Maleriraptor
{{Short description|Genus of herrerasaurian dinosaurs}}
{{Speciesbox
| fossil_range = Late Triassic, {{fossilrange|Norian|Norian|early Norian}}
| image = Maleriraptor (holotype, skeletal reconstruction).jpg
| image_caption = Reconstructed skeleton with known material colored (a) and holotype specimen (b)
| display_parents = 2
| genus = Maleriraptor
| parent_authority = Ezcurra et al., 2025
| species = kuttyi
| authority = Ezcurra et al., 2025
}}
Maleriraptor is an extinct genus of herrerasaurian saurischian dinosaurs from the Late Triassic (Norian) Upper Maleri Formation of India. The genus contains a single species, M. kuttyi, known from a partial skeleton.
Discovery and naming
File:Maleriraptor (geography and stratigraphy).jpg (a) and stratigraphic occurence (b) of Maleriraptor]]
The Maleriraptor holotype specimen, ISIR 282, was collected at some point prior to 1985 from outcrops of the Upper Maleri Formation of the Pranhita–Godavari Basin near Annaram in south-central India. The specimen consists of the first {{dinogloss|sacral|sacral vertebra}}, part of the second sacral rib, a vertebra representing a caudosacral element or the first in the {{dinogloss|caudals|caudal}} series, an anterior (toward the front) caudal vertebra, the right {{dinogloss|ilium}}, both ends of the right {{dinogloss|pubis}}, and part of the left pubis. ISIR 282 is accessioned at the Indian Statistical Institute.{{Cite journal |last1=Ezcurra |first1=Martín D. |last2=Garcia |first2=Maurício Silva |last3=Novas |first3=Fernando E. |last4=Müller |first4=Rodrigo Temp |last5=Agnolín |first5=Federico L. |last6=Chatterjee |first6=Sankar |date=2025-05-07 |title=A new herrerasaurian dinosaur from the Upper Triassic Upper Maleri Formation of south-central India |journal=Royal Society Open Science |language=en |volume=12 |issue=5 |doi=10.1098/rsos.250081 |issn=2054-5703 |doi-access=free|pmid=40370605 |pmc=12077243 }}
ISIR 282 was first described in a 2011 publication by Novas et al. reporting several new dinosauriform specimens from the Upper Maleri Formation. They noted that exhibited a unique character combination that distinguished it from related species. However, the authors refrained from naming the specimen as it could not be compared with Alwalkeria, a similarly-aged saurischian from the slightly older Lower Maleri Formation.{{Cite journal |last1=Novas |first1=Fernando E. |last2=Ezcurra |first2=Martin D. |last3=Chatterjee |first3=Sankar |last4=Kutty |first4=T. S. |date=September 2010 |title=New dinosaur species from the Upper Triassic Upper Maleri and Lower Dharmaram formations of Central India |journal=Earth and Environmental Science Transactions of the Royal Society of Edinburgh |language=en |volume=101 |issue=3–4 |pages=333–349 |doi=10.1017/S1755691011020093 |bibcode=2010EESTR.101..333N |issn=1755-6910}}
File:Maleriraptor kuttyi.png]]
In 2025, Ezcurra and colleagues described Maleriraptor kuttyi as a new genus and species of early saurischian dinosaurs based on these fossil remains. The generic name, Maleriraptor, combines a reference to the discovery of the holotype in the Upper Maleri Formation with the Latin word raptor, meaning "thief". The specific name, kuttyi, honors Tharavat S. Kutty, the discoverer and initial describer of the holotype.
Ezcurra et al. noted that a lack of overlapping material between Maleriraptor and Alwalkeria still precluded comparisons between the two and that the uncertain affinities of the latter did little to clarify their distinction. Regardless, they opted to name Maleriraptor kuttyi, anticipating species-level differences between the Upper and Lower Maleri formations and hoping that future discoveries of more complete Alwalkeria specimens could allow for an improved understanding of the relationships of these taxa.
