List of bats of Madagascar#Family Miniopteridae

Bats are one of the major components of the indigenous mammalian fauna of Madagascar, in addition to tenrecs, lemurs, euplerid carnivores, and nesomyine rodents. Forty-six bat species have so far been recorded on Madagascar, of which thirty-six occur only on the island. However, new species continue to be discovered, causing the number of species to rise rapidly; for example, Nick Garbutt's Mammals of Madagascar (2007) listed only 36 species.Garbutt, 2007, p. 59 Most Malagasy bats have their origins in nearby mainland Africa, but on at least three occasions—Pipistrellus raceyi, Pteropus rufus, and the species pair Emballonura atrata–E. tiavato—bats have colonized Madagascar from Asia.Bates et al., 2006, p. 321

Taxonomic classification

The following bat genera and families include species found on Madagascar (all species counts are for Madagascar only):

Family Pteropodidae (3 endemic species)
Genus Eidolon (1 endemic species)
Genus Pteropus (1 endemic species)
Genus Rousettus (1 endemic species)
Family Hipposideridae (6 endemic species)
Genus Hipposideros (1 extinct endemic species)
Genus Macronycteris (2 endemic species)
Genus Paratriaenops (2 endemic species)
Genus Triaenops (2 endemic species, one of which is extinct)
Family Emballonuridae (2 endemic, 2 non-endemic species)
Genus Coleura (1 non-endemic species)
Genus Emballonura (2 endemic species)
Genus Taphozous (1 non-endemic species)
Family Nycteridae (1 endemic species)
Genus Nycteris (1 endemic species)
Family Myzopodidae (2 endemic species)
Genus Myzopoda (2 endemic species)
Family Molossidae (5 endemic, 3 non-endemic species)
Genus Chaerephon (2 endemic, 1 non-endemic species)
Genus Mops (1 endemic, 1 non-endemic species)
Genus Mormopterus (1 endemic species)
Genus Otomops (1 endemic species)
Genus Tadarida (1 non-endemic species)
Family Miniopteridae (9 endemic, 2 non-endemic species)
Genus Miniopterus (9 endemic, 2 non-endemic species)
Family Vespertilionidae (8 endemic, 4 non-endemic species)
Genus Hypsugo (1 non-endemic species)
Genus Myotis (1 endemic species)
Genus Neoromicia (3 endemic species)
Genus Pipistrellus (1 endemic, 1 non-endemic species)
Genus Scotophilus (3 endemic, 1 non-endemic species)

Key

class="wikitable"

|style="background:#e3e3e3"|Scientific name

|Scientific name of the species

style="background:#e3e3e3"|Classified

|Year when the species was formally described and classified, as well as the binomial authority of the species

style="background:#e3e3e3"|Distribution

|Geographic distribution of the species. Abbreviations used are S, south; N, north; E, east; W, west; C, central; SW, southwest; etcetera.

style="background:#e3e3e3"|Forearm

|Range in forearm length of adult members of the species, in metric and English units

style="background:#e3e3e3"|Conservation status

|Conservation status of the species, per IUCN as of 2008, except as indicated. "Not evaluated" is used to indicate that no IUCN status assessment is available.

Family Pteropodidae

Pteropodidae are a diverse family, with 186 species recognized in 2005,Wilson and Reeder, 2005, p. xxix which occurs across the tropical regions of the Old World. They include the largest bats, but also some smaller species, and are mostly diurnal and frugivorous. Three species are known from Madagascar; each is classified in its own genus and is most closely related to species from outside Madagascar.Nowak, 1994, pp. 48–49; Garbutt, 2007, p. 59

class="wikitable sortable"

!Scientific name

!Classified

!Distribution

!Forearm

!Conservation status

!class="unsortable"|References

Eidolon dupreanum{{#tag:ref|Sometimes included in the mainland African species Eidolon helvum.Simmons, 2005, p. 321|group=Note}}

|1866, Pollen

|Madagascar only

|115 to 130 mm (4.5 to 5.1 in)

|Vulnerable

|Simmons, 2005, p. 321; Garbutt, 2007, p. 62; Andriafidison et al., 2008a

Pteropus rufus

|1803, E. Geoffroy

|Madagascar only

|155 to 175 mm (6.1 to 6.9 in)

|Vulnerable

|Simmons, 2005, p. 343; Garbutt, 2007, p. 60; Andriafidison et al., 2008b

Rousettus madagascariensis{{#tag:ref|Sometimes included in the mainland African species Rousettus lanosus.Simmons, 2005, p. 348|group=Note}}

|1928, G. Grandidier

|Madagascar only; absent in SW

|65 to 75 mm (2.6 to 3.0 in)

