Qiupa Formation

{{Short description|Geological formation in Henan, China}}

{{Infobox rockunit

| name = Qiupa Formation

| image = Stratigraphic occurrence of Yubaatar - sediments.png

| caption = Characteristic red brownish siltstones of the Qiupa Formation

| type = Geological formation

| age = Maastrichtian
~{{fossilrange|72|66}}

| period = Maastrichtian

| prilithology = Mudstone, siltstone

| otherlithology = Sandstone, conglomerate

| namedfor = Qiupa

| namedby =

| region = Henan Province

| country = China

| coordinates = {{coord|33.9|N|111.8|E|display=inline,title}}

| paleocoordinates = {{coord|31.6|N|102.8|E|display=inline}}

| unitof = Tantou Group

| subunits =

| underlies = Gaoyugou Formation

| overlies = Shitaijie Formation

| thickness =

| extent = Tantou Basin

| area =

| map = {{Location map+ | China#China Henan

| relief = 1

| width = 250

| float = center

| places =

{{Location map~ | China#China Henan

| lat_deg = 33.9

| lon_deg = 111.8

| mark = Lightgreen pog.svg

| marksize = 12

}}

}}

| map_caption =

}}

The Qiupa Formation ({{zh|s=秋扒组|p=Qiūpázǔ}}) is a Late Cretaceous Maastrichtian geologic formation in Henan Province, central China. It is rich in dinosaur eggs and bones, such as those of carnivorous and herbivorous dinosaurs. The Qiupa Formation is considered to be Late Maastrichtian in age, about 72 million and 66 million years ago.{{cite book|author=Bureau of Geology and Mineral Resources of Henan Province|date=1989|title=Regional Geology of Henan Province|publisher=Geological Publishing House|location=Beijing|pages=1−772}}{{cite journal |last1=Lü |first1=J. C. |last2=Xu |first2=L. |last3=Chang |first3=H. L. |last4=Jia |first4=S. H. |last5=Zhang |first5=J. M. |last6=Gao |first6=D. S. |last7=Zhang |first7=Y. Y. |last8=Zhang |first8=C. J. |last9=Ding |first9=F. |date=2018 |title=A new alvarezsaurid dinosaur from the Late Cretaceous Qiupa Formation of Luanchuan, Henan Province, central China |journal=China Geology |volume=1 |issue=1 |pages=28–35 |bibcode=2018CGeo....1...28L |doi=10.31035/cg2018005 |doi-access=free}}

Geology

The sedimentation is characterized for the presence of calcareous mudstone interbedded with thin fine conglomerates, brownish red thick-bedded siltstone and parallel and cross laminations. This sedimentation indicates habitats composed by large and shallow meanders with braided river deltas.{{cite journal|last1=Jiang|first1=X.-J.|last2=Liu|first2=Y.-Q.|last3=Ji|first3=S.-A.|last4=Zhang|first4=X.-L.|last5=Xu|first5=L.|last6=Jia|first6=S.-H.|last7=Lü|first7=J.-C.|last8=Yuan|first8=C.-X.|last9=Li|first9=M.|date=2011|title=Dinosaur-bearing strata and K/T boundary in the Luanchuan-Tantou Basin of western Henan Province, China|journal=Science China Earth Sciences|volume=54|issue=1149|pages=1149–1155 |doi=10.1007/s11430-011-4186-1|bibcode=2011ScChD..54.1149J }}

Fossil content

The Qiupa Formation has yielded numerous dinosaur fossils, particularly eggs. The remains of various theropods such as troodontids, dromaeosaurids and oviraptorosaurs have been found. In addition, indeterminate remains of sauropods, ornithopods, ankylosaurs, lizards, turtles and a possible avian bone are reported.{{cite journal|last1=Kobayashi|first1=Y.|last2=Lü|first2=J.|last3=Lee|first3=Y.-N.|last4=Xu|first4=L.|last5=Zhang|first5=X.|date=2008|title=A new basal ornithomimid (Dinosauria: Theropoda) from the Late Cretaceous in Henan Province of China|journal=Journal of Vertebrate Paleontology|volume=28|issue=supp. 3|pages=101A|doi=10.1080/02724634.2008.10010459|url=http://digital.csic.es/bitstream/10261/12900/1/Gongbusaurus.pdf}}

= Dinosaurs =

class="wikitable" align="center"
colspan="5" align="center" | Dinosaurs of the Qiupa Formation
GenusSpeciesMaterialNotesImages
Ankylosauridae indet.{{cite journal |last1=Jia |first1=S.-H. |last2=Lü |first2=J.-C. |last3=Xu |first3=L. |last4=Hu |first4=W.-Y. |last5=Li |first5=J.-H. |last6=Zhang |first6=J.-M. |date=2010 |title=Discovery and significance of ankylosaur specimens from the Late Cretaceous Qiupa Formation in Luanchuan, Henan, China |url=http://dzhtb.cgs.cn/article/id/20100401 |journal=Geological Bulletin of China |volume=29 |issue=4 |pages=483–487}}

| Indeterminate

| "A tooth, well-preserved dorsal vertebra, incomplete dorsal rib and ischium."

