List of camouflage methods

{{short description|List of ways of hiding objects or animals in plain sight}}

{{featured list}}

Camouflage is the concealment of animals or objects of military interest by any combination of methods that helps them to remain unnoticed. This includes the use of high-contrast disruptive patterns as used on military uniforms, but anything that delays recognition can be used as camouflage. Camouflage involves deception, whether by looking like the background or by resembling something else, which may be plainly visible to observers.Cott, 1940. Chapter 1: General Colour Resemblance. pp. 5–19.Forbes, 2009. p. 51. This article lists methods used by animals and the military to escape notice.

Conventions used

File:Antennarius striatus2.JPG, Antennarius striatus, is elaborately camouflaged for life on the subtropical ocean floor.|alt=Elaborately camouflaged frogfish on ocean floor]]

Different camouflage methods employed by terrestrial, aerial, and aquatic animals, and in military usage, are compared in the table. Several methods are often combined, so for example the Bushbuck is both countershaded over its whole body, and disruptively coloured with small pale spots. Until the discovery of countershading in the 1890s, protective coloration was considered to be mainly a matter of colour matching, but while this is certainly important, a variety of other methods are used to provide effective camouflage.

When an entry is marked Dominant, that method is used widely in that environment, in most cases. For example, countershading is very common among land animals, but not for military camouflage. The dominant camouflage methods on land are countershading and disruptive coloration, supported by less frequent usage of many other methods.Cott, 1940. Part 1: Concealment. pp. 5–190. The dominant camouflage methods in the open ocean are transparency,{{cite journal | jstor=1543609 | title=Hidden in Plain Sight: The Ecology and Physiology of Organismal Transparency | author=Johnsen, Sönke | journal=Biological Bulletin |date=December 2001 | volume=201 | issue=3 | pages=301–318 | doi=10.2307/1543609 | pmid=11751243| s2cid=6385064 }} reflection, and counterillumination.{{cite journal | title=Crypsis in the Pelagic Environment | author=McFall-Ngai, Margaret J | journal=American Zoologist | year=1990 | volume=30 | issue=1 | pages=175–188 | doi=10.1093/icb/30.1.175| doi-access=free }} Transparency and reflectivity are dominant in the top {{convert|100|m|ft}} of the ocean; counterillumination is dominant from {{convert|100|m|ft}} down to {{convert|1000|m|ft}}. Most animals of the open sea use one or more of these methods. Military camouflage relies predominantly on disruptive patterns,Newark, 2007. p. 154. though methods such as outline disruption are also used, and others have been prototyped.

In 1890 the English zoologist Edward Bagnall Poulton categorised animal colours by their uses,Poulton, 1890. Fold-out after p. 339. which cover both camouflage and mimicry.Forbes, 2009. pp. 51–52. Poulton's categories were largely followed by Hugh Cott in 1940. Relevant Poulton categories are listed in the table. Where Poulton's definition covers a method but does not name it explicitly, the category is named in parentheses.

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Comparisons

class="sortable wikitable" border="1"

|+ Examples of camouflage methods in animal and military usage

! scope=col width=150px | Method

! scope=col width=150px | Poulton
category

! scope=col | Terrestrial, aerial

! scope=col | Aquatic

! scope=col | Military

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| rowspan=2|Mimesis:
resembling something not of interest to the observer

Special aggressive resemblance:
mimesis by a predator to avoid scaring off prey
Flower mantisForbes, 2009. p. 134.
File:Mantis Hymenopus coronatus 6 Luc Viatour.jpg
Green frogfishCott, 1940. pp. 340–342.
File:Green Frogfish.jpg
SunshieldBarkas, 1952. pp. 202–203.
File:IWM-E-18461-Crusader-camouflaged-19421026.jpg
valign="top" align="center"Special protective resemblance:
resemblance to a specific object by prey to avoid detection by predators
Dead leaf butterfly
File:Kallima inachus qtl1.jpg
Soft coral spider crabCott, 1940. p. 338.
File:Hoplophrys oatesii (Soft coral spider crab).jpg
Observation tree, 1916{{cite web | title=Art of the First World War: André Mare and Leon Underwood | work=The Elm at Vermezeele | publisher=Memorial-Caen | year=1998 | url=http://www.memorial-caen.fr/10EVENT/EXPO1418/gb/texte/027text.html | access-date=8 February 2013 | url-status=dead | archive-url=https://web.archive.org/web/20130529001851/http://www.memorial-caen.fr/10EVENT/EXPO1418/gb/texte/027text.html | archive-date=29 May 2013 }}
File:Andre Mare Camouflaged Iron Observation Tree (The Elm at Vermezeele) 1916.jpg
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| Colour matching:
having similar colours to the environment, also known as background matching

