Mahenge Formation

{{short description|Geological formation in Tanzania}}

{{Infobox rockunit|name=Mahenge Formation|image=|caption=|type=Sedimentary|age={{fossilrange|46.3|45.7|Middle Eocene (Lutetian)}}|prilithology=Mudstone, shale|otherlithology=|namedfor=Mahenge|namedby=|region=|country=Tanzania|coordinates={{coord|4.8|S|34.3|E|display=inline,title}}|unitof=|subunits=|underlies=|overlies=Precambrian basement (Nyanzian Belt)|thickness=|extent=|area=|map={{Location map+ | Tanzania

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{{Location map~ | Tanzania

| lat_deg = -4.8

| lon_deg = 34.3

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}}|map_caption=|period=Lutetian|paleocoordinates={{coord|18.0|S|33.0|E|display=inline}}}}The Mahenge Formation{{Cite journal |last1=McMahan |first1=Caleb D. |last2=Chakrabarty |first2=Prosanta |last3=Sparks |first3=John S. |last4=Smith |first4=W. M. Leo |last5=Davis |first5=Matthew P. |date=2013 |title=Temporal patterns of diversification across global cichlid biodiversity (Acanthomorpha: Cichlidae) |journal=PLOS ONE |volume=8 |issue=8 |pages=e71162 |doi=10.1371/journal.pone.0071162 |doi-access=free |issn=1932-6203 |pmc=3747193 |pmid=23990936|bibcode=2013PLoSO...871162M }} is a Middle Eocene-aged geological formation and Konservat-Lagerstätte from Tanzania.{{Cite journal |last1=Uhl |first1=Dieter |last2=Wuttke |first2=Michael |last3=Smith |first3=Krister T. |last4=Wedmann |first4=Sonja |last5=Lehmann |first5=Thomas |date=2024-12-01 |title=Pre-Quaternary maar lakes/volcanogenic lakes as Konservat Lagerstätten—Messel and beyond |url=https://link.springer.com/article/10.1007/s12549-024-00636-z |journal=Palaeobiodiversity and Palaeoenvironments |language=en |volume=104 |issue=4 |pages=753–761 |doi=10.1007/s12549-024-00636-z |bibcode=2024PdPe..104..753U |issn=1867-1608|url-access=subscription }} It contains a high variety of exceptionally-preserved fossils, including many of the earliest fossil records of dominant freshwater fish groups found in Africa today. It represents one of the most comprehensive Paleogene-aged African fossil deposits south of the Sahara. Based on fossil plant remains found in the formation, the Mahenge lake was situated within legume-dominated woodlands reminiscent of modern miombo forests.{{Cite journal |last1=Kaiser |first1=Thomas M. |last2=Ansorge |first2=Jörg |last3=Arratia |first3=Gloria |last4=Bullwinkel |first4=Volker |last5=Gunnell |first5=Gregg F. |last6=Herendeen |first6=Patrick S. |last7=Jacobs |first7=Bonnie |last8=Mingram |first8=Jens |last9=Msuya |first9=Charles |last10=Musolff |first10=Andreas |last11=Naumann |first11=Rudolf |last12=Schulz |first12=Ellen |last13=Wilde |first13=Volker |date=2006-09-01 |title=The maar lake of Mahenge (Tanzania) unique evidence of Eocene terrestrial environments in sub-Sahara Africa |url=http://www.schweizerbart.de/papers/zdgg/detail/157/55495/The_maar_lake_of_Mahenge_Tanzania_unique_evidence_?af=crossref |journal=Zeitschrift der Deutschen Gesellschaft für Geowissenschaften |language=en |volume=157 |issue=3 |pages=411–431 |doi=10.1127/1860-1804/2006/0157-0411 |bibcode=2006ZDGG..157..411K |issn=1860-1804|url-access=subscription }}

It was deposited within a former maar lake formed from a volcanic intrusion into the Precambrian bedrock via a kimberlite pipe.{{Citation |last1=Harrison |first1=Terry |title=Paleontological Investigations at the Eocene Locality of Mahenge in North-Central Tanzania, East Africa |date=2001 |work=Eocene Biodiversity: Unusual Occurrences and Rarely Sampled Habitats |pages=39–74 |editor-last=Gunnell |editor-first=Gregg F. |url=https://link.springer.com/chapter/10.1007/978-1-4615-1271-4_2 |access-date=2025-05-02 |place=Boston, MA |publisher=Springer US |language=en |doi=10.1007/978-1-4615-1271-4_2 |isbn=978-1-4615-1271-4 |last2=Msuya |first2=Charles P. |last3=Murray |first3=Alison M. |last4=Jacobs |first4=Bonnie Fine |last5=Báez |first5=Ana Maria |last6=Mundil |first6=Roland |last7=Ludwig |first7=Kenneth R.|url-access=subscription }} It has been dated to the Lutetian, roughly 46.0 million years ago. Unusually, relatively few fossil insect remains are known from the lake, unlike with other maar deposits. It is thought that the high diversity of fish in the lake may have led to most insects being consumed before they could be fossilized.