| An ankylosaurid

|

style="background:#e3f5ff;" | Elongatoolithidae sp.{{cite journal |last1=Tanaka |first1=K. |last2=Lü |first2=J.-C. |last3=Kobayashi |first3=Y. |last4=Zelenitsky |first4=D. K. |last5=Xu |first5=L. |last6=Jia |first6=S. |last7=Qin |first7=S. |last8=Tang |first8=M. |date=2011 |title=Description and Phylogenetic Position of Dinosaur Eggshells from the Luanchuan Area of Western Henan Province, China |journal=Acta Geologica Sinica (English Edition) |volume=85 |issue=1 |pages=66–74 |bibcode=2011AcGlS..85...66K |doi=10.1111/j.1755-6724.2011.00379.x}}

|style="background:#e3f5ff;" | Indeterminate

|style="background:#e3f5ff;" | "145 egg shells probably laid by oviraptorids."

|style="background:#e3f5ff;" | Oviraptorid eggs

|style="background:#e3f5ff;" |

Luanchuanraptor{{cite journal |last1=Lü |first1=J.-C. |last2=Xu |first2=L. |last3=Zhang |first3=X.-L. |last4=Ji |first4=Q. |last5=Jia |first5=S.-H. |last6=Hu |first6=W.-Y. |last7=Zhang |first7=J.-M. |last8=Wu |first8=Y.-H. |date=2007 |title=New dromaeosaurid dinosaur from the Late Cretaceous Qiupa Formation of Luanchuan area, western Henan, China |url=http://dzhtb.cgs.cn/en/article/id/200707141 |journal=Geological Bulletin of China |volume=26 |issue=7 |pages=777–786 |s2cid=130054772}}

| L. henanensis

| "Teeth, a frontal and fragmented postcrania."

| A dromaeosaurid

| File:Luanchuanraptor.jpg

Qiupalong{{cite journal |last1=Xu |first1=L. |last2=Kobayashi |first2=Y. |last3=Lü |first3=J.-C. |last4=Lee |first4=Y. N. |last5=Liu |first5=Y. |last6=Tanaka |first6=K. |last7=Zhang |first7=X. |last8=Jia |first8=S. |last9=Zhang |first9=J. |date=2011 |title=A new ornithomimid dinosaur with North American affinities from the Late Cretaceous Qiupa Formation in Henan Province of China |journal=Cretaceous Research |volume=32 |issue=2 |pages=213–222 |bibcode=2011CrRes..32..213X |doi=10.1016/j.cretres.2010.12.004}}

| Q. henanensis

| Partial skeleton preserving the pelvis and the right hindlimb.

| An ornithomimid

| File:Qiupalong Restoration.png

Qiupanykus

| Q. zhangi

| Partial skeleton with vertebrae and hindlimbs.

| An alvarezsaurid

|

"Tyrannosaurus"{{Cite book |last=Dong |first=Zhiming |url=https://books.google.com/books?id=LsYDMgAACAAJ |title=Mesozoic and Cenozoic red beds of South China |publisher=Science Press |year=1979 |location=Beijing |pages=342–350 |language=zh |chapter=Dinosaurs from the Cretaceous of South China |doi=10.5281/zenodo.162178 |chapter-url=https://zenodo.org/records/162178}}

| T. luanchuanensis

| Five teeth.

| A tyrannosaurid

|

Yulong{{cite journal |last1=Lü |first1=J.-C. |last2=Currie |first2=P. J. |last3=Xu |first3=L. |last4=Zhang |first4=X. |last5=Pu |first5=H. |last6=Jia |first6=S. |date=2013 |title=Chicken-sized oviraptorid dinosaurs from central China and their ontogenetic implications |journal=Naturwissenschaften |volume=100 |issue=2 |pages=165–175 |bibcode=2013NW....100..165L |doi=10.1007/s00114-012-1007-0 |pmid=23314810}}{{cite journal |last1=Wei |first1=X. |last2=Kundrát |first2=M. |last3=Xu |first3=L. |last4=Ma |first4=W. |last5=Wu |first5=Y. |last6=Chang |first6=H. |last7=Zhang |first7=J. |last8=Zhou |first8=X. |date=2022 |title=A new subadult specimen of oviraptorid Yulong mini (Theropoda: Oviraptorosauria) from the Upper Cretaceous Qiupa Formation of Luanchuan, central China |journal=Cretaceous Research |volume=138 |bibcode=2022CrRes.13805261W |doi=10.1016/j.cretres.2022.105261 |s2cid=248977151 |number=105261}}

| Y. mini

| Many juvenile specimens and one subadult skeleton.

