Palaeophis
{{Short description|Extinct genus of snakes}}
{{Automatic taxobox
| fossil_range = Ypresian - Priabonian, {{Fossil range|56.0|33.9|earliest=100.5}} Possible Cenomanian - Maastrichtian record
| image = Palaeophiidae - Palaeophis maghrebianus.JPG
| image_caption = Fossil vertebrae of Palaeophis maghrebianus from Khouribga (Morocco)
| image_upright = 1.15
| taxon = Palaeophis
| authority = Owen, 1841
| type_species = {{extinct}}Palaeophis toliapicus
| type_species_authority = Owen, 1841
| subdivision_ranks = Species
| subdivision = {{collapsible list|bullets = true
|title=Species
| †Palaeophis africanus Andrews 1924
| †Palaeophis casei Holman 1982
| †Palaeophis colossaeus Rage 1983
| †Palaeophis ferganicus Averianov 1997
| †Palaeophis grandis Marsh 1869
| †Palaeophis littoralis Cope 1847
| †Palaeophis maghrebianus Arambourg 1952
| †Palaeophis nessovi Averianov 1997
| †Palaeophis oweni Zigno 1881
| †Palaeophis tamdy Averianov 1997
| †Palaeophis toliapicus Owen 1841(type)
| †Palaeophis typhaeus Owen 1850
| †Palaeophis vastaniensis Bajpai & Head 2008
| †Palaeophis virginianus Lynn 1934
| †Palaeophis zhylan Averianov 1997}}
| synonyms = *Dinophis (Marsh, 1869)
- Titanophis (Marsh, 1878)
- Vialovophis (Averianov, 1997)
}}
Palaeophis ('ancient snake') is an extinct genus of marine snake that is the type genus of the extinct snake family Palaeophiidae.
Described species within this genus lived in the Eocene epoch, with some unnamed or questionable records from Cenomanian and Maastrichtian.{{Cite journal |last1=Folie |first1=Annelise |last2=Mees |first2=Florias |last3=De Putter |first3=Thierry |last4=Smith |first4=Thierry |date=2021-07-01 |title=Presence of the large aquatic snake Palaeophis africanus in the middle Eocene marine margin of the Congo Basin, Cabinda, Angola |url=https://www.sciencedirect.com/science/article/pii/S0016699520300954 |journal=Geobios |series=Proceedings of the PalEurAfrica project international symposium Evolution and Paleoenvironment of Early Modern Vertebrates during the Paleogene |language=en |volume=66-67 |pages=45–54 |doi=10.1016/j.geobios.2020.11.002 |bibcode=2021Geobi..66...45F |s2cid=229411514 |issn=0016-6995}} Fossils of species within this genus have been found in England, France, Denmark,{{cite journal|last=Kristensen|first=H. V.|author2=Cuny, G. |author3=Rasmussen, A. R. |author4= Madsen, H |title=Earliest record of the fossil snake Palaeophis from the Paleocene/Eocene boundary in Denmark|journal=Bulletin de la Société Géologique de France|date=2012|volume=183|issue=6|pages=621–625|url=http://bsgf.geoscienceworld.org/content/183/6/621.abstract |doi=10.2113/gssgfbull.183.6.621}} Morocco and Mali.{{cite journal|last=Rage|first=J.-C.|title=Palaeophis colossaeus nov. sp. (le plus grand Seprent connu?) de l'Eocène du Mali et le problème du genre chez les Palaeopheinae|journal=Comptes rendus de l'Académie des Sciences de Paris |date=1983|volume=3|issue=296|pages=1741–1744}} Remains have also been found in North America, including Maryland and Virginia (from the early Eocene Nanjemoy Formation),Blake, S. F. "Note on a vertebra of Palaeophis from the Eocene of Maryland." Journal of the Washington Academy of Sciences 31.12 (1941): 501-503. GeorgiaParmley, Dennis, and Harold W. Reed. "Size and age class estimates of North American Eocene palaeopheid snakes." Georgia Journal of Science 61.4 (2003): 220. and Mississippi.Holman, J. Alan. "Palaeophis casei, new species, a tiny palaeophid snake from the early Eocene of Mississippi." Journal of Vertebrate Paleontology 2.2 (1982): 163-166.