General protective resemblance:
resembling the background in a general way
European tree frogBeddard, 1892. p. 83.
File:Laubfrosch.jpg
Brown troutCott, 1940. p. 28.
File:Salmo trutta.jpg
Khaki uniforms, 1910Newark, 2007. pp. 45–46.
File:Greek artillery, 1910.png
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| Disruptive coloration:
having high contrast coloration that breaks up outlines, so observers fail to recognise the object

General protective resemblance (a type of)Papuan frogmouthCott, 1940. p. 148.
File:Podargus papuensis - Daintree River.jpg
Dominant
Commerson's frogfish{{cite web | url=http://www.flmnh.ufl.edu/fish/gallery/Descript/StriatedFrogfish/StriatedFrogfish.html | title=Striated Frogfish | publisher=Florida Museum of Natural History | access-date=12 February 2013 | author=Bester, Cathleen | archive-url=https://web.archive.org/web/20121216095921/http://www.flmnh.ufl.edu/fish/Gallery/Descript/StriatedFrogfish/StriatedFrogfish.html | archive-date=16 December 2012 | url-status=dead }}
File:Commerson's Frogfish, Kona, Hawaii.jpg
Disruptive Pattern Material{{cite book|author1=Blechman, Hardy |author2=Newman, Alex |year=2004|title= DPM: Disruptive Pattern Material|publisher=DPM Ltd}}
Image:British dpm2.jpg
Dominant
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| Disruptive eye mask:
a disruptive pattern that covers or runs up to the eyes, concealing them

Coincident disruptive pattern (a type of) (Cott)Common frogCott, 1940. pp. 70–88.
File:2014.07.17.-28-Zadlitzgraben Pressel--Grasfrosch-Weibchen.jpg
Jack-knifefish{{cite journal |last1=Barlow |first1=G. W. |title=The attitude of fish eye-lines in relation to body shape and to stripes and bars |journal=Copeia |date=1972 |volume=1972 |issue=1 |pages=4–12 |jstor=1442777|doi=10.2307/1442777 }}
File:Equetus lanceolatus in Madagascar Reef.jpg
Gun barrel of
Sherman Firefly{{cite book | title=Middle East AFV Technical Letter | publisher=The Tank Museum, UK; originally G(Cam) Eighth Army | date=26 January 1945}}
File:Sherman Firefly 9-08-2008 15-05-43.JPG
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| Seasonal variation:
having coloration that varies with season, usually summer to winter

Variable general protective resemblance:
having coloration that resembles the background in each season, in a general way
Arctic hareCott, 1940. p. 23.
File:Arctic Hare 1.jpg
Walleye{{Cite journal |last1=Schaefer |first1=Wayne F. |last2=Schmitz |first2=Mark H. |last3=Blazer |first3=Vicki S. |last4=Ehlinger |first4=Timothy J. |last5=Berges |first5=John A. |date=2014 |title=Localization and seasonal variation of blue pigment (sandercyanin) in walleye (Sander vitreus) |journal=Canadian Journal of Fisheries and Aquatic Sciences |volume=72 |issue=2 |pages=281–289 |doi=10.1139/cjfas-2014-0139 |issn=0706-652X|doi-access=free }}File:Sander_vitreus.jpgSnow overalls{{cite web | url=http://www.dijitalimaj.com/alamyDetail.aspx?img={00C97AE1-77F9-49BC-9CAD-94A91EF188A3} | title=1915 Protective colouring pyrotechnics British Soldier white overalls snow winter clothing uniform camouflage camouflaged | publisher=DijitalImaj | access-date=22 February 2013}} Original publication in "The War Illustrated a Conflict of Nations"
File:Norwegian Winter War Volunteers.jpg
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| Side or Thayer countershading:
having graded toning from dark above to light below, so as to cancel out the apparent effect of self-shadowing when viewed from the side