|Near threatened

|Simmons, 2005, p. 348; Garbutt, 2007, p. 64; Andriafidison et al., 2008c

Family Hipposideridae

Hipposideridae are a moderately diverse family—81 species were listed in 2005—and occur across the Old World tropics. Insectivorous, cave-roosting, and characterized by an elaborate noseleaf, they have often been united with the horseshoe bats (Rhinolophus; absent from Madagascar) into a single family Rhinolophidae, but are currently classified separately. Six species, all endemic, are known from Madagascar, of which four are extant. Macronycteris commersoni is the largest non-pteropodid bat of Madagascar and the extinct Hipposideros besaoka was even larger. The other species belong to the closely related genera Triaenops and Paratriaenops; the latter was split from Triaenops in 2009 and is restricted to Madagascar and the western Seychelles.Nowak, 1994, p. 110; Garbutt, 2007, pp. 68–69; Samonds, 2007; Benda and Vallo, 2009

class="wikitable sortable"

!Scientific name

!Classified

!Distribution

!Forearm

!Conservation status

!class="unsortable"|References

Hipposideros besaoka

|2007, Samonds

|Anjohibe, NW Madagascar only

|–

|Extinct{{#tag:ref|Not in IUCN Red List.|group=Note|name=not-in-IUCN}}

|Samonds, 2007, pp. 49ff., 62

Macronycteris commersoni{{#tag:ref|Previously placed in the genus Hipposideros as H. commersoni and H. cryptovalorona.Foley et al., 2017|group=Note|name=Macronycteris}}

|1813, E. Geoffroy

|Madagascar only

|83 to 97 mm (3.3 to 3.8 in)

|Near threatened

|Simmons, 2005, p. 369; Garbutt, 2007, p. 69; Andriafidison et al., 2008f

Macronycteris cryptovalorona

|2016, Goodman et al.

|Madagascar only

|80 to 81 mm (3.1 to 3.2 in)

|Not evaluated

|Goodman et al., 2016

Paratriaenops auritus{{#tag:ref|Previously placed in the genus Triaenops as Triaenops auritus and Triaenops furculus, but now classified in Paratriaenops.Benda and Vallo, 2009, p. 34|group=Note|name=Triaenops}}

|1912, G. Grandidier

|N and NW Madagascar only

|44 to 51 mm (1.7 to 2.0 in)

|VulnerableAs Triaenops auritus.

|Simmons, 2005, p. 378; Ranivo and Goodman, 2007, p. 964, appendix 2; Benda and Vallo, 2009, p. 34; Andriafidison et al., 2008o

Paratriaenops furcula

|1906, Trouessart

|W and SW Madagascar only

|42 to 49 mm (1.7 to 1.9 in)

|Least concernAs Triaenops furculus.

|Simmons, 2005, p. 378; Ranivo and Goodman, 2007, p. 964, appendix 2; Benda and Vallo, 2009, p. 34; Andriafidison et al., 2008p

Triaenops goodmani

|2007, Samonds

|Anjohibe, NW Madagascar only

|–

|Extinct

|Samonds, 2007, pp. 46ff., 62

Triaenops menamena{{#tag:ref|Populations now included in this species have previously also been included in Triaenops persicus, which is now restricted to the Middle East, but are more conventionally known as Triaenops rufus. This name is, however, in fact a synonym of T. persicus; therefore, the new name menamena was introduced for the Madagascan populations in 2009.Simmons, 2005, p. 379; Goodman and Ranivo, 2009, p. 54; Benda and Vallo, 2009, p. 34|group=Note}}

|2009, Goodman and Ranivo

|N, W, and S Madagascar only

|46 to 56 mm (1.8 to 2.2 in)

|Least concernAs Triaenops rufus.

|Ranivo and Goodman, 2007, appendix 3; Goodman and Ranivo, 2009, p. 54; Benda and Vallo, 2009, p. 34; Andriafidison et al., 2008q

Family Emballonuridae

With 51 species (2005), Emballonuridae are a moderately diverse family. Found in tropical and subtropical regions across the world, they are characterized by a tail that extends beyond the uropatagium (tail membrane), but may be retracted into a sheath. Four species are known from Madagascar, of which two are endemic and two others are shared with mainland Africa.Walker, 1994, pp. 87–88; Garbutt, 2007, p. 64

class="wikitable sortable"

!Scientific name

!Classified

!Distribution

!Forearm

!Conservation status

!class="unsortable"|References

Coleura afra

|1852, Peters

|Ankarana, N Madagascar; Sub-Saharan Africa; Yemen

|45 to 55 mm (1.8 to 2.2 in)

|Least concern

|Simmons, 2005, p. 386; Garbutt, 2007, p. 66; Goodman et al., 2008a; Mickleburgh et al., 2008c

Emballonura atrata

|1836, Eydoux and Gervais

|E Madagascar only

|c. 37 to 40 mm (1.5 to 1.6 in)

|Least concern

|Goodman et al., 2006a; Garbutt, 2007, p. 65; Jenkins et al., 2008a

Emballonura tiavato

|2006, Goodman et al.

