Laosuchus

L. naga was found in the Luang Prabang Basin of Northern Laos, part of the Indochina block. The site was first discovered by J. B. H. Counillon in 1896 as part of the Pavie's third Mission. Counillon was tasked with mapping mineral resources for the French colonial empire. L. naga was discovered during a 2005 expedition to the area, along with remains of dicynodonts.{{Cite journal|last=Steyer|first=J. Sebastien|date=2009|title=The geological and palaeontological exploration of Laos; following in the footsteps of J. B. H. Counillon and A. Pavie|journal=Geological Society, London, Special Publications|language=en|volume=315|issue=1|pages=25–32|doi=10.1144/SP315.3|bibcode=2009GSLSP.315...25S|s2cid=131670024|issn=0305-8719}}{{Cite journal|last1=Bercovici|first1=Antoine|last2=Bourquin|first2=Sylvie|last3=Broutin|first3=Jean|last4=Steyer|first4=Jean-Sébastien|last5=Battail|first5=Bernard|last6=Véran|first6=Monette|last7=Vacant|first7=Renaud|last8=Khenthavong|first8=Bounxou|last9=Vongphamany|first9=Sotsy|date=2012-10-22|title=Permian continental paleoenvironments in Southeastern Asia: New insights from the Luang Prabang Basin (Laos)|journal=Journal of Asian Earth Sciences|language=en|volume=60|pages=197–211|doi=10.1016/j.jseaes.2012.08.019|bibcode=2012JAESc..60..197B|issn=1367-9120}} It was later described by Arbez, Sidor, and Steyer in 2018.{{Cite journal|last1=Arbez|first1=Thomas|last2=Sidor|first2=Christian A.|last3=Steyer|first3=J.-Sébastien|date=2019-07-18|title=Laosuchus naga gen. et sp. nov., a new chroniosuchian from South-East Asia (Laos) with internal structures revealed by micro-CT scan and discussion of its palaeobiology|url=https://hal.sorbonne-universite.fr/hal-02048152/file/Arbez%20et%20al%202018%20Laosuchus%20HAL.pdf|journal=Journal of Systematic Palaeontology|language=en|volume=17|issue=14|pages=1165–1182|doi=10.1080/14772019.2018.1504827|bibcode=2019JSPal..17.1165A |issn=1477-2019|s2cid=91670454}} Its name comes from the Nāga, a snake-like deity that appears in multiple east Asian religions. In 2021 a new species L. hun was described from the Naobaogou Formation of the Daqing Mountains of Inner Mongolia, China.{{Cite journal|last1=Liu|first1=J.|last2=Chen|first2=J.|year=2021|title=The tetrapod fauna of the upper Permian Naobaogou Formation of China: 7. Laosuchus hun sp. nov. (Chroniosuchia) and interrelationships of chroniosuchians|journal=Journal of Systematic Palaeontology|volume=18|issue=24|pages=2043–2058|doi=10.1080/14772019.2021.1873435|s2cid=232116225}}

Laosuchus naga is represented by a single skull and articulated left hemimandible designated as specimen MDS-LPQ 2005-09, stored at the Musée des Dinosaures in Savannakhet. The skull, roughly 26 centimeters in length, is similar in shape to that of crocodiles. Its long snout bore marginal labyrinthodont teeth with an average height of 9 millimeters. Its nares are similar in shape to Madygenerpeton pustulatus. Like M. pustulatus, it also has oval-shaped orbits that are raised above the skull roof, but the orbits are proportionally smaller. Its choanae are relatively long compared to other chroniosuchians.

L. naga has a number of traits that make it unique among chroniosuchia. It lacks palatal tusks, bearing only small denticles on the palate. Its pineal foramen is significantly reduced with a 1mm diameter(M. pustulatus, C. dongusensis, and B. schumanni have diameters of 2mm, 2.5mm, and 3.5mm respectivelyClack, J., & Klembara, J. (2009). AN ARTICULATED SPECIMEN OF CHRONIOSAURUS DONGUSENSIS AND THE MORPHOLOGY AND RELATIONSHIPS OF THE CHRONIOSUCHIDS. Patterns And Processes In Early Vertebrate Evolution, 81(81), 15-42.{{Cite journal|last1=Schoch|first1=Rainer R.|last2=Voigt|first2=Sebastian|last3=Buchwitz|first3=Michael|date=2010-11-01|title=A chroniosuchid from the Triassic of Kyrgyzstan and analysis of chroniosuchian relationships|journal=Zoological Journal of the Linnean Society|language=en|volume=160|issue=3|pages=515–530|doi=10.1111/j.1096-3642.2009.00613.x|issn=0024-4082|doi-access=free}}{{Cite journal|last1=Witzmann|first1=Florian|last2=Schoch|first2=Rainer R.|date=2018-07-15|title=Skull and postcranium of the bystrowianid Bystrowiella schumanni from the Middle Triassic of Germany, and the position of chroniosuchians within Tetrapoda|journal=Journal of Systematic Palaeontology|volume=16|issue=9|pages=711–739|doi=10.1080/14772019.2017.1336579|bibcode=2018JSPal..16..711W |s2cid=90784937|issn=1477-2019}}). A transverse flange extends from the pterygoid, contacting the maxilla. CT-scanning revealed an autapomorphic internal crest on the dorsal palate that follows the internal margin of the choanae. The parasphenoid bears a thin ventro-medial ridge, a condition also seen in Discosauriscus austriacus{{Cite journal|last=Klembara|first=Jozef|date=1997-03-29|title=The cranial anatomy of Discosauricsus Kuhn, a seymouriamorph tetrapod from the Lower Permian of the Boskovice Furrow (Czech Republic)|journal=Philosophical Transactions of the Royal Society of London. Series B: Biological Sciences|volume=352|issue=1351|pages=257–302|doi=10.1098/rstb.1997.0021|pmc=1691931}} Its tabular horn and posterior squamosal contact, closing the otic notch.

