class="sortable wikitable" border="1"
|+ Examples of camouflage methods in animal and military usage
! scope=col width=150px | Method
! scope=col width=150px | Poulton category
! scope=col | Terrestrial, aerial
! scope=col | Aquatic
! scope=col | Military |
valign="top" align="center"
| rowspan=2|Mimesis: resembling something not of interest to the observer | Special aggressive resemblance: mimesis by a predator to avoid scaring off prey | Flower mantis[Forbes, 2009. p. 134.] File:Mantis Hymenopus coronatus 6 Luc Viatour.jpg | Green frogfish[Cott, 1940. pp. 340–342.] File:Green Frogfish.jpg | Sunshield[Barkas, 1952. pp. 202–203.] File:IWM-E-18461-Crusader-camouflaged-19421026.jpg |
valign="top" align="center" | Special protective resemblance: resemblance to a specific object by prey to avoid detection by predators | Dead leaf butterfly File:Kallima inachus qtl1.jpg | Soft coral spider crab[Cott, 1940. p. 338.] File:Hoplophrys oatesii (Soft coral spider crab).jpg | Observation tree, 1916[{{cite web | title=Art of the First World War: André Mare and Leon Underwood | work=The Elm at Vermezeele | publisher=Memorial-Caen | year=1998 | url=http://www.memorial-caen.fr/10EVENT/EXPO1418/gb/texte/027text.html | access-date=8 February 2013 | url-status=dead | archive-url=https://web.archive.org/web/20130529001851/http://www.memorial-caen.fr/10EVENT/EXPO1418/gb/texte/027text.html | archive-date=29 May 2013 }}] File:Andre Mare Camouflaged Iron Observation Tree (The Elm at Vermezeele) 1916.jpg |
valign="top" align="center"
| Colour matching: having similar colours to the environment, also known as background matching | General protective resemblance: resembling the background in a general way | European tree frog[Beddard, 1892. p. 83.] File:Laubfrosch.jpg | Brown trout[Cott, 1940. p. 28.] File:Salmo trutta.jpg | Khaki uniforms, 1910[Newark, 2007. pp. 45–46.] File:Greek artillery, 1910.png |
valign="top" align="center"
| Disruptive coloration: having high contrast coloration that breaks up outlines, so observers fail to recognise the object | General protective resemblance (a type of) | Papuan frogmouth[Cott, 1940. p. 148.] File:Podargus papuensis - Daintree River.jpg Dominant | Commerson's frogfish[{{cite web | url=http://www.flmnh.ufl.edu/fish/gallery/Descript/StriatedFrogfish/StriatedFrogfish.html | title=Striated Frogfish | publisher=Florida Museum of Natural History | access-date=12 February 2013 | author=Bester, Cathleen | archive-url=https://web.archive.org/web/20121216095921/http://www.flmnh.ufl.edu/fish/Gallery/Descript/StriatedFrogfish/StriatedFrogfish.html | archive-date=16 December 2012 | url-status=dead }}] File:Commerson's Frogfish, Kona, Hawaii.jpg | Disruptive Pattern Material[{{cite book|author1=Blechman, Hardy |author2=Newman, Alex |year=2004|title= DPM: Disruptive Pattern Material|publisher=DPM Ltd}}] Image:British dpm2.jpg Dominant |
valign="top" align="center"
| Disruptive eye mask: a disruptive pattern that covers or runs up to the eyes, concealing them | Coincident disruptive pattern (a type of) (Cott) | Common frog[Cott, 1940. pp. 70–88.] File:2014.07.17.-28-Zadlitzgraben Pressel--Grasfrosch-Weibchen.jpg | Jack-knifefish[{{cite journal |last1=Barlow |first1=G. W. |title=The attitude of fish eye-lines in relation to body shape and to stripes and bars |journal=Copeia |date=1972 |volume=1972 |issue=1 |pages=4–12 |jstor=1442777|doi=10.2307/1442777 }}] File:Equetus lanceolatus in Madagascar Reef.jpg | Gun barrel of Sherman Firefly[{{cite book | title=Middle East AFV Technical Letter | publisher=The Tank Museum, UK; originally G(Cam) Eighth Army | date=26 January 1945}}] File:Sherman Firefly 9-08-2008 15-05-43.JPG |
valign="top" align="center"