|N and W Madagascar only

|35 to 41 mm (1.4 to 1.6 in)

|Least concern

|Goodman et al., 2006a; Garbutt, 2007, p. 66; Jenkins et al., 2008b

Taphozous mauritianus

|1818, E. Geoffroy

|Madagascar and other W Indian Ocean islands; Sub-Saharan Africa

|58 to 64 mm (2.3 to 2.5 in)

|Least concern

|Simmons, 2005, p. 384; Garbutt, 2007, p. 67; Hutson et al., 2008c

Family Nycteridae

Nycteridae is a small family of 16 species (2005) in a single genus found in Africa and east to the Sunda Islands. They are characterized by a groove on their face and are insectivorous. A single, poorly known species has been recorded from Madagascar.Walker, 1994, pp. 101–102; Garbutt, 2007, p. 68

class="wikitable sortable"

!Scientific name

!Classified

!Distribution

!Forearm

!Conservation status

!class="unsortable"|References

Nycteris madagascariensis{{#tag:ref|Sometimes included in the mainland African species Nycteris macrotis.Simmons, 2005, pp. 392–393|group=Note}}

|1937, G. Grandidier

|N Madagascar only

|50 to 52 mm (about 2.0 in)

|Data deficient

|Simmons, 2005, p. 393; Garbutt, 2007, p. 68; Hutson et al., 2008a

Family Myzopodidae

This family, characterized by suction disks on the hand and feet, is unique to Madagascar. (It does, however, have a fossil record in Africa extending from the late Eocene to the Pleistocene.{{Cite journal | last1 = Gunnell | first1 = G. F. | last2 = Simmons | first2 = N. B. | last3 = Seiffert | first3 = E. R. | date = 2014-02-04 | title = New Myzopodidae (Chiroptera) from the Late Paleogene of Egypt: Emended Family Diagnosis and Biogeographic Origins of Noctilionoidea | journal = PLoS ONE | volume = 9 | issue = 2 | pages = e86712 | doi = 10.1371/journal.pone.0086712 | pmid=24504061 | pmc=3913578| bibcode = 2014PLoSO...986712G | doi-access = free }}) A single species has historically been recognized, but eastern and western populations were classified as separate species in 2007.Garbutt, 2007, p. 72; Goodman et al., 2007

class="wikitable sortable"

!Scientific name

!Classified

!Distribution

!Forearm

!Conservation status

!class="unsortable"|References

Myzopoda aurita

|1878, Milne-Edwards and Grandidier

|E Madagascar only

|46 to 49 mm (1.8 to 1.9 in)

|Least concern

|Simmons, 2005, p. 394; Garbutt, 2007, p. 72; Goodman et al., 2007; Jenkins et al., 2008c

Myzopoda schliemanni

|2007, Goodman et al.

|W Madagascar only

|45 to 49 mm (1.8 to 1.9 in)

|Least concern

|Garbutt, 2007, p. 73; Goodman et al., 2007; Jenkins et al., 2008d

Family Molossidae

This diverse family of 100 species (2005) occurs across the world in tropical regions. The tail conspicuously projects from the uropatagium and the wings are long. Eight species are known from Madagascar, four of which are endemic.Garbutt, 2007, pp. 80–84

class="wikitable sortable"

!Scientific name

!Classified

!Distribution

!Forearm

!Conservation status

!class="unsortable"|References

Chaerephon atsinanana{{#tag:ref|Previously included in the mainland African Chaerephon pumilus (or Tadarida pumila).Mickleburgh et al., 2008; Goodman et al., 2010c, pp. 1–2|group=Note}}

|2010, Goodman et al.

|E Madagascar

|37 to 42 mm (1.5 to 1.7 in)

|Goodman et al., 2010c, pp. 1–2, table 2

Chaerephon jobimena

|2004, Goodman and Cardiff

|Madagascar only

|45 to 48 mm (1.8 to 1.9 in)

|Least concern

|Goodman and Cardiff, 2004, table 1; Garbutt, 2007, p. 82; Andriafidison et al., 2014

Chaerephon leucogaster

|1870, A. Grandidier

|W Madagascar; Sub-Saharan Africa; Mayotte

|33 to 38 mm (1.3 to 1.5 in)

|Not evaluated{{#tag:ref|Included in Tadarida pumila.Mickleburgh et al., 2008b|group=Note}}

|Goodman and Cardiff, 2004, p. 227; Simmons, 2005, p. 434; Mickleburgh et al., 2008b; Ratrimomanarivo et al., 2009, table 1; Goodman et al., 2010b, p. 128

Mops leucostigma

|1918, G.M. Allen

|Madagascar only

|42 to 45 mm (1.7 to 1.8 in)

|Least concernAs Tadarida leucostigma.