Bushbuck (also has white distractive markings){{cite journal |author1=Ruxton, Graeme D |author2=Speed, Michael P |author3=Kelly, David J | title=What, if anything, is the adaptive function of countershading?| journal=Animal Behaviour | year=2004 | volume=68 |issue=3 | pages=445–451 | doi=10.1016/j.anbehav.2003.12.009|s2cid=43106264 | url=http://academic.brooklyn.cuny.edu/biology/jbasil/documents/COUNTERSHADINGCLASS10.pdf}}
File:Tragelaphus scriptus (male) cropped.jpg
Dominant

Blue sharkCott, 1940. pp. 40–41.
File:Blue shark.jpg
Hugh Cott's gunsForbes, 2009. pp. 149–150.
File:Countershaded Rail-mounted Gun Camouflaged by Hugh Cott 1940.jpg
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| Above/below countershading:
having different colours or patterns above and below, to camouflage the upperside for observers from above, and the underside for observers from below

Gull (white underside to match sky, improves fishing success){{cite journal |last1=Rowland |first1=Hannah M. |title=Review From Abbott Thayer to the present day: what have we learned about the function of countershading? |journal=Philosophical Transactions of the Royal Society B |date=2009 |volume=364 |issue=1516 |pages=519–527 |doi=10.1098/rstb.2008.0261 |url=http://rstb.royalsocietypublishing.org/content/364/1516/519.full.pdf+html |pmid=19000972 |pmc=2674085}}
File:Gull in flight.jpg
Penguins{{cite journal | title=From Abbott Thayer to the present day: what have we learned about the function of countershading? | author=Rowland, Hannah M | journal=Philosophical Transactions of the Royal Society B |date=February 2009 | volume=364 | issue=1516 | pages=519–527 | doi=10.1098/rstb.2008.0261 | pmc=2674085 | pmid=19000972}}
File:Penguins walking -Moltke Harbour, South Georgia, British overseas territory, UK-8.jpg
Supermarine Spitfire{{cite book | title=Malta Spitfire Aces | publisher=Osprey Publishing | author=Nichols, Steve | year=2008 | page=16}}
File:Spitfire mk2a p7350 arp.jpg
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| Counterillumination:
generating light to raise the brightness of an object to match a brighter background, as of a marine animal's underside against the sea surface

Sparkling enope squidHerring, 2002. pp. 190–195.{{cite web | url=http://ocean.si.edu/ocean-photos/midwater-squid-abralia-veranyi | title=Midwater Squid, Abralia veranyi | publisher=Smithsonian National Museum of Natural History | access-date=28 November 2011}}
File:Squid Counterillumination.png
Dominant (100–1000m)
 
Yehudi lights
(prototype){{cite book | url=http://www.dtic.mil/dtic/tr/fulltext/u2/221102.pdf | archive-url=https://web.archive.org/web/20131023061821/http://www.dtic.mil/dtic/tr/fulltext/u2/221102.pdf | url-status=dead | archive-date=October 23, 2013 | work=Visibility Studies and Some Applications in the Field of Camouflage | publisher=Office of Scientific Research and Development, National Defence Research Committee | title=Camouflage of Sea-Search Aircraft | year=1946 | access-date=12 February 2013 |last1=Bush |first1=Vannevar |author1-link=Vannevar Bush |author2=Conant, James |author3=Harrison, George | pages=225–240|display-authors=etal}}
File:Principle of Yehudi Lights with Avenger head-on view.jpg
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| Transparency:
letting so much light through that the object is hard to see in typical lighting conditions

General protective resemblance (a type of)Glass frogs{{cite web |last=Naish |first=D. |title=Green-boned glass frogs, monkey frogs, toothless toads |url=http://scienceblogs.com/tetrapodzoology/2007/11/09/green-boned-glass-frogs/ |work=Tetrapod zoology |publisher=scienceblogs.com |access-date=14 February 2013}}
File:Hyalinobatrachium uranoscopum01a.jpg
Comb jellies
File:Mnemiopsis leidyi.jpg
Dominant (0–100m)
1916 trials{{cite web |title=American Art Native Americans George de Forest Brush Arapahoes |url=http://www.ebay.com/itm/American-Art-Native-Americans-George-de-Forest-Brush-Arapahoes-cm1676-/360508931653 |access-date=4 September 2015 |quote=In 1916, Brush acquired a small Morane-Borel monoplane (also known as a Morane-Saulnier). He experimented with the possibility of making its wings and fuselage transparent, to reduce its visibility.}} See also the Etrich Taube, a 1910 design whose translucency effectively camouflaged it above 1200 feet (400 metres).{{cite web |last1=Naughton |first1=Russell |title=Igo Etrich (1879 - 1967) and his 'Taube' |url=http://www.ctie.monash.edu.au/hargrave/etrich_taube.htm |publisher=Monash University |date=1 January 2002}}
File:Emile Taddéoli monoplane.jpg
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| Reflection (silvering):
reflecting enough light, usually from the sides, to make the object show as a (reflected) patch of the environment