| Seasonal variation: having coloration that varies with season, usually summer to winter | Variable general protective resemblance: having coloration that resembles the background in each season, in a general way | Arctic hare[Cott, 1940. p. 23.] File:Arctic Hare 1.jpg | Walleye[{{Cite journal |last1=Schaefer |first1=Wayne F. |last2=Schmitz |first2=Mark H. |last3=Blazer |first3=Vicki S. |last4=Ehlinger |first4=Timothy J. |last5=Berges |first5=John A. |date=2014 |title=Localization and seasonal variation of blue pigment (sandercyanin) in walleye (Sander vitreus) |journal=Canadian Journal of Fisheries and Aquatic Sciences |volume=72 |issue=2 |pages=281–289 |doi=10.1139/cjfas-2014-0139 |issn=0706-652X|doi-access=free }}]File:Sander_vitreus.jpg | Snow overalls[{{cite web | url=http://www.dijitalimaj.com/alamyDetail.aspx?img={00C97AE1-77F9-49BC-9CAD-94A91EF188A3} | title=1915 Protective colouring pyrotechnics British Soldier white overalls snow winter clothing uniform camouflage camouflaged | publisher=DijitalImaj | access-date=22 February 2013}} Original publication in "The War Illustrated a Conflict of Nations"] File:Norwegian Winter War Volunteers.jpg |
valign="top" align="center"
| Side or Thayer countershading: having graded toning from dark above to light below, so as to cancel out the apparent effect of self-shadowing when viewed from the side | — | Bushbuck (also has white distractive markings)[{{cite journal |author1=Ruxton, Graeme D |author2=Speed, Michael P |author3=Kelly, David J | title=What, if anything, is the adaptive function of countershading?| journal=Animal Behaviour | year=2004 | volume=68 |issue=3 | pages=445–451 | doi=10.1016/j.anbehav.2003.12.009|s2cid=43106264 | url=http://academic.brooklyn.cuny.edu/biology/jbasil/documents/COUNTERSHADINGCLASS10.pdf}}] File:Tragelaphus scriptus (male) cropped.jpg Dominant
| Blue shark[Cott, 1940. pp. 40–41.] File:Blue shark.jpg | Hugh Cott's guns[Forbes, 2009. pp. 149–150.] File:Countershaded Rail-mounted Gun Camouflaged by Hugh Cott 1940.jpg |
valign="top" align="center"
| Above/below countershading: having different colours or patterns above and below, to camouflage the upperside for observers from above, and the underside for observers from below | — | Gull (white underside to match sky, improves fishing success)[{{cite journal |last1=Rowland |first1=Hannah M. |title=Review From Abbott Thayer to the present day: what have we learned about the function of countershading? |journal=Philosophical Transactions of the Royal Society B |date=2009 |volume=364 |issue=1516 |pages=519–527 |doi=10.1098/rstb.2008.0261 |url=http://rstb.royalsocietypublishing.org/content/364/1516/519.full.pdf+html |pmid=19000972 |pmc=2674085}}] File:Gull in flight.jpg | Penguins[{{cite journal | title=From Abbott Thayer to the present day: what have we learned about the function of countershading? | author=Rowland, Hannah M | journal=Philosophical Transactions of the Royal Society B |date=February 2009 | volume=364 | issue=1516 | pages=519–527 | doi=10.1098/rstb.2008.0261 | pmc=2674085 | pmid=19000972}}] File:Penguins walking -Moltke Harbour, South Georgia, British overseas territory, UK-8.jpg | Supermarine Spitfire[{{cite book | title=Malta Spitfire Aces | publisher=Osprey Publishing | author=Nichols, Steve | year=2008 | page=16}}] File:Spitfire mk2a p7350 arp.jpg |
valign="top" align="center"
| Counterillumination: generating light to raise the brightness of an object to match a brighter background, as of a marine animal's underside against the sea surface | — | — | Sparkling enope squid[Herring, 2002. pp. 190–195.][{{cite web | url=http://ocean.si.edu/ocean-photos/midwater-squid-abralia-veranyi | title=Midwater Squid, Abralia veranyi | publisher=Smithsonian National Museum of Natural History | access-date=28 November 2011}}] File:Squid Counterillumination.png Dominant (100–1000m) | Yehudi lights (prototype)[{{cite book | url=http://www.dtic.mil/dtic/tr/fulltext/u2/221102.pdf | archive-url=https://web.archive.org/web/20131023061821/http://www.dtic.mil/dtic/tr/fulltext/u2/221102.pdf | url-status=dead | archive-date=October 23, 2013 | work=Visibility Studies and Some Applications in the Field of Camouflage | publisher=Office of Scientific Research and Development, National Defence Research Committee | title=Camouflage of Sea-Search Aircraft | year=1946 | access-date=12 February 2013 |last1=Bush |first1=Vannevar |author1-link=Vannevar Bush |author2=Conant, James |author3=Harrison, George | pages=225–240|display-authors=etal}}] File:Principle of Yehudi Lights with Avenger head-on view.jpg |