|Simmons, 2005, p. 442; Garbutt, 2007, p. 83; Andriafidison et al., 2008n

Mops midas

|1843, Sundevall

|Madagascar; Sub-Saharan Africa; Saudi Arabia

|62 to 63 mm (2.4 to 2.5 in)

|Least concernAs Tadarida midas.

|Simmons, 2005, p. 442; Garbutt, 2007, p. 83; Jenkins et al., 2008e

Mormopterus jugularis

|1865, Peters

|Madagascar only

|37 to 40 mm (1.5 to 1.6 in)

|Least concern

|Simmons, 2005, p. 445; Garbutt, 2007, p. 80; Andriafidison et al., 2008j

Otomops madagascariensis{{#tag:ref|Previously included in the mainland African Otomops martiensseni.Andriafidison et al., 2008k|group=Note}}

|1953, Dorst

|Madagascar only

|59 to 65 mm (2.3 to 2.6 in)

|Least concern

|Simmons, 2005, p. 447; Garbutt, 2007, p. 84; Andriafidison et al., 2008k

Tadarida fulminans

|1903, Thomas

|Madagascar; Sub-Saharan Africa north to Kenya

|57 to 60 mm (2.2 to 2.4 in)

|Least concern

|Simmons, 2005, p. 450; Garbutt, 2007, p. 81 Cotterill et al., 2008

Family Miniopteridae

This family contains a single genus, Miniopterus, with 19 species recognized in 2005,Simmons, 2005, pp. 519–522 which was classified in Vespertilionidae until recently.Miller-Butterworth et al., 2007, p. 1553 Insectivorous and characterized by long fingers, the species are all quite similar, leading to a confused classification. On Madagascar, four species were recognized as recently as 2007, but systematic research has led the number to increase to eleven, of which nine are restricted to Madagascar and two shared with the Comoros.Garbutt, 2007, pp. 77–80; subsequent revisions cited below

class="wikitable sortable"

!Scientific name

!Classified

!Distribution

!Forearm

!Conservation status

!class="unsortable"|References

Miniopterus aelleni

|2009, Goodman et al.

|N and W Madagascar; Anjouan, Comoros

|35 to 41 mm (1.4 to 1.6 in)

|Not evaluated

|Goodman et al., 2009, p. 6, table 3

Miniopterus brachytragos

|2009, Goodman et al.

|N and W Madagascar only

|35 to 38 mm (1.4 to 1.5 in)

|Not evaluated

|Goodman et al., 2009, fig. 1, table 3

Miniopterus egeri

|2011, Goodman et al.

|E Madagascar only

|37 to 40 mm (1.5 to 1.6 in)

|Not evaluated

|Goodman et al., 2011, p. 1, table 5

Miniopterus gleni

|1995, Peterson et al.

|Madagascar only, except S

|47 to 50 mm (1.9 to 2.0 in)

|Least concern

|Simmons, 2005, p. 519; Andriafidison et al., 2008g; Goodman et al., 2010a, fig. 1, table 1

Miniopterus griffithsi

|2009, Goodman et al.

|S Madagascar only

|48 to 50 mm (1.9 to 2.0 in)

|Not evaluated

|Goodman et al., 2010a, fig. 1, table 1

Miniopterus griveaudi

|1959, Harrison

|N and W Madagascar; Comoros

|35 to 38 mm (1.4 to 1.5 in)

|Data deficient

|Juste, 2008; Goodman et al., 2009, p. 5, table 3

Miniopterus mahafaliensis

|2009, Goodman et al.

|S Madagascar only

|35 to 40 mm (1.4 to 1.6 in)

|Not evaluated

Miniopterus majori

|1906, Thomas

|Madagascar only{{#tag:ref|Although often cited for the Comoros, its occurrence there is questionable.Goodman et al., 2010b, p. 132|group=Note}}

|43 to 47 mm (1.7 to 1.9 in)

|Least concern

|Simmons, 2005, p. 520; Jenkins and Rakotoarivelo, 2008a; Goodman et al., 2010b, p. 132

Miniopterus manavi

|1906, Thomas

|Central Highlands, Madagascar only

|38 to 39 mm (1.5 to 1.5 in)

|Least concern

|Goodman et al., 2009, p. 5, table 3; Andriafidison et al., 2008h

Miniopterus petersoni

|2008, Goodman et al.

|SE Madagascar only

|38 to 43 mm (1.5 to 1.7 in)

|Data deficient

|Goodman et al., 2008b, fig. 1, table 1; 2009, p. 5; Jenkins and Rakotoarivelo, 2008b

Miniopterus sororculus

|2007, Goodman et al.

