Tim Birkhead

Birkhead was awarded a Bachelor's degree in Biology from Newcastle University in 1972, followed by a Doctor of Philosophy degree from University of Oxford in 1976 for research on the breeding biology and survival of guillemots Uria aalge supervised by E.K. Dunn and Chris Perrins.{{cite thesis |degree=DPhil |first=Timothy Robert|last=Birkhead |title=Breeding biology and survival of guillemots (Uria aalge) |publisher=University of Oxford |year=1976 |url=https://ora.ox.ac.uk/objects/uuid:05606087-da38-464f-8d70-b7af1ca05451|website=ora.ox.ac.uk |oclc=44837387|id={{EThOS|uk.bl.ethos.449886}} {{free access}}}} He was subsequently awarded a Doctor of Science from Newcastle in 1989.

Birkhead's research on promiscuity in birds redefined the mating systems of birds.{{citation needed|date=February 2018}} Focusing initially on the adaptive significance of male promiscuity and female promiscuity, he later switched to the study of mechanisms and resolved the mechanisms of sperm competition in birds. He provided some of the first evidence of: cryptic female choice in birds; strategic sperm allocation, and he also provided the first estimates of the quantitative genetics of sperm traits in birds.{{citation needed|date=February 2018}}

Birkhead's research also resolved the issue of polyspermy in birds and provided the first evidence for morphological sperm selection in the female reproductive tract.{{Cite journal|last1=Hemmings|first1=N.|last2=Birkhead|first2=T. R.|date=2015-11-07|title=Polyspermy in birds: sperm numbers and embryo survival|journal=Proceedings of the Royal Society B|language=en|volume=282|issue=1818|pages=20151682|doi=10.1098/rspb.2015.1682|issn=0962-8452|pmid=26511048|pmc=4650155}} His long term study of the population biology of common guillemots on the island of Skomer off Wales has run since 1972, and is currently in need of support.{{Cite journal|last=Birkhead|first=Tim|date=2014-10-23|title=Stormy outlook for long-term ecology studies|journal=Nature|language=en|volume=514|issue=7523|pages=405|doi=10.1038/514405a|pmid=25341754|bibcode=2014Natur.514..405B|doi-access=free}}{{Cite web|url=https://www.justgiving.com/fundraising/guillemotsskomer|title=Read Tim's story|author=Anon|website=justgiving.com|access-date=2018-01-08}}

His recent research is on the adaptive significance of egg shape in birds, including the common guillemot whose pyriform egg has long been thought to allow it to either spin-like- a-top or roll-in-an-arc to prevent it rolling off the cliff ledge. However, there is no evidence for either of these ideas.{{Cite journal|last1=Birkhead|first1=Tim R.|last2=Thompson|first2=Jamie E.|last3=Biggins|first3=John D.|date=2017-07-01|title=Egg shape in the Common Guillemot Uria aalge and Brunnich's Guillemot U. lomvia: not a rolling matter?|journal=Journal of Ornithology|language=en|volume=158|issue=3|pages=679–685|doi=10.1007/s10336-017-1437-8|bibcode=2017JOrni.158..679B |s2cid=9084008|issn=2193-7192|url=http://eprints.whiterose.ac.uk/112880/1/TRB%20et%20all%20G%20eggs%202.pdf}}{{Cite web|url=https://britishbirds.co.uk/article/vulgar-errors-point-guillemots-egg/|website=britishbirds.co.uk|title=Vulgar errors – the point of a Guillemot's egg - British Birds|access-date=2018-01-08|language=en-US}} Instead, Birkhead and colleagues have identified the main advantage of a pyriform shape: stability. The pyriform shape makes the egg inherently more stable, especially on the sloping surfaces on which guillemots commonly breed.{{Cite journal|last1=Birkhead|first1=Tim R.|last2=Thompson|first2=Jamie E.|last3=Montgomerie|first3=Robert|date=2018-10-01|title=The pyriform egg of the Common Murre (Uria aalge) is more stable on sloping surfacesEl huevo piriforme de Uria aalge es más estable en superficies inclinadasCommon Murre egg shape and stability|journal=The Auk|language=en|volume=135|issue=4|pages=1020–1032|doi=10.1642/AUK-18-38.1|s2cid=92507158|issn=0004-8038|doi-access=free}}

