Lycopodium clavatum

Lycopodium clavatum is a spore-bearing vascular plant, growing mainly prostrate along the ground with stems up to {{convert|1|m|in|abbr=on}} long; the stems are much branched, and densely clothed with small, spirally arranged microphyll leaves. The leaves are 3–5 mm long and 0.7–1 mm broad, tapered to a fine hair-like white point. The branches bearing strobili or spore cones turn erect, reaching {{convert|5|–|15|cm|in|abbr=on}} above ground, and their leaves are modified as sporophylls that enclose the spore capsules or sporangia. The spore cones are yellow-green, {{convert|2|–|3|cm|in|abbr=on}} long, and {{convert|5|mm|in|abbr=on}} broad. The horizontal stems produce roots at frequent intervals along their length, allowing the stem to grow indefinitely along the ground. The stems superficially resemble small seedlings of coniferous trees, though it is not related to these.

File:LycopodiumClavatum.jpg]]

Lycopodium clavatum has a widespread distribution across several continents.{{Cite web|url=http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=200002751|title=Lycopodium clavatum in Flora of North America @ efloras.org|website=www.efloras.org|access-date=2019-05-29}}{{Cite web|url=http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200002751|title=Lycopodium clavatum in Flora of China @ efloras.org|website=www.efloras.org|access-date=2019-05-29}}[http://luirig.altervista.org/flora/taxa/index1.php?scientific-name=lycopodium+clavatum Altervista Flora Italiana, Lycopodium clavatum L.] includes photos and European distribution mapJørgensen, P. M., M. H. Nee & S. G. Beck. (eds.) 2014. Catálogo de las plantas vasculares de Bolivia, Monographs in systematic botany from the Missouri Botanical Garden 127(1–2): i–viii, 1–1744. Missouri Botanical Garden Press, St. LouisMickel, J. T. & J. M. Beitel. 1988. Pteridophyte Flora of Oaxaca, Mexico. Memoirs of The New York Botanical Garden 46: 1–568Gibbs Russell, G. E., W. G. M. Welman, E. Retief, K. L. Immelman, G. Germishuizen, B. J. Pienaar, M. Van Wyk & A. Nicholas. 1987. List of species of southern African plants. Memoirs of the Botanical Survey of South Africa 2(1–2): 1–152(pt. 1), 1–270(pt. 2). There are distinct subspecies and varieties in different parts of its range:

• Lycopodium clavatum subsp. clavatum
• Lycopodium clavatum subsp. clavatum var. clavatum (Europe, Asia, North America)
• Lycopodium clavatum subsp. clavatum var. aristatum (Mexico, Caribbean, Central America, northern South America south to northern Argentina)
• Lycopodium clavatum subsp. clavatum var. asiaticum (Japan, northeast China)
• Lycopodium clavatum subsp. clavatum var. borbonicum (central and southern Africa)
• Lycopodium clavatum subsp. clavatum var. kiboanum (mountains of tropical Africa)
• Lycopodium clavatum subsp. contiguum (southern Central America, northern South America; syn. Lycopodium contiguum)

Although globally widespread, like many clubmosses, it is confined to undisturbed sites, disappearing from farmed areas and sites with regular burning. As a result, it is endangered in many areas. In the UK it is one of 101 species named as a high priority for conservation by the wild plant charity Plantlife.

;Other common names

Common names for this species include wolf's-foot clubmoss, common clubmoss, wolf-paw clubmoss, running ground-pine, running pine, running moss, princess pine, and others.

;Use

The dried spores of this moss are explosive if present at high density in air. They were used as flash powder in early photography and are still used in magic acts. See lycopodium powder.

The plant has been used in Finnish traditional medicine as a diuretic and as a remedy for rickets.{{cite book |last1=Piirainen |first1=Mikko |last2=Piirainen |first2=Pirkko |last3=Vainio |first3=Hannele |title=Kotimaan luonnonkasvit |trans-title=Native wild plants |date=1999 |isbn=951-0-23001-4 |page=18 |language=fi |location=Porvoo, Finland |publisher=WSOY}}

;Active constituents

Bioactive secondary metabolites in clubmosses include triterpenoids with acetylcholinesterase inhibitor activity isolated from this species.Rollinger JM, Ewelt J, Seger C, Sturm S, Ellmerer EP, Stuppner H (2005). Planta Med;71(11):1040-3. {{PMID|16320206}}

Image:Lycopodium clavatum clavatum2.jpg|L. clavatum ssp. clavatum var. clavatum, with strobili

Image:Lycopodium_clavatum.jpg|L. clavatum with strobili

{{Reflist|2}}

{{Commons|Lycopodium clavatum}}

• [http://www.rook.org/earl/bwca/nature/ferns/lycopodclav-WIP.html Lycopodium clavatum, running ground pine, Flora, fauna, earth, and sky...The natural history of the northwoods]
• [https://web.archive.org/web/20050330163625/http://www.bioimages.org.uk/html/T52435.HTM Bioimagesphotos]
• [http://ucjeps.berkeley.edu/cgi-bin/get_JM_treatment.pl?61,64,65 Jepson Manual Treatment, University of California]
• [http://calphotos.berkeley.edu/cgi/img_query?query_src=photos_index&where-taxon=Lycopodium+clavatum Calphotos Photo gallery, University of California]
• [http://www.tropicos.org/ImageFullView.aspx?imageid=100129998 photo of herbarium specimen at Missouri Botanical Garden, collected in Dominican Republic in 1967]

{{Taxonbar|from=Q466426}}

Category:Lycopodiaceae

Category:Medicinal plants

Category:Flora of Asia

Category:Flora of Africa

Category:Flora of Northern America

Category:Flora of the Caribbean

Category:Flora of Southern America

Category:Plants described in 1753

Category:Taxa named by Carl Linnaeus