|Central Highlands, Madagascar only

|42 to 45 mm (1.7 to 1.8 in)

|Least concern

|Goodman et al., 2008b, fig. 1, table 1; Jenkins et al., 2008f

Family Vespertilionidae

With 407 species (2005; including Miniopterus, which is now classified in its own family), Vespertilionidae is the largest bat family. Characterized by a tail contained in the uropatagium, they occur around the world in many habitats and are insectivorous.Garbutt, 2007, p. 73 Madagascar hosts an endemic species of the extremely widespread genus Myotis, four species (three endemic) of the house bat Scotophilus, and at least six (four endemic) of small vespertilionids ("pipistrelles") in the genera Hypsugo, Eptesicus, Neoromicia, and Pipistrellus.Garbutt, 2007, pp. 73–77; Bates et al., 2006, p. 299 The classification of the "pipistrelles" is confused, leading to many changing identifications.Bates et al., 2006, pp. 299–300 In addition to the six "pipistrelles" listed here, the African Neoromicia nanus has also been recorded from Madagascar, but the identification of the Madagascar records needs to be confirmed.Bates et al., 2006, p. 320; Hutson et al., 2008b

class="wikitable sortable"

!Scientific name

!Classified

!Distribution

!Forearm

!Conservation status

!class="unsortable"|References

Hypsugo anchietaePreviously placed in the genus Pipistrellus as Pipistrellus anchietae.

|1900, Seabra

|SW Madagascar; southern Africa

|28 to 31 mm (1.1 to 1.2 in)

|Least concernAs Pipistrellus anchietae.

|Simmons, 2005, p. 489; Bates et al., 2006, table 1, fig. 7, p. 319; Jacobs et al., 2008

Myotis goudoti

|1834, A. Smith

|Madagascar only

|32 to 41 mm (1.3 to 1.6 in)

|Least concern

|Simmons, 2005, p. 508; Garbutt, 2007, p. 74; Andriafidison et al., 2008i

Neoromicia malagasyensisPreviously included in Eptesicus, as Eptesicus malagasyensis and/or considered a subspecies of Neoromicia somalicus (=Eptesicus somalicus) from mainland Africa.

|1995, Peterson et al.

|Isalo, SC Madagascar only

|30 to 32 mm (1.2 to 1.3 in)

|EndangeredAs Eptesicus malagasyensis.

|Simmons, 2005, p. 495; Bates et al., 2006, table 1, pp. 313, 315; Andriafidison et al., 2008d

Neoromicia matrokaPreviously included in Eptesicus or Pipistrellus and/or considered a subspecies of Neoromicia capensis (=Eptesicus capensis; Pipistrellus capensis).

|1905, Thomas and Schwann

|E Madagascar only

|31 to 33 mm (1.2 to 1.3 in)

|Least concernAs Eptesicus matroka.

|Simmons, 2005, p. 456; Bates et al., 2006, table 1, fig. 7, pp. 312–313; Andriafidison et al., 2008e

Neoromicia robertsiSpecimens identified as this species were previously called Neoromicia melckorum, and they have also been included in the genus Eptesicus or Pipistrellus.

|2012, Goodman et al.

|E Madagascar only

|34 to 38 mm (1.3 to 1.5 in)

|Not evaluated

|Simmons, 2005, p. 494; Bates et al., 2006, table 1, fig. 7, pp. 315–316; Goodman et al., 2012

Pipistrellus hesperidusPreviously included in the European species Pipistrellus kuhlii.

|1840, Temminck

|W Madagascar; Sub-Saharan Africa

|29 to 31 mm (1.1 to 1.2 in)

|Least concern

|Simmons, 2005, p. 474; Bates et al., 2006, table 1, fig. 7, pp. 317–318; Mickleburgh et al., 2008a

Pipistrellus raceyi

|2006, Bates et al.

|W and E Madagascar only

|28 to 31 mm (1.1 to 1.2 in)

|Least concern

|Bates et al., 2006, table 1, fig. 7; Jenkins et al., 2008g

Scotophilus cf. borbonicusIt is uncertain whether the single known specimen from Madagascar belongs to this species, otherwise known from Réunion by a single specimen.

|1803, E. Geoffroy

|Sarodrano, SW Madagascar; Réunion

|51 to 52 mm (about 2.0 in)

|Data deficient

|Simmons, 2005, p. 465; Goodman et al., 2005, table 1, pp. 871–873; Garbutt, 2007, p. 76; Andriafidison et al., 2008k

Scotophilus marovaza

|2006, Goodman et al.

|W Madagascar only

|41 to 45 mm (1.6 to 1.8 in)

|Least concern

|Goodman et al., 2006b, p. 21, table 1; Jenkins et al., 2008h

Scotophilus robustus

|1881, Milne-Edwards

|Madagascar only

|62 to 65 mm (2.4 to 2.6 in)

|Least concern

|Simmons, 2005, p. 467; Goodman et al., 2005, table 1, pp. 873–875; Garbutt, 2007, p. 76; Andriafidison et al., 2008l

Scotophilus tandrefana

|2005, Goodman et al.