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• Nettleship, D. N. & Birkhead, T. R. (eds) (1985) The Atlantic Alcidae. Academic Press. pp 574. {{ISBN|9780125156714}}
• Birkhead, T. R. & Møller, A. P. (1992). Sperm Competition in Birds: Evolutionary Causes and Consequences. Academic Press. pp. 280. {{ISBN|9780121005405}}
• Birkhead, T. R. & Møller A. P. (eds) (1998) Sperm Competition and Sexual Selection. Academic Press. pp 826. {{ISBN|0121005437}}
• Birkhead, T. R., Hosken, D. & Pitnick, S. (eds) (2009). Sperm Biology: An Evolutionary Perspective. London: Academic Press. pp 642. {{ISBN|9780123725684}}
• Birkhead, T. R. (ed) (2016). Virtuoso by Nature: the Scientific Worlds of Francis Willughby FRS (1635-1672). Brill. pp 439. {{ISBN|9789004285316}}
• Promiscuity: An Evolutionary History of Sperm Competition{{cite book| title=Promiscuity: An Evolutionary History of Sperm Competition| publisher=Harvard University Press| year=2002| isbn=978-0-674-00666-9}}
• Promiscuity (Faber & Faber 2000), {{ISBN|9780571193608}} which makes the concept of post-copulatory sexual selection accessible to the non-specialist.
• Ten Thousand Birds: Ornithology since Darwin (Princeton 2014) (with J. Wimpenny and R. Montgomerie),{{cite journal|author=Bodio, Stephen J.|title=Book Review: Ten Thousand Birds: Ornithology since Darwin by Tim Birkhead, Jo Wimpenny, and Bob Montgomerie|date=Summer 2014|journal=Living Bird Magazine|url=https://www.allaboutbirds.org/news/book-review-ten-thousand-birds-ornithology-since-darwin-by-tim-birkhead-jo-wimpenny-and-bob-montgomerie/#}} {{ISBN|978-0-691-15197-7}}, won the USA Prose Award. PROSE Award (American Publishers Awards for Professional and Scholarly Excellence) for the best book in 2014 in the History of Science, Medicine and Technology category; CHOICE (magazine of the American Library Association) list of Outstanding Academic Titles, 2014 in Zoology; Runner-up for BB/BTO Best Bird Book of 2014.
• The red canary: the story of the first genetically engineered animal, Phoenix, 2004, {{ISBN|978-0-7538-1772-8}}, describes the power of selective breeding and how the interaction between professional scientist and an amateur bird-keeper created the red canary. The book won the Consul Cremer Prize (2003).
• {{cite book| title=The Wisdom of Birds: An Illustrated History of Ornithology| publisher=Bloomsbury| year= 2008| isbn=978-0-7475-9256-3}}; Bloomsbury Publishing, 2011, {{ISBN|978-0-7475-9822-0}}, The book describes how we know what we know about the biology of birds, focussing on evolutionary explanations. The Wisdom of Birds won the Best Bird book of the Year Award (2009) from the British Trust for Ornithology and British Birds.{{citation needed|date=February 2018}}
• {{cite book| title=The Magpies: The Ecology and Behaviour of Black-Billed and Yellow-Billed Magpies| publisher=A&C Black|year= 2010|isbn=978-1-4081-3777-2}}
• {{cite book| title=Great Auk Islands; a Field Biologist in the Arctic| publisher=A & C Black, 2011|isbn= 978-1-4081-3786-4| date=2010-10-30}}
• Bird Sense: What it Is Like to Be a Bird, Bloomsbury Publishing Plc, 2012, {{ISBN|978-1-4088-2013-1}} was rated best natural history book of 2012 by the Independent and Guardian Newspapers, and was awarded a Best Bird Book of 2012 prize by British Birds and the British Trust for Ornithology, and was short-listed for the Royal Society Winton Book Prize in 2013.[https://www.theguardian.com/books/2013/nov/26/royal-society-winton-prize-sean-carroll "Royal Society Winton Prize goes to 'rock star' science book"], "The Guardian", London, 26 November 2013. Retrieved on 8 May 2019.
• The Most Perfect Thing: the Inside (and Outside) of a Bird's Egg, Bloomsbury Publishing Plc, 2016,{{cite journal | last1=Ju | first1=Chenghui | last2=Lahti | first2=David C. | title=The Most Perfect Thing: Inside (and Outside) a Bird's Egg by Tim Birkhead. 2016. Bloomsbury, New York, NY, USA. xvi + 304 pp., 15 color and 3 black-and-white photographs. $27 (hardcover). ISBN 978-1632863690. | journal=The Auk | publisher=Oxford University Press (OUP) | volume=134 | issue=4 | year=2017 | issn=0004-8038 | doi=10.1642/auk-17-112.1 | pages=922–924| doi-access=free }}{{cite news|author=Preston, Alex|author-link=Alex Preston (author)|date=April 17, 2016|newspaper=The Guardian|title=Review of The Most Perfect Thing: Inside (and Outside) a Bird's Egg by Tim Birkhead|url=https://www.theguardian.com/books/2016/apr/17/most-perfect-thing-inside-birds-egg-tim-birkhead-review-alex-preston}} {{ISBN|978-1-4088-5126-5}} short-listed for the Royal Society Insight Investment Science Book Prize (2016),{{Cite web|url=https://royalsociety.org/news/2016/08/shortlist-for-science-book-prize-2016-unveiled/|title=Shortlist for The Royal Society Insight Investment Science Book Prize 2016 unveiled|website=royalsociety.org|access-date=2016-09-22}} and winner of the Zoological Society of London's 2017 prize for communicating zoology.{{citation needed|date=February 2018}} David Attenborough described it as ‘Magnificent: science without any high falutin’ technology’.{{citation needed|date=February 2018}}
• The Wonderful Mr Willughby: The First True Ornithologist, Bloomsbury Publishing Plc, 2018, {{ISBN|978-1-4088-7850-7}}
• [https://books.google.com/books/about/Birds_and_Us.html?id=4ylgEAAAQBAJ Birds and Us: A 12,000-Year History from Cave Art to Conservation], Princeton University Press, 2022.{{cite news|author=Freeman, Frank|title=Review: 'Birds and Us,' by Tim Birkhead|date=December 17, 2022|newspaper=Star Tribune|location=Minnesota|url=https://www.startribune.com/review-birds-and-us-by-tim-birkhead/600236396/}} {{isbn|978-0-691-23992-7}}{{cite web|title=Review of Birds and Us by Tim Birkhead|website=Kirkus Reviews|year=2022|url=https://www.kirkusreviews.com/book-reviews/tim-birkhead/birds-and-us/}}