General protective resemblance (a type of)Pilchard
File:Sardina pilchardus 2011.jpg
Dominant (0–100m)
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| Self-decoration:
covering oneself in materials from the environment

Adventitious protection:
covering oneself in materials that are not part of the body
Masked hunter bug{{cite journal|author=Wierauch, C|year=2006|title=Anatomy of disguise: camouflaging structures in nymphs of Some Reduviidae (Heteroptera) |journal=American Museum Novitates| issue=3542| pages=1–18| doi=10.1206/0003-0082(2006)3542[1:AODCSI]2.0.CO;2|url=http://digitallibrary.amnh.org/bitstream/2246/5820/1//v3/dspace/updateIngest/pdfs/N3542.pdf|hdl=2246/5820|s2cid=7894145 }}
File:Reduvius personatus, Masked Hunter Bug nymph camouflaged with sand grains.JPG
Decorator crabs{{cite book |author1=Hultgren, Kristin |author2=Jay Stachowicz |year=2011 |chapter=Camouflage in decorator crabs: integrating ecological, behavioural and evolutionary approaches |title=Animal Camouflage |editor1=Martin Stevens |editor1-link=Martin Stevens (biologist) |editor2=Sami Merilaita |isbn=978-0-521-19911-7 |publisher=Cambridge University Press |chapter-url=http://www-eve.ucdavis.edu/stachowicz/papers/Hultgren_Stachowicz_2011_BookChapter.pdf |access-date=2013-02-12 |archive-date=2017-08-15 |archive-url=https://web.archive.org/web/20170815183654/http://www-eve.ucdavis.edu/stachowicz/papers/Hultgren_Stachowicz_2011_BookChapter.pdf |url-status=dead }}
File:Graceful decorator crab (Oregonia gracilis) with sponge.jpg
Ghillie suitForbes, 2009. pp. 102–103.
File:Marine sniper ghillie suit.JPG
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| Concealment of shadow:
having features such as flanges or a flattened body to reduce or hide the shadow

Flying lizard{{cite book | title=Nature's Treasurehouse: The Wildlife of Indonesia | publisher=Gramedia Pustaka Utama | author=MacKinnon, Kathy | year=1992 | pages=38}}
File:2005-Draco-dussumieri.jpg
Tasselled wobbegong{{cite web | url=http://www.thedeep.co.uk/downloads/38?type=pressmedia | title=New shark at The Deep heralds summer event | publisher=The Deep | date=21 July 2011 | access-date=18 April 2013 | author=Martin, Linda | archive-date=10 August 2014 | archive-url=https://web.archive.org/web/20140810161723/http://www.thedeep.co.uk/downloads/38?type=pressmedia | url-status=dead }}
File:Eucrossorhinus dasypogon.jpg
Camouflage nettingCott, 1940. p. 111.
File:Øvelse på Evjemoen Tropp 4.2 - camouflage nettings.jpg
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| Irregular outline:
having a broken or complex outline (that may help delay recognition by an observer)

Special protective resemblance (types of)Comma butterflyCott, 1940. p. 75.
File:Polygonia c-album LC0238.jpg
Leafy sea dragonCott, 1940. pp. 341–342.
File:Leafy Sea Dragon.jpg
Scrim, branchesCott, 1940. pp. 359, 362.
File:Battle of Lake Khasan-Camouflaged soviet tanks.jpg
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| Distraction:
having coloration that distracts an observer's attention away from a feature of the object (such as the head or eye)