|W Madagascar only

|44 to 47 mm (1.7 to 1.9 in)

|Data deficient

|Goodman et al., 2005, table 1, p. 875; Garbutt, 2007, p. 76; Jenkins et al., 2008i

Notes

References

Literature cited

=General=

Garbutt, N. 2007. Mammals of Madagascar: A Complete Guide. London: A & C Black, 304 pp. {{ISBN|978-0-7136-7043-1}}
Goodman, S.M., Weyeneth, N., Ibrahim, Y., Saïd, I. and Ruedi, M. 2010b. [https://dx.doi.org/10.3161/150811010X504635 A review of the bat fauna of the Comoro Archipelago] (subscription required). Acta Chiropterologica 12(1):117–141.
Nowak, R.M. 1994. Walker's Bats of the World. Baltimore: The Johns Hopkins University Press, 287 pp. {{ISBN|978-0-8018-4986-2}}
Simmons, N.B. 2005. Order Chiroptera. Pp. 312–529 in Wilson, D.E. and Reeder, D.M. (eds.). [http://www.bucknell.edu/msw3 Mammal Species of the World: a taxonomic and geographic reference]. 3rd ed. Baltimore: The Johns Hopkins University Press, 2 vols., 2142 pp. {{ISBN|978-0-8018-8221-0}}
Wilson, D.E. and Reeder, D.M. (eds.). 2005. [http://www.bucknell.edu/msw3 Mammal Species of the World: a taxonomic and geographic reference]. 3rd ed. Baltimore: The Johns Hopkins University Press, 2 vols., 2142 pp. {{ISBN|978-0-8018-8221-0}}

=Pteropodidae=

Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008a. {{IUCNlink|7083|Eidolon dupreanum}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 26, 2010.
Andriafidison, D, Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Rabearivelo, A., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008b. {{IUCNlink|18756|Pteropus rufus}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 26, 2010.
Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008c. {{IUCNlink|19750|Rousettus madagascariensis}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 26, 2010.

=Hipposideridae=

Andriafidison, D, Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008f. {{IUCNlink|10120|Hipposideros commersoni}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008o. {{IUCNlink|40025|Triaenops auritus}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008p. {{IUCNlink|22089|Triaenops furculus}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008q. {{IUCNlink|40026|Triaenops rufus}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Benda, P. and Vallo, P. 2009. [https://web.archive.org/web/20160916211407/http://www.nm.cz/download/pm/zoo/benda_lit/Benda2009fzm.pdf Taxonomic revision of the genus Triaenops (Chiroptera: Hipposideridae) with description of a new species from southern Arabia and definitions of a new genus and tribe]. Folia Zoologica 58 (Monograph 1):1–45.
{{Cite journal|author=Foley, N.M. |author2=Goodman, S.M. |author3=Whelan, C.V. |author4=Puechmaille, S.J. |author5=Teeling, E. |year= 2017|title= Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae) (subscription required)|journal= Acta Chiropterologica|volume= 19|issue= 1|pages= 1–18|doi= 10.3161/15081109acc2017.19.1.001|s2cid= 89997441}}
Goodman, S.M. and Ranivo, J. 2009. [https://dx.doi.org/10.1515/MAMM.2009.011 The geographical origin of the type specimens of Triaenops rufus and T. humbloti (Chiroptera: Hipposideridae) reputed to be from Madagascar and the description of a replacement species name] (subscription required). Mammalia 73:47–55.
{{cite journal|author=Goodman, S.M. |author2=Schoeman, M.C. |author3=Rakotoarivelo, A. |author4=Willows-Munro, S. |title= How many species of Hipposideros have occurred on Madagascar since the Late Pleistocene? (subscription required)|journal= Zoological Journal of the Linnean Society|volume= 177|issue= 2|year= 2016|pages= 428–449|doi= 10.1111/zoj.12368|doi-access= free}}
Ranivo, J. and Goodman S.M. 2006. Révision taxinomique des Triaenops malgaches (Mammalia, Chiroptera, Hipposideridae). Zoosystema 28(4):963–985.
Samonds, K.E. 2007. [https://web.archive.org/web/20090718195034/http://redpath-staff.mcgill.ca/Samonds/PDFS/11%20Samonds%202007.pdf Late Pleistocene bat fossils from Anjohibe Cave, northwestern Madagascar]. Acta Chiropterologica 9(1):39–65.

