Sacabambaspis

{{Automatic taxobox

| fossil_range = {{Fossil range|461|450|Dapingian-Sandbian}}

| image = Sacabambaspis janvieri cast (cropped).jpg

| image_caption = Sacabambaspis janvieri fossil cast of a specimen from Bolivia.

| taxon = Sacabambaspis

| authority = Gagnier, Blieck & Rodrigo, 1986

| type_species = †Sacabambaspis janvieri

| type_species_authority = Gagnier, Blieck & Rodrigo, 1986

| subdivision_ranks = Species

| subdivision = *S. janvieri Gagnier, Blieck & Rodrico, 1986

}}

Sacabambaspis is an extinct genus of jawless fish that lived in the Ordovician period. Sacabambaspis lived in shallow waters on the continental margins of Gondwana.{{Cite journal |last1=Sansom |first1=Ivan J. |last2=Miller |first2=C. Giles |last3=Heward |first3=Alan |last4=Davies |first4=Neil S. |last5=Booth |first5=Graham A. |last6=Fortey |first6=Richard A. |last7=Paris |first7=Florentin |title=Ordovician Fish from the Arabian Peninsula |date=2009 |journal=Palaeontology |language=en |volume=52 |issue=2 |pages=337–342 |doi=10.1111/j.1475-4983.2009.00846.x|s2cid=128704925 |doi-access=free |bibcode=2009Palgy..52..337S }} It is the best known arandaspid with many specimens known.

Description

{{multiple image

| align = left

| direction = vertical

| width = 250

| image1 = Sacabambaspis.jpg

| footer = Life restoration

}}

Sacabambaspis was approximately {{convert|25|cm|in|abbr=in|frac=4}} in length. The body shape of Sacabambaspis vaguely resembled that of a tadpole with an oversized head, flat body, wriggling tail, and lack of fins. It had characteristic, frontally positioned eyes, like car headlamps.Janvier, Philippe (1997) [http://tolweb.org/Arandaspida/16907 Arandaspida] The Tree of Life Web Project.

=Armor=

Sacabambaspis had a head shield made from a large upper (dorsal) plate that rose to a slight ridge in the midline and a deep curved lower (ventral) plate. This headshield is ornamented with characteristic oak-leaf shaped or tear-drop shaped tubercles.{{cite book|last=Benton|first=Michael J.|title=Vertebrate Palaeontology 3rd Edition|year=2005|publisher=Blackwell Publishing|isbn=978-1-4051-4449-0|page=47|url=https://books.google.com/books?id=VThUUUtM8A4C&q=Sacabambaspis+sensory+structure&pg=PA47}}{{cite book|last=Taylor & Francis|title=PE Ahlberg (ed.) Major Events in Early Vertebrate Evolution|year=2001|publisher=Sansom IJ, MM Smith & MP Smith|pages=156–171}} It also had narrow branchial plates which link these two along the sides, and cover the gill area. The eyes were far forward and between them are possibly two small nostrils and they, which are surrounded by what is thought to be endoskeletal bone and putative nostrils, are found at the extreme anterior of the head, one of the diagnostic features of the arandaspids. The rest of the body was covered by long, strap-like scales behind the head.

=Tail=

The tail consists of relatively large dorsal and ventral webs and an elongated notochordal lobe, the posterior end of which is bordered by a small fin web. This tail structure clearly differs from that of heterostracans, which are currently grouped with arandaspids and astraspids in the clade Pteraspidomorphi (Gagnier 1993, 1995; Donoghue & Smith 2001; Sansom et al. 2005), in which the caudal fin looks diphycercal (i.e. symmetrical) and is strengthened by a few large radials (Janvier 1996).{{cite journal|title=The tail of the Ordovician fish Sacabambaspis|pmc=2373808|journal=Biology Letters|year=2006|publisher=Alan Pradel, Ivan. J Sansom, Pierre-Yves Gagnier, Ricardo Cespedes, and Philippe Janvier|last1=Pradel|first1=A.|last2=Sansom|first2=I. J.|last3=Gagnier|first3=P. Y.|last4=Cespedes|first4=R.|last5=Janvier|first5=P.|volume=3|issue=1|pages=72–75|doi=10.1098/rsbl.2006.0557|pmid=17443969}}