Classification
File:Upper Maleri Formation dinosaurs (Maleriraptor and Jaklapallisaurus).jpg]]
In their initial description of ISIR 282, Novas et al. (2011) reported it as belonging to an indeterminate dinosauriform; their phylogenetic analysis including the specimen recovered it in a polytomy with Silesaurus, Ornithischia, Theropoda, Sauropodomorpha, and three herrerasaurs. They argued that the open {{dinogloss|acetabulum}} and straight shaft of the pubis indicated a position more derived than silesaurids and that several anatomical characters excluded it from sauropodomorphs, but regarded its phylogenetic placement as "highly problematic". In 2021, Novas et al. reported ISIR 282 as belonging to an indeterminate herrerasaur without detailed discussion.{{Cite journal |last1=Novas |first1=Fernando E. |last2=Agnolin |first2=Federico L. |last3=Ezcurra |first3=Martín D. |last4=Temp Müller |first4=Rodrigo |last5=Martinelli |first5=Agustín G. |last6=Langer |first6=Max C. |date=October 2021 |title=Review of the fossil record of early dinosaurs from South America, and its phylogenetic implications |journal=Journal of South American Earth Sciences |language=en |volume=110 |pages=103341 |doi=10.1016/j.jsames.2021.103341|bibcode=2021JSAES.11003341N }}
To more accurately determine the relationships of Maleriraptor, Ezcurra et al. (2025) scored it an updated version of the Ezcurra et al. (2023){{Cite journal |last1=Ezcurra |first1=Martín D. |last2=Marke |first2=Daniel |last3=Walsh |first3=Stig A. |last4=Brusatte |first4=Stephen L. |date=2023-11-20 |title=A revision of the 'coelophysoid-grade' theropod specimen from the Lower Jurassic of the Isle of Skye (Scotland ) |journal=Scottish Journal of Geology |language=en |volume=59 |issue=1–2 |doi=10.1144/sjg2023-012 |bibcode=2023ScJG...59...12E |issn=0036-9276|url=https://www.pure.ed.ac.uk/ws/files/384816568/Ezcurra_et_al_MS_R1.pdf }} phylogenetic dataset. This matrix placed Maleriraptor as an early-diverging member of the saurischian clade Herrerasauria. Scoring it in the Garcia et al. (2024){{Cite journal |last1=Garcia |first1=Mauricio Silva |last2=Cabreira |first2=Sérgio Furtado |last3=da Silva |first3=Lúcio Roberto |last4=Pretto |first4=Flávio Augusto |last5=Müller |first5=Rodrigo Temp |date=2024-11-16 |title=A saurischian (Archosauria, Dinosauria) ilium from the Upper Triassic of southern Brazil and the rise of Herrerasauria |journal=The Anatomical Record |language=en |volume=307 |issue=4 |pages=1011–1024 |doi=10.1002/ar.25342 |pmid=37971103 |issn=1932-8486}} matrix yielded similar results, with Maleriraptor as a non-herrerasaurid herresaur. The results using the Ezcurra et al. (2023) dataset are displayed in the cladogram below:
{{clade
|{{clade
|label1=Eusaurischia
|1={{clade
|1=Theropoda
|2=Sauropodomorpha }}
|label3=Herrerasauria
|3={{clade
|1=CAPPA/UFSM 0373 {{small|(Santa Maria Fm. taxon)}}
|2={{clade
|1=Maleriraptor
|2={{clade
|1=TTU-P10082 {{small|(Cooper Canyon Fm. taxon{{Cite journal |last1=Sarıgül |first1=Volkan |date=2017-02-01 |title=New theropod fossils from the Upper Triassic Dockum Group of Texas, USA, and a brief overview of the Dockum theropod diversity |journal=PaleoBios |volume=34 |doi=10.5070/P9341033817 |issn=2373-8189}})}}
|2={{clade
|1={{clade
|2=Tawa
|3=Chindesaurus }}
|label4=Herrerasauridae
|4={{clade
|2={{clade
|2={{clade
|1=UFSM 11330 {{small|(Santa Maria Fm. taxon{{Cite journal |last1=Garcia |first1=Maurício S. |last2=Müller |first2=Rodrigo T. |last3=Pretto |first3=Flávio A. |last4=Da-Rosa |first4=Átila A. S. |last5=Dias-Da-Silva |first5=Sérgio |date=2021-03-09 |title=Taxonomic and phylogenetic reassessment of a large-bodied dinosaur from the earliest dinosaur-bearing beds (Carnian, Upper Triassic) from southern Brazil |journal=Journal of Systematic Palaeontology |volume=19 |issue=1 |pages=1–37 |doi=10.1080/14772019.2021.1873433 |bibcode=2021JSPal..19....1G |issn=1477-2019}})}}
|2=Gnathovorax
}} }} }} }} }} }} }} }}|label1=Saurischia}}
References
{{Reflist}}
{{Avemetatarsalia}}
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