=Emballonuridae=

Goodman, S.M., Cardiff, S.G., Ranivo, J., Russell, A.L. and Yoder, A.D. 2006a. [http://hdl.handle.net/2246/5816 A new species of Emballonura (Chiroptera, Emballonuridae) from the dry regions of Madagascar]. American Museum Novitates 3538:1–24.
Goodman, S.M., Cardiff, S.G. and Ratrimomanarivo, F.H. 2008a. [https://dx.doi.org/10.1017/S1477200008002715 First record of Coleura (Chiroptera: Emballonuridae) on Madagascar and identification and diagnosis of members of the genus] (subscription required). Systematics and Biodiversity 6(2):283–292.
Hutson, A.M., Racey, P., Ravino, J., Mickleburgh, S., Bergmans, W. and Fahr, J. 2008c. {{IUCNlink|21460|Taphozous mauritianus}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Jenkins, R.K.B., Rakotoarivelo, A.R., Ratrimomanarivo, F.H. and Cardiff, S.G. 2008a. {{IUCNlink|7671|Emballonura atrata}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Jenkins, R.K.B., Rakotoarivelo, A.R., Ratrimomanarivo, F.H. and Cardiff, S.G. 2008b. {{IUCNlink|136835|Emballonura tiavato}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Mickleburgh, S., Hutson, A.M., Racey, P.A., Cardiff, S. and Bergmans, W. 2008c. {{IUCNlink|5113|Coleura afra}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.

=Nycteridae=

Hutson, T., Racey, P.A. and Ravino, J. 2008a. {{IUCNlink|40022|Nycteris madagascariensis}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.

=Myzopodidae=

Goodman, S.M., Rakotondraparany, F. and Kofoky, A. 2007. [https://dx.doi.org/10.1016/j.mambio.2006.08.001 The description of a new species of Myzopoda (Myzopodidae: Chiroptera) from western Madagascar] (subscription required). Mammalian Biology 72(2):65–81.
Jenkins, R.K.B., Rakotoarivelo, A.R., Ratrimomanarivo, F.H. and Cardiff, S.G. 2008c. {{IUCNlink|14288|Myzopoda aurita}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Jenkins, R.K.B., Rakotoarivelo, A.R., Ratrimomanarivo, F.H. and Cardiff, S.G. 2008d. {{IUCNlink|136465|Myzopoda schliemanni}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.

=Molossidae=

Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008i. {{IUCNlink|13882|Mormopterus jugularis}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008j. {{IUCNlink|136564|Otomops madagascariensis}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008n. {{IUCNlink|40024|Tadarida leucostigma}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2014. {{IUCNlink|136393|Chaerephon jobimena}}. IUCN Red List of Threatened Species. Version 2014.3. Downloaded May 28, 2015.
Cotterill, F.P.D, Hutson, A.M., Racey, P.A. and Ravino, J. 2008. {{IUCNlink|21316|Tadarida fulminans}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Goodman, S.M. and Cardiff, S.G. 2004. A new species of Chaerephon (Molossidae) from Madagascar with notes on other members of the family. Acta Chiropterologica 6(2):227–248.
Goodman, S.M., Buccas, W., Naidoo, T., Ratrimomanarivo, F., Taylor, P.J. and Lamb, J. 2010c. [http://www.mapress.com/zootaxa/2010/2/zt02551p036.pdf Patterns of morphological and genetic variation in western Indian Ocean members of the Chaerephon 'pumilus' complex (Chiroptera: Molossidae), with the description of a new species from Madagascar] (subscription required). Zootaxa 2551:1–36.
Jenkins, R.K.B., Racey, P.A., Ranivo, J., Ratimomanarivo, F., Mickleburgh, S., Hutson, A.M., Bergmans, W., Cotterill, F.P.D. and Fahr, J. 2008e. {{IUCNlink|13841|Tadarida midas}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Mickleburgh, S., Hutson, A.M., Racey, P.A., Ravino, J., Bergmans, W., Cotterill, F.P.D. and Gerlach, J. 2008b. {{IUCNlink|4317|Tadarida pumila}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Ratrimomanarivo, F.H., Goodman, S.M., Stanley, W.T., Naidoo, T., Taylor, P.J. and Lamb, J. 2009. [https://dx.doi.org/10.3161/150811009X465677 Geographic and phylogeographic variation in Chaerephon leucogaster (Chiroptera: Molossidae) of Madagascar and the western Indian Ocean islands of Mayotte and Pemba] (subscription required). Acta Chiropterologica 11(1):25–52.

