Scenella
{{Short description|Genus of molluscs (fossil)}}
{{Taxobox
| fossil_range = {{fossil range|Cambrian Stage 3|Late Ordovician|ref=}}
| image = Scenella tenuistriata.png
| image_caption = Dorsal view of the shell of Scenella tenuistriata
| image2 = Scenella tenuistriata 2.png
| image2_caption = Lateral view of the shell of Scenella tenuistriata
| regnum = Animalia
| phylum = Mollusca
| ordo =
| superfamilia = † Scenelloidea
| familia = † Scenellidae
| familia_authority = S. A. Miller, 1889Miller S. A. (after October 1889). North American geology and palaeontology: 389.
| genus= Scenella
| genus_authority = Billings, 1872
| subdivision_ranks = Species
| synonyms =
- Helcionellinae Wenz, 1938
- Hampilinae Kobayashi, 1958
- Securiconidae Missarzhevsky, 1989
}}
Scenella is an extinct genus of fossil invertebrate animal which is generally considered to be a mollusc; at various times it has been suggested that this genus belongs with the gastropods, the monoplacophorans, or the helcionellids, although no firm association with any of these classes has been established.{{cite journal | year = 1984 | last1 = Yochelson | first1 = E. L. | last2 = Cid | first2 = D. G. L. | title = Reevaluation of the systematic position of Scenella | journal = Lethaia|volume =17 |issue =4 | pages = 331–340 | url = http://www3.interscience.wiley.com/journal/120036307/abstract | doi = 10.1111/j.1502-3931.1984.tb00679.x| doi-broken-date = 3 April 2025 | url-access = subscription }}{{dead link|date=February 2019|bot=medic}}{{cbignore|bot=medic}} An affinity with the hydrozoa (as a flotation device) has been considered, although some authors oppose this hypothesis.{{Cite journal| doi = 10.1017/s0022336000033837| journal = Journal of Paleontology| year = 1992| volume = 66| issue = 2| title = Scenella and "A Chondrophorine (Medusoid Hydrozoan) from the Basal Cambrian (Placentian) of Newfoundland"| author = Landing| author2 = Narbonne| jstor = 1305918| pages = 338| bibcode = 1992JPal...66..338L| s2cid = 132296800}} A gastropod affinity is defended on the basis of six pairs of internal muscle scars,{{Cite journal| journal = Journal of Paleontology| year = 1954| volume = 28| issue = 1| title = Internal Shell Structures in the Middle Cambrian Gastropod Scenella and the Problematic Genus Stenothecoides| author = Rasetti| jstor = 1300208| pages = 59–66}} whilst the serially-repeated nature of these scars suggests to other authors a monoplacophoran affinity. However the specimens showing this scarring have not been convincingly shown to belong to the genus Scenella. A similarity to the Ediacaran Ovatoscutum has also been drawn.{{cite journal|author = Benjamin M. Waggoner|author2= Allen G. Collins|title = A new chondrophorine (Cnidaria, Hydrozoa) from the Cadiz Formation (Middle Cambrian) of California|journal = Paläontologische Zeitschrift|issn = 0031-0220|doi = 10.1007/BF02985970|pages = 7–17|issue = 1–2 |date = March 1995|volume = 69|bibcode= 1995PalZ...69....7W|s2cid= 128662109}}
Description
The shell of Scenella is elongated along its anterior-posterior axis, and comprises concentric rings around a conical central peak. Radial and concentric corrugations exist in some species. Some specimens are preserved as organic films, others appear to have been infilled with calcite. They are usually preserved point-upwards, with their long axes consistently oriented; this probably represents their most stable position under their depositional current. Soft parts have never been reported in association with Scenella, suggesting that the preserved fragments separated quickly from the associated tissue prior to burial.
Fossil occurrence
Scenella lived from the Cambrian to the Ordovician. Its remains have been found in Antarctica, Asia, Europe, and North America. Individual fossils are common throughout the Burgess shale, where they often occur in dense aggregations.{{cite journal
| last1=Babcock |first1=L. E.
| last2=Robison |first2=R. A.
| year = 1988
| title = Taxonomy and paleobiology of some Middle Cambrian Scenella (Cnidaria) and Hyolithids (Mollusca) from western North America
| url = http://kuscholarworks.ku.edu/dspace/bitstream/1808/3638/1/paleo.paper.121.pdf
| volume = 121 | pages = 1–22
| journal = University of Kansas Paleontological Contributions
| hdl=1808/3638}} Where they overlap, specimens deform as by draping. Specimens are sometimes cracked or torn, with margins often damaged by folding or "tattering". 1206 specimens of Scenella are known from the Greater Phyllopod bed, where they comprise 2.29% of the community.{{cite journal|last1=Caron |first1=Jean-Bernard|last2=Jackson |first2=Donald A.|title=Taphonomy of the Greater Phyllopod Bed community, Burgess Shale|journal=PALAIOS |volume=21 |issue=5 |pages=451–65|date=October 2006|doi=10.2110/palo.2003.P05-070R|jstor=20173022|bibcode=2006Palai..21..451C |s2cid=53646959 }}
Taxonomy
File:Scenella.jpg, Middle Cambrian, Walcott Quarry, near Field, British Columbia]]
Scenella is the only genus in the family Scenellidae. This family has no subfamilies and Scenella is the type genus of the family Scenellidae.
The taxonomy of the Gastropoda by Bouchet & Rocroi, 2005 categorizes Scenellidae in the superfamilia Scenelloidea within the Paleozoic molluscs of uncertain systematic position.
Species
Species in the genus Scenella include:[https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=7864 Scenella]. Paleobiology Database, accessed 16 August 2009.
References
{{Reflist}}
External links
{{Commons category|Scenella}}
- {{Cite web|date=2011|title=Scenella amii|work=Burgess Shale Fossil Gallery|publisher=Virtual Museum of Canada|url=http://burgess-shale.rom.on.ca/en/fossil-gallery/view-species.php?id=111|archive-url=https://web.archive.org/20201112025257/http://burgess-shale.rom.on.ca/en/fossil-gallery/view-species.php?id=111|archive-date=2020-11-12|url-status=dead}}
- [http://www.fieldmuseum.org/evolvingplanet/popUps/CO20.html Scenella] at the Field Museum's Evolving Planet
{{Taxonbar|from=Q3951822}}
Category:Burgess Shale fossils
Category:Prehistoric mollusc genera
Category:Cambrian invertebrates
Category:Ordovician invertebrates
Category:Extinct animals of Antarctica
Category:Paleozoic animals of Asia
Category:Prehistoric animals of Europe
Category:Paleozoic animals of North America
Category:Cambrian Series 2 first appearances
Category:Late Ordovician extinctions
Category:Paleozoic life of Ontario
Category:Paleozoic life of Newfoundland and Labrador