Scenella

{{Short description|Genus of molluscs (fossil)}}

{{Taxobox

| fossil_range = {{fossil range|Cambrian Stage 3|Late Ordovician|ref=}}

| image = Scenella tenuistriata.png

| image_caption = Dorsal view of the shell of Scenella tenuistriata

| image2 = Scenella tenuistriata 2.png

| image2_caption = Lateral view of the shell of Scenella tenuistriata

| regnum = Animalia

| phylum = Mollusca

| ordo =

| superfamilia = † Scenelloidea

| familia = † Scenellidae

| familia_authority = S. A. Miller, 1889Miller S. A. (after October 1889). North American geology and palaeontology: 389.

| genus= Scenella

| genus_authority = Billings, 1872

| subdivision_ranks = Species

| synonyms_ref = Bouchet P. & Rocroi J.-P. (Ed.); Frýda J., Hausdorf B., Ponder W., Valdés Á. & Warén A. 2005. [http://www.vliz.be/Vmdcdata/imis2/ref.php?refid=78278 Classification and nomenclator of gastropod families]. Malacologia: International Journal of Malacology, 47(1-2). ConchBooks: Hackenheim, Germany. {{ISBN|3-925919-72-4}}. ISSN 0076-2997. 397 pp.

| synonyms =

  • Helcionellinae Wenz, 1938
  • Hampilinae Kobayashi, 1958
  • Securiconidae Missarzhevsky, 1989

}}

Scenella is an extinct genus of fossil invertebrate animal which is generally considered to be a mollusc; at various times it has been suggested that this genus belongs with the gastropods, the monoplacophorans, or the helcionellids, although no firm association with any of these classes has been established.{{cite journal | year = 1984 | last1 = Yochelson | first1 = E. L. | last2 = Cid | first2 = D. G. L. | title = Reevaluation of the systematic position of Scenella | journal = Lethaia|volume =17 |issue =4 | pages = 331–340 | url = http://www3.interscience.wiley.com/journal/120036307/abstract | doi = 10.1111/j.1502-3931.1984.tb00679.x| doi-broken-date = 3 April 2025 | url-access = subscription }}{{dead link|date=February 2019|bot=medic}}{{cbignore|bot=medic}} An affinity with the hydrozoa (as a flotation device) has been considered, although some authors oppose this hypothesis.{{Cite journal| doi = 10.1017/s0022336000033837| journal = Journal of Paleontology| year = 1992| volume = 66| issue = 2| title = Scenella and "A Chondrophorine (Medusoid Hydrozoan) from the Basal Cambrian (Placentian) of Newfoundland"| author = Landing| author2 = Narbonne| jstor = 1305918| pages = 338| bibcode = 1992JPal...66..338L| s2cid = 132296800}} A gastropod affinity is defended on the basis of six pairs of internal muscle scars,{{Cite journal| journal = Journal of Paleontology| year = 1954| volume = 28| issue = 1| title = Internal Shell Structures in the Middle Cambrian Gastropod Scenella and the Problematic Genus Stenothecoides| author = Rasetti| jstor = 1300208| pages = 59–66}} whilst the serially-repeated nature of these scars suggests to other authors a monoplacophoran affinity. However the specimens showing this scarring have not been convincingly shown to belong to the genus Scenella. A similarity to the Ediacaran Ovatoscutum has also been drawn.{{cite journal|author = Benjamin M. Waggoner|author2= Allen G. Collins|title = A new chondrophorine (Cnidaria, Hydrozoa) from the Cadiz Formation (Middle Cambrian) of California|journal = Paläontologische Zeitschrift|issn = 0031-0220|doi = 10.1007/BF02985970|pages = 7–17|issue = 1–2 |date = March 1995|volume = 69|bibcode= 1995PalZ...69....7W|s2cid= 128662109}}

Description

The shell of Scenella is elongated along its anterior-posterior axis, and comprises concentric rings around a conical central peak. Radial and concentric corrugations exist in some species. Some specimens are preserved as organic films, others appear to have been infilled with calcite. They are usually preserved point-upwards, with their long axes consistently oriented; this probably represents their most stable position under their depositional current. Soft parts have never been reported in association with Scenella, suggesting that the preserved fragments separated quickly from the associated tissue prior to burial.