There are a number of traits that L. naga share with other chroniosuchians. It bears a subtriangular fontanelle on its premaxilla, a poorly ossified braincase, a well developed pterygoid flange, and a relatively narrow parasphenoid bearing a crest between the basicranial joint.

CT-scanning also revealed the presence of canals within the mandible, snout, and bones in front of the orbit. The canals in the skull roof are interpreted as being related to a lateral line system. Canals in the tip of the snout and mandible are more complex and could be neurovascular canals, which modern animals use for thermoreception, electroreception, or mechanoreception. The paleontologists who described L. naga suggest that the lateral line system was used to detect prey beneath the water surface while the neurovascular system could let it detect movement at the water surface similar to modern crocodiles.

L. hun is distinguished from L. naga by several traits, including reduced palatal dentition, with a few denticles present on the vomer and pterygoid bones, and an irregular posterior cheek margin.

The two families within chroniosuchia, Chroniosuchidae and Bystrowianidae, are differentiated primarily based on postcranial elements, such as the shape of their vertebrae and the degree of overlap between their dorsal osteoderms.{{Cite journal|last1=Buchwitz|first1=Michael|last2=Foth|first2=Christian|last3=Kogan|first3=Ilja|last4=Voigt|first4=Sebastian|date=2012|title=On the use of osteoderm features in a phylogenetic approach on the internal relationships of the Chroniosuchia (Tetrapoda: Reptiliomorpha)|journal=Palaeontology|language=en|volume=55|issue=3|pages=623–640|doi=10.1111/j.1475-4983.2012.01137.x|issn=1475-4983|doi-access=free|bibcode=2012Palgy..55..623B }}{{Cite journal|last1=Jiang|first1=Shan|last2=Ji|first2=Shu-an|last3=Mo|first3=Jinyou|date=2017|title=First record of Bystrowianid Chroniosuchians (Amphibia: Anthracosauromorpha) from the Middle Permian of China|journal=Acta Geologica Sinica - English Edition|volume=91|issue=5|pages=1523–1529|doi=10.1111/1755-6724.13397|bibcode=2017AcGlS..91.1523J |issn=1755-6724}} Thus, while L. naga has numerous characteristics supporting its inclusion to chroniosuchia, the lack of postcranial elements in addition to several cranial traits preclude their inclusion to either family. As a result, L. naga was classified as Chroniosuchia incertae sedis. In the 2021 description of L. hun postcranial remains indicated that Laosuchus belonged to the Chroniosuchidae.

The depositional environment that L. naga occurred in consisted of braided rivers transitioning into alluvial plains, with an input of volcanic sediment. In addition to this, the presence of a lateral line system and poorly ossified braincase imply that L. naga spent much of its time in the water. According to the paleontologists who described L. naga, Its placement in nonmarine sediment provides a line of support for the scenario that the North China Block, South China Block, and Indochina block were connected like a peninsula and linked to Laurussia during the Permian and Triassic.{{Cite journal|last=Metcalfe|first=I.|date=2006-01-01|title=Palaeozoic and Mesozoic tectonic evolution and palaeogeography of East Asian crustal fragments: The Korean Peninsula in context|journal=Gondwana Research|series=Tectonic Evolution of Korean Peninsula and Adjacent Crustal Fragments in Asia|language=en|volume=9|issue=1|pages=24–46|doi=10.1016/j.gr.2005.04.002|bibcode=2006GondR...9...24M|issn=1342-937X}}

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• [https://pbs.twimg.com/media/DoKX3J_X0AA8dwL.jpg Photographs and illustrations of the specimen, published by the Journal of Systematic Palaeontology twitter account]

{{Early tetrapods}}

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Category:Chroniosuchia

Category:Fossil taxa described in 2018

Category:Fossils of Laos

Category:Triassic tetrapods of Asia

Category:Permian tetrapods of Asia