Paleobiota

= Bony fish =

class="wikitable" align="center"

!Genus

!Species

!Material

!Notes

!Images

Chaulopareion

|C. mahengeense

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|An osteoglossid.{{Cite journal |last1=MURRAY |first1=ALISON M. |last2=WILSON |first2=MARK V. H. |date=2005-06-01 |title=Description of a new Eocene osteoglossid fish and additional information on †Singida jacksonoides Greenwood and Patterson, 1967 (Osteoglossomorpha), with an assessment of their phylogenetic relationships |url=https://academic.oup.com/zoolinnean/article-abstract/144/2/213/2627644?redirectedFrom=fulltext |journal=Zoological Journal of the Linnean Society |volume=144 |issue=2 |pages=213–228 |doi=10.1111/j.1096-3642.2005.00170.x |issn=0024-4082}}

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Chrysichthys

|C. mahengeensis

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|A claroteid catfish.{{Cite journal |last1=Murray |first1=Alison M |last2=Budney |first2=Lisa A |date=2003 |title=A new species of catfish (Claroteidae, Chrysichthys) from an Eocene crater lake in East Africa |url=https://cdnsciencepub.com/doi/abs/10.1139/e03-031 |journal=Canadian Journal of Earth Sciences |volume=40 |issue=7 |pages=983–993 |doi=10.1139/e03-031 |bibcode=2003CaJES..40..983M |issn=0008-4077|url-access=subscription }}

|frameless

Eocitharinus

|E. macrognathus

|

|A citharinid.{{Cite journal |last=Murray |first=Alison M. |date=2003 |title=A new Eocene citharinoid fish (Ostariophysi: Characiformes) from Tanzania |url=https://www.tandfonline.com/doi/abs/10.1671/1854 |journal=Journal of Vertebrate Paleontology |volume=23 |issue=3 |pages=501–507 |doi=10.1671/1854 |bibcode=2003JVPal..23..501M |issn=0272-4634|url-access=subscription }}

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Mahengecharax

|M. carrolli

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|A potential alestid characiform, taxonomic identity disputed.{{Cite journal |last=Murray |first=Alison M |date=2003 |title=A new characiform fish (Teleostei: Ostariophysi) from the Eocene of Tanzania |url=https://cdnsciencepub.com/doi/abs/10.1139/e02-108 |journal=Canadian Journal of Earth Sciences |volume=40 |issue=4 |pages=473–481 |doi=10.1139/e02-108 |bibcode=2003CaJES..40..473M |issn=0008-4077|url-access=subscription }}{{Cite journal |last1=ZANATA |first1=ANGELA M. |last2=VARI |first2=RICHARD P. |date=2005-09-01 |title=The family Alestidae (Ostariophysi, Characiformes): a phylogenetic analysis of a trans-Atlantic clade |url=https://academic.oup.com/zoolinnean/article-abstract/145/1/1/2627449?redirectedFrom=fulltext |journal=Zoological Journal of the Linnean Society |volume=145 |issue=1 |pages=1–144 |doi=10.1111/j.1096-3642.2005.00183.x |issn=0024-4082|url-access=subscription }}

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rowspan="5" |Mahengechromis

|M. brachycranium

|

| rowspan="5" |A cichlid, forming a species flock.{{Cite journal |last=Murray |first=Alison M. |date=2001-04-07 |title=The oldest fossil cichlids (Teleostei: Perciformes): indication of a 45 million-year-old species flock |journal=Proceedings of the Royal Society of London. Series B: Biological Sciences |volume=268 |issue=1468 |pages=679–684 |doi=10.1098/rspb.2000.1570 |pmc=1088656 |pmid=11321055}}{{Cite web |last=Murray |first=Alison M. |others=Robert L. (Supervisor) Carroll |title=The Eocene cichlids (Perciformes:Labroidei) of Mahenge, Tanzania / |url=https://escholarship.mcgill.ca/concern/theses/d791sh906 |access-date=2025-04-22 |website=escholarship.mcgill.ca}}