Description
File:Palaeophis maghrebianus life restoration.jpg
File:Palaeophis toliapicus.jpg
These species varied broadly in size; Palaeophis casei is the smallest at 1.3 metres of length, while the largest species, Palaeophis colossaeus, is estimated to have been {{cvt|8.1|-|12.3|m|ft}} long based on isolated vertebrae,{{Cite journal |last1=McCartney |first1=Jacob |last2=Roberts |first2=Eric |last3=Tapanila |first3=Leif |last4=O'Leary |first4=Maureen |date=2018 |title=Large palaeophiid and nigerophiid snakes from Paleogene Trans-Saharan Seaway deposits of Mali |journal=Acta Palaeontologica Polonica |language=en |volume=63 |doi=10.4202/app.00442.2017 |s2cid=59147071 |issn=0567-7920|doi-access=free }} making it one of the largest known snakes. However, most species of the genus were not as big.{{cite journal |author=Holman, J. Alan |year=1982 |title=Palaeophis casei, new species, a tiny palaeophid snake from the early Eocene of Mississippi |journal=Journal of Vertebrate Paleontology |volume=2 |issue=2 |pages=163–166 |jstor=4522892 |doi=10.1080/02724634.1982.10011927|bibcode=1982JVPal...2..163H }}{{cite journal| last = Rage| first = Jean-Claude|title = Early Eocene snakes from Kutch, Western India, with a review of the Palaeophiidae| journal = Geodiversitas| volume = 25| issue = 4| pages = 695–716|publisher = Editions scientifiques du Muséum, Paris, France| year = 2003| url = https://www.researchgate.net/publication/287476886| issn = 1280-9659| access-date = 12 May 2010|display-authors=etal}} There are many species of Palaeophis but they can be separated into two assemblages of species or grades. In which the primitive grade include species whose vertebrae are weakly laterally compressed and have less developed and low process of vertebrae. Subsequently the advanced grade are characterized by vertebrae presenting a strong lateral compression which translate to being much better adaption to aquatic life.
Biology
Species of Palaeophis were specialised aquatic animals, as their fossils occur primarily in marine strata, though at least some estuarine remains have also been found. Different species are thought to have occupied different ecological niches.
Studies on Palaeophis vertebrae show a high degree of vascularisation, suggesting that it had a considerably faster metabolism and growth rate than modern snakes. This may suggest that palaeophiids, like other marine reptiles such as mosasaurs, might have developed towards endothermy.{{cite journal|last1=Houssaye|first1=Alexandra|last2=Rage|first2=Jean-Claude |last3=Bardet|first3=Nathalie |last4=Vincent|first4=Peggy|last5=Amaghzaz|first5=Mbarek |last6=Meslouh|first6=Said |year=2013 |title=New highlights about the enigmatic marine snake Palaeophis maghrebianus (Palaeophiidae; Palaeophiinae) from the Ypresian (Lower Eocene) phosphates of Morocco |journal=Palaeontology |volume=56 |issue=3 |pages=647–661 | doi=10.1111/pala.12008|doi-access= |bibcode=2013Palgy..56..647H |s2cid=129055598 }}
References
{{Commons category|Palaeophis}}
{{Portal|Paleontology}}
{{Reflist|2}}
{{Fossil snakes}}
{{Taxonbar|from=Q2449270}}
Category:Paleocene reptiles of Europe
Category:Paleogene reptiles of Africa
Category:Eocene reptiles of Europe
Category:Taxa named by Richard Owen
Category:Fossil taxa described in 1841
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