 | date = Nov 2000 | pmid = 11071635 | doi = 10.1182/blood.V96.10.3406 | url = https://pure.eur.nl/en/publications/395fb5fc-60e3-45d7-a9b1-fc7b9cc6b4bc | hdl = 1765/9530 | hdl-access = free }}

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Further reading

{{refbegin|35em}}

  • {{cite journal | vauthors = Jouvin MH, Numerof RP, Kinet JP | title = Signal transduction through the conserved motifs of the high affinity IgE receptor Fc epsilon RI | journal = Seminars in Immunology | volume = 7 | issue = 1 | pages = 29–35 | date = Feb 1995 | pmid = 7612892 | doi = 10.1016/1044-5323(95)90005-5 }}
  • {{cite journal | vauthors = Hibbs ML, Dunn AR | title = Lyn, a src-like tyrosine kinase | journal = The International Journal of Biochemistry & Cell Biology | volume = 29 | issue = 3 | pages = 397–400 | date = Mar 1997 | pmid = 9202419 | doi = 10.1016/S1357-2725(96)00104-5 }}
  • {{cite book | vauthors = Blasioli J, Goodnow CC | chapter = Lyn/CD22/SHP-1 and Their Importance in Autoimmunity | title = Current Directions in Autoimmunity | volume = 5 | pages = 151–160 | year = 2002 | pmid = 11826756 | doi = 10.1159/000060551 | isbn = 978-3-8055-7308-5 }}
  • {{cite journal | vauthors = Greenway AL, Holloway G, McPhee DA, Ellis P, Cornall A, Lidman M | title = HIV-1 Nef control of cell signalling molecules: multiple strategies to promote virus replication | journal = Journal of Biosciences | volume = 28 | issue = 3 | pages = 323–335 | date = Apr 2003 | pmid = 12734410 | doi = 10.1007/BF02970151 | s2cid = 33749514 }}
  • {{cite journal | vauthors = Tolstrup M, Ostergaard L, Laursen AL, Pedersen SF, Duch M | title = HIV/SIV escape from immune surveillance: focus on Nef | journal = Current HIV Research | volume = 2 | issue = 2 | pages = 141–151 | date = Apr 2004 | pmid = 15078178 | doi = 10.2174/1570162043484924 }}
  • {{cite journal | vauthors = Joseph AM, Kumar M, Mitra D | title = Nef: "necessary and enforcing factor" in HIV infection | journal = Current HIV Research | volume = 3 | issue = 1 | pages = 87–94 | date = Jan 2005 | pmid = 15638726 | doi = 10.2174/1570162052773013 }}
  • {{cite journal | vauthors = Stove V, Verhasselt B | title = Modelling thymic HIV-1 Nef effects | journal = Current HIV Research | volume = 4 | issue = 1 | pages = 57–64 | date = Jan 2006 | pmid = 16454711 | doi = 10.2174/157016206775197583 }}

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