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M. curvifrons

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M. ellipticus

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M. plethos

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M. rotundus

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Mahengichthys

|M. singidaensis

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|A shellear.{{cite book |last1=Davis |first1=M.P. |title=Mesozoic Fishes 5 - Global Diversity and Evolution |last2=Arratia |first2=G. |last3=Kaiser |first3=T.M. |publisher=F. Pfeil |year=2013 |isbn=978-3-89937-159-8 |pages=325–362 |chapter=The first fossil shellear (Gonorynchiformes: Kneriidae) from the Eocene lake of Mahenge (Tanzania)}}

|

Palaeodenticeps

|P. tanganikae

|

|A relative of the denticle herring.

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Singida

|S. jacksonoides

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|A singidid osteoglossiform.{{Cite journal |last1=MURRAY |first1=ALISON M. |last2=WILSON |first2=MARK V. H. |date=2005-06-01 |title=Description of a new Eocene osteoglossid fish and additional information on †Singida jacksonoides Greenwood and Patterson, 1967 (Osteoglossomorpha), with an assessment of their phylogenetic relationships |url=https://academic.oup.com/zoolinnean/article-abstract/144/2/213/2627644?redirectedFrom=fulltext |journal=Zoological Journal of the Linnean Society |volume=144 |issue=2 |pages=213–228 |doi=10.1111/j.1096-3642.2005.00170.x |issn=0024-4082}}

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= Amphibians =

class="wikitable" align="center"

!Genus

!Species

!Material

!Notes

!Images

Singidella

|S. laticostata

|

|A pipid frog.{{Cite journal |last1=Báez |first1=Ana Maria |last2=Harrison |first2=Terry |date=2005 |title=A New Pipine Frog from an Eocene Crater Lake in North-Central Tanzania |url=https://onlinelibrary.wiley.com/doi/abs/10.1111/j.1475-4983.2005.00477.x |journal=Palaeontology |language=en |volume=48 |issue=4 |pages=723–737 |doi=10.1111/j.1475-4983.2005.00477.x |bibcode=2005Palgy..48..723B |issn=1475-4983}}

|frameless

= Mammals =

class="wikitable" align="center"

!Genus

!Species

!Material

!Notes

!Images

Tanzanycteris

|T. mannardi

|

|A tanzanycterid bat.{{Cite web |title=OLDEST PLACENTAL MAMMAL FROM SUB-SAHARAN AFRICA: EOCENE MICROBAT FROM TANZANIA - EVIDENCE FOR EARLY EVOLUTION OF SOPHISTICATED ECHOLOCATION by Gregg F. Gunnell, Bonnie Fine Jacobs, Patrick S. Herendeen, Jason J. Head, Elizabeth Kowalski, Charles P. Msuya, Ferdinand A. Mizambwa, Terry Harrison, Jörg Habersetzer, and Gerhard Storch |url=https://palaeo-electronica.org/2002_2/africa/issue2_02.htm |access-date=2025-04-22 |website=palaeo-electronica.org}}

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= Insects =

class="wikitable" align="center"

!Genus

!Species

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!Images

Mahengea

|M. mckayi

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|A scarab beetle.{{Cite journal |last1=Strümpher |first1=Werner P. |last2=Scholtz |first2=Clarke H. |last3=Schlüter |first3=Thomas |date=2023-03-29 |title=An Eocene fossil scarab beetle (Coleoptera: Scarabaeoidea) from Tanzania |url=https://sajs.co.za/article/view/14681 |journal=South African Journal of Science |volume=119 |issue=3/4 |doi=10.17159/sajs.2023/14681 |issn=1996-7489|doi-access=free |hdl=2263/90265 |hdl-access=free }}

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= Plants =

class="wikitable" align="center"

!Genus

!Species

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!Images

Acacia

|A. mahengensis

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|An acacia.{{Cite journal |last1=Herendeen |first1=P. S. |last2=Jacobs |first2=B. F. |date=2000 |title=Fossil legumes from the Middle Eocene (46.0 Ma) Mahenge Flora of Singida, Tanzania |url=https://pubmed.ncbi.nlm.nih.gov/10991905 |journal=American Journal of Botany |volume=87 |issue=9 |pages=1358–1366 |doi=10.2307/2656727 |jstor=2656727 |issn=0002-9122 |pmid=10991905|bibcode=2000AmJB...87.1358H }}

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Aphanocalyx

|A. singidaensis

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|A legume.

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cf. Cynometra

|C. sp.

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|A legume.

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References