| An oviraptorid

| File:Yulong.png

Yuornis{{Cite journal |last1=Xu |first1=Li |last2=Buffetaut |first2=Eric |last3=O’Connor |first3=Jingmai |last4=Zhang |first4=Xingliao |last5=Jia |first5=Songhai |last6=Zhang |first6=Jiming |last7=Chang |first7=Huali |last8=Tong |first8=Haiyan |date=2021 |title=A new, remarkably preserved, enantiornithine bird from the Upper Cretaceous Qiupa Formation of Henan (central China) and convergent evolution between enantiornithines and modern birds |url=https://www.researchgate.net/publication/354585552 |journal=Geological Magazine |language=en |volume=158 |issue=11 |pages=2087–2094 |doi=10.1017/S0016756821000807 |bibcode=2021GeoM..158.2087X |issn=0016-7568}}

|Y. junchangi

|Partial skeleton.

|An enantiornithine bird

|

= Mammals =

class="wikitable" align="center"
colspan="5" align="center" | Mammals of the Qiupa Formation
GenusSpeciesMaterialNotesImages
Lotheridium{{cite journal |last1=Bi |first1=S. |last2=Jin |first2=X. |last3=Li |first3=S. |last4=Du |first4=T. |date=2015 |title=A new Cretaceous Metatherian mammal from Henan, China |journal=PeerJ |volume=3 |pages=e896 |doi=10.7717/peerj.896 |pmc=4400878 |pmid=25893149 |doi-access=free}}

| L. mengi

| Nearly complete skull with jaws.

| A deltatheridiid

| File:Lotheridium.png

Yubaatar{{cite journal |last1=Xu |first1=L. |last2=Zhang |first2=X. |last3=Pu |first3=H. |last4=Jia |first4=S. |last5=Zhang |first5=J. |last6=Lü |first6=J.-C. |last7=Meng |first7=J. |date=2015 |title=Largest known Mesozoic multituberculate from Eurasia and implications for multituberculate evolution and biology |journal=Scientific Reports |volume=5 |issue=1495 |page=14950 |bibcode=2015NatSR...514950X |doi=10.1038/srep14950 |pmc=4615031 |pmid=26492455 |doi-access=free}}

| Y. zhongyuanensis

| "Virtually complete crania and fragmented poscrania."

| A cimolodont

| 200px

= Squamates =

class="wikitable" align="center"
colspan="5" align="center" | Squamates of the Qiupa Formation
GenusSpeciesMaterialNotesImages
Funiusaurus{{cite journal |last1=Xu |first1=L. |last2=Lü |first2=J.-C. |last3=Jia |first3=S. |last4=Zhang |first4=J. |last5=Pu |first5=H. |last6=Zhang |first6=X. |date=2014 |title=A New Lizard (Lepidosauria: Squamata) from the Upper Cretaceous of Henan, China |url=https://www.researchgate.net/publication/265912600 |journal=Acta Geologica Sinica |volume=88 |issue=4 |pages=1041–1050 |bibcode=2014AcGlS..88.1041X |doi=10.1111/1755-6724.12271}}

| F. luanchuanensis

| Partial skull.

| A polyglyphanodont

|

Tianyusaurus{{cite journal |last1=Lü |first1=J.-C. |last2=Ji |first2=S. |last3=Dong |first3=Z. |last4=Wu |first4=X.-C. |date=2008 |title=An Upper Cretaceous lizard with a lower temporal arcade |url=https://www.researchgate.net/publication/5514468 |journal=Naturwissenschaften |volume=95 |issue=7 |pages=663–669 |bibcode=2008NW.....95..663L |doi=10.1007/s00114-008-0364-1 |pmid=18338150}}

| T. zhengi

| Skull and shoulder girdle.

| A polyglyphanodont

| File:Tianyusaurus.jpg

Zhongyuanxi{{cite journal |last1=Xu |first1=L. |last2=Dong |first2=L. |last3=Gao |first3=D. |last4=Li |first4=Y. |last5=Wu |first5=Y. |last6=Chang |first6=F. |last7=Li |first7=J. |last8=Evans |first8=S. |year=2025 |title=A new anguimorph lizard from the Upper Cretaceous of Henan Province, China |journal=Journal of Systematic Palaeontology |volume=23 |issue=1 |at=2470790 |doi=10.1080/14772019.2025.2470790|bibcode=2025JSPal..2370790X }}

| Z. jiai

| Partial skull.

| A stem-varanid anguimorph

|

{{paleobiota-key-compact}}

References