Eyespots of
peacock butterfly{{cite journal |doi=10.1098/rspb.2004.3034 |last1=Vallin|first1=A.|first2=S. |last2=Jakobsson|first3=J. |last3=Lind|first4=C. |last4=Wiklund |title=Prey survival by predator intimidation: an experimental study of peacock butterfly defence against blue tits |journal=Proceedings of the Royal Society B: Biological Sciences |volume=272 |year=2005 |pages=1203–1207 |issue=1569 |pmid=16024383 |pmc=1564111}}
File:Inachis io qtl2.jpg
Foureye butterflyfishCott, 1940. p. 373.
File:Chaetodon capistratus1.jpg
False bow wave in
ship camouflage{{cite web | url=http://www.history.navy.mil/photos/sh-usn/usnsh-n/ca26.htm | title=USS Northampton (CA-26, originally CL-26), 1930–1942 | publisher=Naval Historical Center | year=2002 | access-date=13 June 2012 | url-status=dead | archive-url=https://web.archive.org/web/20120721234920/http://www.history.navy.mil/photos/sh-usn/usnsh-n/ca26.htm | archive-date=21 July 2012 }}
File:USS Northampton (CA-26) at Brisbane on 5 August 1941 (NH 94596).jpg
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| Distractive markings
Small conspicuous marks that distract an observer's attention from recognising the object as a whole{{cite journal |last1=Dimitrova |first1=M. |last2=Stobbe |first2=N. |last3=Schaefer |first3=H. M. |last4=Merilaita |first4=S. |title=Concealed by conspicuousness: distractive prey markings and backgrounds |journal=Proceedings of the Royal Society B: Biological Sciences |volume=276 |issue=1663 |year=2009 |pages=1905–1910 |doi=10.1098/rspb.2009.0052|pmid=19324754 |pmc=2674505 }}

Snowy owl with distractively marked plumage{{cite book |author1=Thayer, Gerald H. |author2=Thayer, Abbott H. |author2-link=Abbott Handerson Thayer |title=Concealing Coloration in the Animal Kingdom: An Exposition of the Laws of Disguise Through Color and Pattern; Being a Summary of Abbott H. Thayer's Disclosures |publisher=Macmillan |location=New York |year=1909 |url=https://archive.org/details/cu31924022546406 |pages=[https://archive.org/details/cu31924022546406/page/n291 151]–152, 246–247}}
File:Bubo scandiacus male Muskegon (cropped).jpg
Snow camouflage using small distractive marks
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| Active camouflage:
changing the coloration rapidly enough to maintain resemblance to the current background while moving

Variable aggressive resemblance, variable protective resemblance:
varying coloration to resemble the background, in predator and prey respectively
Veiled chameleonForbes, 2009. p. 236.
File:Yemen Chameleon (cropped).jpg
OctopusesForbes, 2009. pp. 236–239.
File:Octopus2.jpg
Adaptiv{{cite web|url=http://www.baesystems.com/image/BAES_019603/innovation-adaptiv-car-signature |title=Innovation Adaptiv Car Signature |publisher=BAE Systems |year=2012 |access-date=12 February 2013 |url-status=dead |archive-url=https://web.archive.org/web/20130618173128/http://www.baesystems.com/image/BAES_019603/innovation-adaptiv-car-signature |archive-date=18 June 2013 }}
(see that article for image)
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| Motion camouflage:
following a track such that the object remains between a starting point and the target (e.g. prey) at all times, rather than going straight for the target

Hoverfly{{cite journal | title=Strategies for active camouflage of motion

|author1=Srinivasan, M. V. |author2=Davey, M. | journal=Proceedings of the Royal Society B | year=1995 | volume=259 | pages=19–25 | issue=1354 | doi=10.1098/rspb.1995.0004 | bibcode=1995RSPSB.259...19S|s2cid=131341953 }}
File:Syritta pipiens-pjt1.jpg

