Orthalicoidea
{{Short description|Superfamily of gastropods}}
{{Automatic taxobox
|taxon = Orthalicoidea
|image = OrthalicidSnailPeru.jpg
|image_caption = A large orthalicid from eastern Peru (probably of genus Sultana), recovering from substantial shell damage.
|authority = Albers, 1860
|synonyms_ref =
|synonyms =
|subdivision_ranks = Families
|subdivision = See text
|diversity = ~1750 taxon names are available
|display_parents = 3
}}
Orthalicoidea is a superfamily of air-breathing land snails, terrestrial gastropod mollusks in the infraorder Orthalicoidei of the suborder Helicina.MolluscaBase eds. (2021). MolluscaBase. Simpulopsidae Schileyko, 1999. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=870047 on 2021-02-05
Distribution
The Orthalicoidea is a dominant faunal element in the Neotropics, but also has a number of genera with a Gondwanan distribution.
Taxonomy
= 2005 taxonomy =
This taxonomy, as accepted by Bouchet & Rocroi, was based on the study by Nordsieck, published in 1986,{{cite journal | author=Nordsieck H.| title=The system of the Stylommatophora (Gastropoda), with special regard to the systematic position of the Clausiliidae, II. Importance of the shell and distribution| journal=Archiv für Molluskenkunde| year=1986| volume=117| issue=1–3| pages=93–116}} and the classification as formulated by Schileyko in 1999.{{cite journal | author=Schileyko A. A.| title=Treatise on recent terrestrial pulmonate molluscs| journal=Ruthenica| year=1998–2003| volume=suppl. 2| pages=1–1626}} However Bouchet & Rocroi diverged from this classification in uniting the families Bulimulidae and Orthalacidae. They also considered the family Placostylidae as a distinct family from Orthalicidae, and the family Coelociontidae a distinct family from Urocoptidae. However, the position of the families Megaspiridae and Grangerellidae is doubtful.{{Bouchet 2005}}
The following seven families have been recognized in the taxonomy of Bouchet & Rocroi (2005):
- Family Cerionidae Pilsbry, 1901
- Family Coelociontidae Iredale, 1937
- † Family Grangerellidae Russell, 1931
- Family Megaspiridae Pilsbry, 1904
- Family Orthalicidae Albers, 1860
- Family Placostylidae Pilsbry, 1946
- Family Urocoptidae Pilsbry, 1898 (1868)
- Vidaliellidae H. Nordsieck, 1986 †
= 2008 taxonomy =
Uit de Weerd moved two families Urocoptidae and Cerionidae to newly established superfamily Urocoptoidea based on molecular phylogeny research in 2008.Uit de Weerd D. R. (2008). "Delimitation and phylogenetics of the highly diverse land snail family Urocoptidae (Gastropoda, Pulmonata) based on 28S rRNA sequence data: A reunion with Cerion". Journal of Molluscan Studies 74: 317-329. {{doi|10.1093/mollus/eyn023}}.
= 2009 taxonomy =
Family Prestonellidae van Bruggen, 1978 - A study by Herbert and Mitchell, published in 2009, places this enigmatic family, of which the phylogenetic relationships were previously unknown, in the Gondwanan superfamily Orthalicoidea,{{cite journal | doi=10.1111/j.1095-8312.2008.01109.x |author1=Herbert D. G. |author2=Mitchell A. |name-list-style=amp | title=Phylogenetic relationships of the enigmatic land snail genus Prestonella: the missing African element in the Gondwanan superfamily Orthalicoidea (Mollusca: Stylommatophora)| journal=Zoological Journal of the Linnean Society| year=2009| volume=96| issue=1| pages=203–221| doi-access=free}} while it was previously tentatively placed as a synonym of Aillyidae.
= 2010 taxonomy =
Breure et al. (2010) moved Prestonella and Bothriembryon to Placostylidae and they elevated Bulimulinae to Bulimulidae, Odontostomini to Odontostomidae, Amphibuliminae to Amphibulimidae. They also removed Coelociontidae from Orthalicoidea.Breure A. S. H., Groenenberg D. S. J. & Schilthuizen M. (2010). "New insights in the phylogenetic relations within the Orthalicoidea (Gastropoda, Stylommatophora) based on 28S sequence data". Basteria 74(1-3): 25-31.
= 2012 taxonomy =
Breure & Romero (2012) confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae, renamed Placostylidae to Bothriembryontidae.Breure A. S. H. & Romero P. (2012). "Support and surprises: molecular phylogeny of the land snail superfamily Orthalicoidea using a three-locus gene analysis with a divergence time analysis and ancestral area reconstruction (Gastropoda: Stylommatophora)". Archiv für Molluskenkunde: International Journal of Malacology 141(1): 1-20. {{doi|10.1127/arch.moll/1869-0963/141/001-020}}. Therefore, there are recognized seven extant families and one extinct family within Orthalicoidea:
- Amphibulimidae P. Fischer, 1873
- Bothriembryontidae Iredale, 1937
- Bulimulidae Tryon, 1867
- † Grangerellidae Russell, 1931
- Megaspiridae Pilsbry, 1904
- Odontostomidae Pilsbry & Vanatta, 1898
- Orthalicidae Albers, 1860
- Simpulopsidae Schileyko, 1999
- † Vidaliellidae H. Nordsieck, 1986
References
This article incorporates CC-BY-3.0 text from the referenceBreure A. S. H. (2011). "Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with description of two new species". ZooKeys 101: 1-50. {{doi|10.3897/zookeys.101.1133|doi-access=free}}.
{{Reflist}}
Further reading
- Borrero F. J. & Breure A. S. H. ([http://www.ashbreure.nl/page1/page1.html Submitted]). "The Orthalicoidea (Mollusca: Gastropoda: Stylommatophora) from Colombia and adjacent areas: a review of taxonomy and biogeography. 1. Dryptus Albers, 1860 and Plekocheilus Guilding, 1828".
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