valign="top" align="center"
| Transparency: letting so much light through that the object is hard to see in typical lighting conditions | General protective resemblance (a type of) | Glass frogs[{{cite web |last=Naish |first=D. |title=Green-boned glass frogs, monkey frogs, toothless toads |url=http://scienceblogs.com/tetrapodzoology/2007/11/09/green-boned-glass-frogs/ |work=Tetrapod zoology |publisher=scienceblogs.com |access-date=14 February 2013}}] File:Hyalinobatrachium uranoscopum01a.jpg | Comb jellies File:Mnemiopsis leidyi.jpg Dominant (0–100m) | 1916 trials[{{cite web |title=American Art Native Americans George de Forest Brush Arapahoes |url=http://www.ebay.com/itm/American-Art-Native-Americans-George-de-Forest-Brush-Arapahoes-cm1676-/360508931653 |access-date=4 September 2015 |quote=In 1916, Brush acquired a small Morane-Borel monoplane (also known as a Morane-Saulnier). He experimented with the possibility of making its wings and fuselage transparent, to reduce its visibility.}} See also the Etrich Taube, a 1910 design whose translucency effectively camouflaged it above 1200 feet (400 metres).{{cite web |last1=Naughton |first1=Russell |title=Igo Etrich (1879 - 1967) and his 'Taube' |url=http://www.ctie.monash.edu.au/hargrave/etrich_taube.htm |publisher=Monash University |date=1 January 2002}}] File:Emile Taddéoli monoplane.jpg |
valign="top" align="center"
| Reflection (silvering): reflecting enough light, usually from the sides, to make the object show as a (reflected) patch of the environment | General protective resemblance (a type of) | — | Pilchard File:Sardina pilchardus 2011.jpg Dominant (0–100m) | — |
valign="top" align="center"
| Self-decoration: covering oneself in materials from the environment | Adventitious protection: covering oneself in materials that are not part of the body | Masked hunter bug[{{cite journal|author=Wierauch, C|year=2006|title=Anatomy of disguise: camouflaging structures in nymphs of Some Reduviidae (Heteroptera) |journal=American Museum Novitates| issue=3542| pages=1–18| doi=10.1206/0003-0082(2006)3542[1:AODCSI]2.0.CO;2|url=http://digitallibrary.amnh.org/bitstream/2246/5820/1//v3/dspace/updateIngest/pdfs/N3542.pdf|hdl=2246/5820|s2cid=7894145 }}] File:Reduvius personatus, Masked Hunter Bug nymph camouflaged with sand grains.JPG | Decorator crabs[{{cite book |author1=Hultgren, Kristin |author2=Jay Stachowicz |year=2011 |chapter=Camouflage in decorator crabs: integrating ecological, behavioural and evolutionary approaches |title=Animal Camouflage |editor1=Martin Stevens |editor1-link=Martin Stevens (biologist) |editor2=Sami Merilaita |isbn=978-0-521-19911-7 |publisher=Cambridge University Press |chapter-url=http://www-eve.ucdavis.edu/stachowicz/papers/Hultgren_Stachowicz_2011_BookChapter.pdf |access-date=2013-02-12 |archive-date=2017-08-15 |archive-url=https://web.archive.org/web/20170815183654/http://www-eve.ucdavis.edu/stachowicz/papers/Hultgren_Stachowicz_2011_BookChapter.pdf |url-status=dead }}] File:Graceful decorator crab (Oregonia gracilis) with sponge.jpg
| Ghillie suit[Forbes, 2009. pp. 102–103.] File:Marine sniper ghillie suit.JPG |
valign="top" align="center"