=Miniopteridae=

Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008g. {{IUCNlink|29608|Miniopterus gleni}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008h. {{IUCNlink|40040|Miniopterus manavi}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Goodman, S.M., Bradman, H.M., Maminirina, C.P., Ryan, K.E., Christidis, L. and Appleton, B. 2008b. A new species of Miniopterus (Chiroptera: Miniopteridae) from lowland southeastern Madagascar. Mammalian Biology 73:199–213.
Goodman, S.M., Maminirina, C.P., Bradman, H.M., Christidis, L. and Appleton, B. 2009. The use of molecular phylogenetic and morphological tools to identify cryptic and paraphyletic species: Examples from the diminutive long-fingered bats (Chiroptera: Miniopteridae: Miniopterus) on Madagascar. American Museum Novitates 3669:1–34.
Goodman, S.M., Maminirina, C.P., Bradman, H.M., Christidis, L. and Appleton, B.R. 2010a. [https://dx.doi.org/10.1111/j.1439-0469.2009.00524.x Patterns of morphological and genetic variation in the endemic Malagasy bat Miniopterus gleni (Chiroptera: Miniopteridae), with the description of a new species, M. griffithsi] (subscription required). Journal of Zoological Systematics and Evolutionary Research 48(1):75–86.
Goodman, S.M., Ramasindrazana, B., Maminirina, C.P., Schoeman, M.C. and Appleton, B. 2011. [http://mapress.com/zootaxa/2011/f/zt02880p019.pdf Morphological, bioacoustical, and genetic variation in Miniopterus bats from eastern Madagascar, with the description of a new species]. Zootaxa 2880:1–19.
Jenkins, R.K.B. and Rakotoarivelo, A.R. 2008a. {{IUCNlink|40039|Miniopterus majori}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Jenkins, R.K.B. and Rakotoarivelo, A.R. 2008b. {{IUCNlink|136788|Miniopterus petersoni}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Jenkins, R.K.B., Rakotoarivelo, A.R., Ratrimomanarivo, F.H. and Cardiff, S.G. 2008f. {{IUCNlink|136401|Miniopterus sororculus}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.

=Vespertilionidae=

Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008d. {{IUCNlink|136820|Eptesicus malagasyensis}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008e. {{IUCNlink|40033|Eptesicus matroka}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008i. {{IUCNlink|14163|Myotis goudoti}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Andriafidison, D, Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008k. {{IUCNlink|20064|Scotophilus borbonicus}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Andriafidison, D., Cardiff, S.G., Goodman, S.M., Hutson, A.M., Jenkins, R.K.B., Kofoky, A.F., Racey, P.A., Ranivo, J., Ratrimomanarivo, F.H. and Razafimanahaka, H.J. 2008l. {{IUCNlink|20072|Scotophilus robustus}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Bates, P.J.J., Ratrimomanarivo, F.H., Harrison, D.L. and Goodman, S.M. 2006. [https://www.researchgate.net/profile/Paul_Bates6/publication/232678520_A_description_of_a_new_species_of_Pipistrellus_Chiroptera_Vespertilionidae_from_Madagascar_with_a_review_of_related_Vespertilioninae_from_the_island/links/54aa9cd70cf2bce6aa1d509f/A-description-of-a-new-species-of-Pipistrellus-Chiroptera-Vespertilionidae-from-Madagascar-with-a-review-of-related-Vespertilioninae-from-the-island.pdf A description of a new species of Pipistrellus (Chiroptera: Vespertilionidae) from Madagascar with a review of related Vespertilioninae from the island]. Acta Chiropterologica 8(2):299–324.
Goodman S.M., Jenkins R.K.B. and Ratrimomanarivo F.H. 2005. A review of the genus Scotophilus (Mammalia, Chiroptera, Vespertilionidae) on Madagascar, with the description of a new species. Zoosystema 27(4):867–882.
Goodman, S.M., Ratrimomanarivo, F.H. and Randrianandrianina, F.H. 2006b. A new species of Scotophilus (Chiroptera: Vespertilionidae) from western Madagascar. Acta Chiropterologica 8(1):21–37.
Goodman, S.M., Taylor, P.J., Ratrimomanarivo, F. and Hoofer, S.R. 2012. [http://www.mapress.com/zootaxa/2012/2/zt03250p025.pdf The genus Neoromicia (family Vespertilionidae) in Madagascar, with the description of a new species] (subscription required). Zootaxa 3250:1–25.
Hutson, A.M., Racey, P.A., Goodman, S. and Jacobs, D. 2008b. {{IUCNlink|44923|Pipistrellus nanus}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Jacobs, D., Cotterill, F.P.D., Taylor, P.J., Monadjem, A. and Griffin, M. 2008. {{IUCNlink|44851|Pipistrellus anchietae}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Jenkins, R.K.B., Rakotoarivelo, A.R., Ratrimomanarivo, F.H. and Cardiff, S.G. 2008g. {{IUCNlink|136646|Pipistrellus raceyi}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Jenkins, R.K.B., Rakotoarivelo, A.R., Ratrimomanarivo, F.H. and Cardiff, S.G. 2008h. {{IUCNlink|136774|Scotophilus marovaza}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Jenkins, R.K.B., Rakotoarivelo, A.R., Ratrimomanarivo, F.H. and Cardiff, S.G. 2008i. {{IUCNlink|136675|Scotophilus tandrefana}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.
Mickleburgh, S., Hutson, A.M. and Racey, P.A. 2008a. {{IUCNlink|136741|Pipistrellus hesperidus}}. IUCN Red List of Threatened Species. Version 2009.2. Downloaded May 27, 2010.