Discovery and species

File:Sacabambaspis janvieri many specimens.JPG

Sacabambaspis is named after the village of Sacabamba, Cochabamba Department, Bolivia, where the first fossils of the genus were found.{{Cite journal |last1=Gagnier |first1=Pierre-Yves |last2=Blieck |first2=Alain R. M. |last3=S. |first3=Gabriela Rodrigo |date=1986 |title=First Ordovician vertebrate from South America |url=https://www.sciencedirect.com/science/article/pii/S0016699586800584 |journal=Geobios |language=en |volume=19 |issue=5 |pages=629–634 |doi=10.1016/S0016-6995(86)80058-4 |bibcode=1986Geobi..19..629G |issn=0016-6995}} S. janvieri (Gagnier, Blieck & Rodrico, 1986), the type species of the genus, is known from the Anzaldo Formation of Bolivia. There are 30 known specimens of this Bolivian species, all crammed into a very confined area, believed to be the result of a fish kill, probably due a sudden inflow of freshwater from a large storm. They were found associated with a large number of lingulid brachiopods, also killed at the same time.{{cite web|title=Australia: The Land Where Time Began|url=http://austhrutime.com/arandapsis.htm|accessdate=28 October 2013}}

Indeterminate specimens (described as "Sacabambaspis sp.") have been found in many countries corresponding to the margin of Gondwana. Young, 1997 described fossils of the genus from the Stokes Siltstone and Carmichael Sandstone of Central Australia.{{Cite journal |last=Young |first=G. C. |date=1997-04-16 |title=Ordovician microvertebrate remains from the Amadeus Basin, central Australia |url=https://www.researchgate.net/publication/254313959 |journal=Journal of Vertebrate Paleontology |volume=17 |issue=1 |pages=1–25 |doi=10.1080/02724634.1997.10010948 |bibcode=1997JVPal..17....1Y |issn=0272-4634}} Isolated scales found in the Horn Creek Siltstone from Central Australia have a very similar ornamentation to the Bolivian scales. Specimens have also been reported from Argentina. Sansom et al., 2009 described specimens from the Amdeh Formation of Oman on the Arabian Peninsula. The Oman discoveries showed that the fish were present all around the periphery of the ancient continent of Gondwana and not just in the southern regions as had previously been shown by the findings from South America and Australia.{{cite web |last=Miller |first=George |date=24 June 2011 |title=How can something that small appear on TV? |url=http://www.nhm.ac.uk/natureplus/blogs/micropalaeo/tags/sacabambaspis?fromGateway=true |archive-url=https://web.archive.org/web/20181007005435/http://www.nhm.ac.uk/natureplus/blogs/micropalaeo/tags/sacabambaspis?fromGateway=true |archive-date=7 October 2018 |website=NaturePlus |publisher= |accessdate=28 October 2013}}

Paleobiology

=Feeding=

Although it had no jaws, the mouth of Sacabambaspis janvieri was lined with nearly 60 rows of small bony oral plates which were probably movable in order to provide more efficient suction feeding through expansion and contraction of the oral cavity and pharynx.

=Sensory system=

The fossils of Sacabambaspis show clear evidence of a sensory structure (lateral line system). This is a line of pores within each of which are open nerve endings that can detect slight movements in the water, produced for example by predators. The arrangement of these organs in regular lines allows the fish to detect the direction and distance from which a disturbance in the water is coming.

In popular culture

In 2023, Sacabambaspis became a source of memes and fanarts after a viral tweet from August 2022 about a model of the Sacabambaspis from the Natural History Museum of Helsinki in Finland.{{cite web |url=https://automaton-media.com/en/news/20230616-19550/ |title=470-million-year-old fish Sacabambaspis goes viral in Japan, Among Us dev joins in on fun |author=Amber V |publisher=Automaton Media |access-date=8 February 2024 |date=16 June 2023 }} A song inspired by the meme, entitled The Sacabambaspis Song ({{lang|ja|サカサカバンバンバスピスピス}}) was released by Japanese music producer Yakamochi on 13 June 2023, using the Japanese-language Neutrino singing voicebanks of Kotonoha Akane and Aoi (as of March 2025). It had gained over 4.1 million views on YouTube.{{cite AV media|via=YouTube|date=13 June 2023|title=サカサカバンバンバスピスピス / 琴葉姉妹（neutrino / AI_VOICE）|url=https://www.youtube.com/watch?v=aPkwGhwU_FY|access-date=25 March 2024}}