Fossil occurrence

Scenella lived from the Cambrian to the Ordovician. Its remains have been found in Antarctica, Asia, Europe, and North America. Individual fossils are common throughout the Burgess shale, where they often occur in dense aggregations.{{cite journal

| last1=Babcock |first1=L. E.

| last2=Robison |first2=R. A.

| year = 1988

| title = Taxonomy and paleobiology of some Middle Cambrian Scenella (Cnidaria) and Hyolithids (Mollusca) from western North America

| url = http://kuscholarworks.ku.edu/dspace/bitstream/1808/3638/1/paleo.paper.121.pdf

| volume = 121 | pages = 1–22

| journal = University of Kansas Paleontological Contributions

| hdl=1808/3638}} Where they overlap, specimens deform as by draping. Specimens are sometimes cracked or torn, with margins often damaged by folding or "tattering". 1206 specimens of Scenella are known from the Greater Phyllopod bed, where they comprise 2.29% of the community.{{cite journal|last1=Caron |first1=Jean-Bernard|last2=Jackson |first2=Donald A.|title=Taphonomy of the Greater Phyllopod Bed community, Burgess Shale|journal=PALAIOS |volume=21 |issue=5 |pages=451–65|date=October 2006|doi=10.2110/palo.2003.P05-070R|jstor=20173022|bibcode=2006Palai..21..451C |s2cid=53646959 }}

Taxonomy

File:Scenella.jpg, Middle Cambrian, Walcott Quarry, near Field, British Columbia]]

Scenella is the only genus in the family Scenellidae. This family has no subfamilies and Scenella is the type genus of the family Scenellidae.

The taxonomy of the Gastropoda by Bouchet & Rocroi, 2005 categorizes Scenellidae in the superfamilia Scenelloidea within the Paleozoic molluscs of uncertain systematic position.

Species

Species in the genus Scenella include:[https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=7864 Scenella]. Paleobiology Database, accessed 16 August 2009.

class="wikitable sortable"
Species

! Authority

! Range

Scenella affinis

| Ulrich and Schfield, 1897

|

Scenella amii

| (Matthew, 1902) Babcock and Robinson 1988

| Burgess Shale (Mid Cambrian, {{Ma|Burgess Shale}})

Scenella anomala

| (Billings, 1865) Wagner 2008

Scenella augusta

| (Billings, 1865) Wagner 2008

Scenella barrandei

| (Linnarsson, 1879) (Bergerg-Madsen and Peel, 1986){{Cite journal

| vauthors = BERGERG-MADSEN V, PEEL JS

| year = 1986

| title = Scenella barrandei (Mollusca) from the Middle Cambrian of Baltoscandia

| journal = Norsk Geologisk Tidsskrift

| pages = 81–86

| volume = 66

| issue = 1

}}

Scenella beloitensis

| Ulrich and Scofield (1897)

Scenella clotho

| Walcott, 1936

Scenella compressa

| Ulrich and Scofield (1897)

Scenella conica

| Whiteaves, 1884

Scenella conula

| (Walcott, 1884)

Scenella conicum

| (Whitfield, 1886) Ulrich and Scofield (1897)

Scenella hujingtanensis

| Yu, 1979

Scenella magnifica

| Ulrich and Scofield (1897)

S.? mira

| Vasil’yeva, 1994

Scenella montrealensis

| Billings (1865)

Scenella obtusa

| (Sardeson, 1892) Ulrich and Scofield (1897)

Scenella orithyia

| (Billings, 1865) Ulrich and Scofield (1897)

Scenella pretensa

| Raymond, 1905

Scenella radialis

| Ulrich and Scofield (1897)

Scenella radians

| Babcock & Robinson

Scenella radiata

| Yu, 1979

Scenella reticulata

| (Billings, 1872) (type){{cite journal|journal = Nat. Can.|series=2nd series|volume=vi|title=On some fossils from the primordial rocks of Newfoundland|pages=465–479|year=1872|last=Billings}}

Scenella retusa

| (Ford, 1873){{cite journal|journal=American Journal of Science|year=1873|author=Ford|title=On some new species of fossils from the Primordial or Potsdam group of Rensselaer County, New York (lower Potsdam)|series=3|volume=5|issue=27|pages=211–215|doi=10.2475/ajs.s3-5.27.211|bibcode=1873AmJS....5..211F|s2cid=130771815 |url=https://zenodo.org/record/1450106}}

Scenella unicarinata

| (Kobayashi, 1934) Wagner 2008

Scenella varians

| Walcott 1908

Scenella venillia

| (Billings, 1865) Wagner 2008

Scenella tenuistriata

| Chapman, 1911

| Mid CambrianChapman F. (1911). "New or Little Known Victorian Fossils in the National Museum". Proceedings of the Royal Society of Victoria, Melbourne [https://archive.org/details/proceedingsofroy23roya 23]: [https://archive.org/stream/proceedingsofroy23roya#page/n427/mode/2up 305]-324. plate 58-61.; Scenella tenuistriata at [https://archive.org/stream/proceedingsofroy23roya#page/312/mode/2up 313]-314, [https://archive.org/stream/proceedingsofroy23roya#page/n450/mode/1up plate LIX.] figure 18 a,b.

References

{{Reflist}}