| Concealment of shadow: having features such as flanges or a flattened body to reduce or hide the shadow | — | Flying lizard[{{cite book | title=Nature's Treasurehouse: The Wildlife of Indonesia | publisher=Gramedia Pustaka Utama | author=MacKinnon, Kathy | year=1992 | pages=38}}] File:2005-Draco-dussumieri.jpg | Tasselled wobbegong[{{cite web | url=http://www.thedeep.co.uk/downloads/38?type=pressmedia | title=New shark at The Deep heralds summer event | publisher=The Deep | date=21 July 2011 | access-date=18 April 2013 | author=Martin, Linda | archive-date=10 August 2014 | archive-url=https://web.archive.org/web/20140810161723/http://www.thedeep.co.uk/downloads/38?type=pressmedia | url-status=dead }}] File:Eucrossorhinus dasypogon.jpg | Camouflage netting[Cott, 1940. p. 111.] File:Øvelse på Evjemoen Tropp 4.2 - camouflage nettings.jpg |
valign="top" align="center"
| Irregular outline: having a broken or complex outline (that may help delay recognition by an observer) | Special protective resemblance (types of) | Comma butterfly[Cott, 1940. p. 75.] File:Polygonia c-album LC0238.jpg | Leafy sea dragon[Cott, 1940. pp. 341–342.] File:Leafy Sea Dragon.jpg | Scrim, branches[Cott, 1940. pp. 359, 362.] File:Battle of Lake Khasan-Camouflaged soviet tanks.jpg |
valign="top" align="center"
| Distraction: having coloration that distracts an observer's attention away from a feature of the object (such as the head or eye) | — | Eyespots of peacock butterfly[{{cite journal |doi=10.1098/rspb.2004.3034 |last1=Vallin|first1=A.|first2=S. |last2=Jakobsson|first3=J. |last3=Lind|first4=C. |last4=Wiklund |title=Prey survival by predator intimidation: an experimental study of peacock butterfly defence against blue tits |journal=Proceedings of the Royal Society B: Biological Sciences |volume=272 |year=2005 |pages=1203–1207 |issue=1569 |pmid=16024383 |pmc=1564111}}] File:Inachis io qtl2.jpg | Foureye butterflyfish[Cott, 1940. p. 373.] File:Chaetodon capistratus1.jpg | False bow wave in ship camouflage[{{cite web | url=http://www.history.navy.mil/photos/sh-usn/usnsh-n/ca26.htm | title=USS Northampton (CA-26, originally CL-26), 1930–1942 | publisher=Naval Historical Center | year=2002 | access-date=13 June 2012 | url-status=dead | archive-url=https://web.archive.org/web/20120721234920/http://www.history.navy.mil/photos/sh-usn/usnsh-n/ca26.htm | archive-date=21 July 2012 }}] File:USS Northampton (CA-26) at Brisbane on 5 August 1941 (NH 94596).jpg |
valign="top" align="center"
| Distractive markings Small conspicuous marks that distract an observer's attention from recognising the object as a whole[{{cite journal |last1=Dimitrova |first1=M. |last2=Stobbe |first2=N. |last3=Schaefer |first3=H. M. |last4=Merilaita |first4=S. |title=Concealed by conspicuousness: distractive prey markings and backgrounds |journal=Proceedings of the Royal Society B: Biological Sciences |volume=276 |issue=1663 |year=2009 |pages=1905–1910 |doi=10.1098/rspb.2009.0052|pmid=19324754 |pmc=2674505 }}] | — | Snowy owl with distractively marked plumage[{{cite book |author1=Thayer, Gerald H. |author2=Thayer, Abbott H. |author2-link=Abbott Handerson Thayer |title=Concealing Coloration in the Animal Kingdom: An Exposition of the Laws of Disguise Through Color and Pattern; Being a Summary of Abbott H. Thayer's Disclosures |publisher=Macmillan |location=New York |year=1909 |url=https://archive.org/details/cu31924022546406 |pages=[https://archive.org/details/cu31924022546406/page/n291 151]–152, 246–247}}] File:Bubo scandiacus male Muskegon (cropped).jpg | — | Snow camouflage using small distractive marks |
valign="top" align="center"
| Active camouflage: changing the coloration rapidly enough to maintain resemblance to the current background while moving | Variable aggressive resemblance, variable protective resemblance: varying coloration to resemble the background, in predator and prey respectively | Veiled chameleon[Forbes, 2009. p. 236.] File:Yemen Chameleon (cropped).jpg | Octopuses[Forbes, 2009. pp. 236–239.] File:Octopus2.jpg | Adaptiv[{{cite web|url=http://www.baesystems.com/image/BAES_019603/innovation-adaptiv-car-signature |title=Innovation Adaptiv Car Signature |publisher=BAE Systems |year=2012 |access-date=12 February 2013 |url-status=dead |archive-url=https://web.archive.org/web/20130618173128/http://www.baesystems.com/image/BAES_019603/innovation-adaptiv-car-signature |archive-date=18 June 2013 }}] (see that article for image) |