Air-to-air missile{{cite journal|author1=Ghose, K |author2=Horiuchi, TK |author3=Krishnaprasad, PS |author4=Moss, CF |title=Echolocating Bats Use a Nearly Time-Optimal Strategy to Intercept Prey| journal=PLOS Biology |volume=4 |issue= 5 |page=e108 |doi=10.1371/journal.pbio.0040108| year=2006| pmid=16605303| pmc=1436025 |doi-access=free }}
File:F-15 firing AIM-7Ms.jpg
Used primarily for efficiency
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| Motion dazzle:
rapidly moving a bold pattern of contrasting stripes, confusing an observer's visual processing{{cite journal | title=Dazzle Camouflage Affects Speed Perception |author1=Scott-Samuel, NE |author2=Baddeley, R |author3=Palmer, CE |author4=Cuthill, IC | journal=PLOS ONE |date=June 2011 | volume=6 | issue=6 | doi=10.1371/journal.pone.0020233 | editor1-last=Burr | editor1-first=David C | page=e20233 | pmid=21673797 | pmc=3105982| bibcode=2011PLoSO...620233S |doi-access=free }}{{cite journal | title=Motion camouflage induced by zebra stripes |author1=How, Martin J. |author2=Zanker, Johannes M. | journal=Zoology | year=2014 | pages=163–70 | doi=10.1016/j.zool.2013.10.004 | pmid=24368147 | volume=117| issue=3 |url=https://espace.library.uq.edu.au/view/UQ:319813/UQ319813.pdf }}

Zebra
File:Chapman-Zebra.jpg
Proposal only

(NB: Marine
Dazzle camouflage
did not claim
this effect)
valign="top" align="center"Dazzle camouflage:
bold patterns of contrasting stripes, deceiving enemy about ship's heading
Ship camouflage, mainly WW1{{cite book | author=Wilkinson, Norman | title=A Brush with Life | publisher=Seeley Service | year=1969 | page=79}}
File:USS West Mahomet (ID-3681) cropped.jpg
Dominant 1917–18
valign="top" align="center"Ultra-blackness:
extremely black surface, matching very dark background
Black panther{{Cite web |last1=Magazine |first1=Smithsonian |last2=Black |first2=Riley |title=Why Are Black Leopards So Rare? |url=https://www.smithsonianmag.com/science-nature/why-are-black-leopards-so-rare-180973820/ |access-date=2023-08-15 |website=Smithsonian Magazine |language=en}}File:Black_panther_(4530714641).jpgDeep-sea fish{{cite journal |last1=Davis |first1=Alexander L. |last2=Thomas |first2=Kate N. |last3=Goetz |first3=Freya E. |last4=Robison |first4=Bruce H. |last5=Johnsen |first5=Sönke |last6=Osborn |first6=Karen J. |title=Ultra-black Camouflage in Deep-Sea Fishes |journal=Current Biology |year=2020 |volume=30 |issue=17 | pages=3470–3476.e3 |issn=0960-9822 |doi=10.1016/j.cub.2020.06.044|pmid=32679102 |doi-access=free }}
File:Humpback anglerfish.png|alt=Black-camouflaged deep-sea fish]]
Night fightersStephenson, Hubert Kirk. (1948) Applied Physics, pp. 200, 258. Science in World War II; Office of Scientific Research and Development. Volume 6 of Science in World War II (Atlantic Monthly Press Book). Editors: Chauncey Guy Suits and George Russell Harrison. Little, Brown.
File:Hurricane XII RCAF 5589.jpg

References

{{reflist|33em}}

Bibliography

  • Barkas, Geoffrey; Barkas, Natalie (1952). The Camouflage Story (from Aintree to Alamein). Cassell.
  • Beddard, Frank Evers (1892). Animal Coloration: an account of the principal facts and theories relating to the colours and markings of animals. Swan Sonnenschein.
  • Cott, Hugh (1940). Adaptive Coloration in Animals. Oxford University Press.
  • Forbes, Peter (2009). Dazzled and Deceived: Mimicry and Camouflage. Yale. {{ISBN|978-0-300-12539-9}}.
  • Herring, Peter (2002). The Biology of the Deep Ocean. Oxford University Press. {{ISBN|978-0-198-54956-7}}.
  • Newark, Tim (2007). Camouflage. Thames and Hudson. {{ISBN|978-0-500-51347-7}}.
  • Poulton, Edward Bagnall (1890). The Colours of Animals: their meaning and use especially considered in the case of insects. Kegan Paul, Trench, Trübner.
  • Stevens, Martin; Merilaita, Sami (2011). Animal Camouflage. Cambridge University Press. {{ISBN|978-0-521-19911-7}}.
  • Wickler, Wolfgang (1968). Mimicry in plants and animals. McGraw-Hill. {{ISBN|978-1-114-82438-6}}

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{{DEFAULTSORT:Camouflage methods}}

Methods

Category:Biology-related lists

Category:Military lists