valign="top" align="center"
| Motion camouflage: following a track such that the object remains between a starting point and the target (e.g. prey) at all times, rather than going straight for the target | — | Hoverfly[{{cite journal | title=Strategies for active camouflage of motion
]|author1=Srinivasan, M. V. |author2=Davey, M. | journal=Proceedings of the Royal Society B | year=1995 | volume=259 | pages=19–25 | issue=1354 | doi=10.1098/rspb.1995.0004 | bibcode=1995RSPSB.259...19S|s2cid=131341953 }} File:Syritta pipiens-pjt1.jpg | — | Air-to-air missile[{{cite journal|author1=Ghose, K |author2=Horiuchi, TK |author3=Krishnaprasad, PS |author4=Moss, CF |title=Echolocating Bats Use a Nearly Time-Optimal Strategy to Intercept Prey| journal=PLOS Biology |volume=4 |issue= 5 |page=e108 |doi=10.1371/journal.pbio.0040108| year=2006| pmid=16605303| pmc=1436025 |doi-access=free }}] File:F-15 firing AIM-7Ms.jpg Used primarily for efficiency |
valign="top" align="center"
| Motion dazzle: rapidly moving a bold pattern of contrasting stripes, confusing an observer's visual processing[{{cite journal | title=Dazzle Camouflage Affects Speed Perception |author1=Scott-Samuel, NE |author2=Baddeley, R |author3=Palmer, CE |author4=Cuthill, IC | journal=PLOS ONE |date=June 2011 | volume=6 | issue=6 | doi=10.1371/journal.pone.0020233 | editor1-last=Burr | editor1-first=David C | page=e20233 | pmid=21673797 | pmc=3105982| bibcode=2011PLoSO...620233S |doi-access=free }}][{{cite journal | title=Motion camouflage induced by zebra stripes |author1=How, Martin J. |author2=Zanker, Johannes M. | journal=Zoology | year=2014 | pages=163–70 | doi=10.1016/j.zool.2013.10.004 | pmid=24368147 | volume=117| issue=3 |url=https://espace.library.uq.edu.au/view/UQ:319813/UQ319813.pdf }}] | — | Zebra File:Chapman-Zebra.jpg | — | Proposal only
(NB: Marine Dazzle camouflage did not claim this effect) |
valign="top" align="center" | Dazzle camouflage: bold patterns of contrasting stripes, deceiving enemy about ship's heading | — | — | — | Ship camouflage, mainly WW1[{{cite book | author=Wilkinson, Norman | title=A Brush with Life | publisher=Seeley Service | year=1969 | page=79}}] File:USS West Mahomet (ID-3681) cropped.jpg Dominant 1917–18 |
valign="top" align="center" | Ultra-blackness: extremely black surface, matching very dark background | — | Black panther[{{Cite web |last1=Magazine |first1=Smithsonian |last2=Black |first2=Riley |title=Why Are Black Leopards So Rare? |url=https://www.smithsonianmag.com/science-nature/why-are-black-leopards-so-rare-180973820/ |access-date=2023-08-15 |website=Smithsonian Magazine |language=en}}]File:Black_panther_(4530714641).jpg | Deep-sea fish[{{cite journal |last1=Davis |first1=Alexander L. |last2=Thomas |first2=Kate N. |last3=Goetz |first3=Freya E. |last4=Robison |first4=Bruce H. |last5=Johnsen |first5=Sönke |last6=Osborn |first6=Karen J. |title=Ultra-black Camouflage in Deep-Sea Fishes |journal=Current Biology |year=2020 |volume=30 |issue=17 | pages=3470–3476.e3 |issn=0960-9822 |doi=10.1016/j.cub.2020.06.044|pmid=32679102 |doi-access=free }}] File:Humpback anglerfish.png|alt=Black-camouflaged deep-sea fish]] | Night fighters[Stephenson, Hubert Kirk. (1948) Applied Physics, pp. 200, 258. Science in World War II; Office of Scientific Research and Development. Volume 6 of Science in World War II (Atlantic Monthly Press Book). Editors: Chauncey Guy Suits and George Russell Harrison. Little, Brown.] File:Hurricane XII RCAF 5589.jpg |