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Birkhead has combined his enthusiasm for research with a passion for undergraduate teaching. He has taught courses on ecology, evolution, statistics, birds, behavioural ecology, animal behaviour and the history and philosophy of science. His teaching has been recognised by four awards, including a National Teaching Fellowship in 2017.

Starting in 1992 and continuing until 2015 (when he handed over to a steering group) Birkhead organised (with Professor Harry Moore) a small (~60) biennial meeting on reproductive biology in the Peak District National Park known as Biology of Spermatozoa (BoS). Delegates are from a diverse range of backgrounds and include clinicians, reproductive physiologists, andrologists, theoreticians and evolutionary biologists. The format and interdisciplinary nature of the meeting was successful in terms of exchanging ideas, techniques and establishing collaborations.{{Cite web|url=http://www.bos.group.shef.ac.uk/|title=BoS {{!}} Biology of Spermatozoa meetings|website=www.bos.group.shef.ac.uk|language=en-US|access-date=2018-01-08}}

Between 2002 and 2010 Birkhead had a monthly column in Times Higher Education.{{cite web|url=http://www.timeshighereducation.co.uk/story.asp?storycode=400029|title=Tim Birkhead|date=4 January 2008|publisher=}} His articles were concerned with various aspects of higher education: undergraduate teaching, administration and, occasionally, research.

He has written for The Guardian,{{citation needed|date=February 2018}} The Independent,{{citation needed|date=February 2018}} the BBC, The Biologist, Natural History and Evolve.{{citation needed|date=February 2018}}

He has featured on numerous BBC Radio 4 programmes, including Start the Week — with Jeremy Paxman;{{when|date=February 2018}} The Life Scientific with Jim Al-Khalili;{{Cite web|url=http://www.bbc.co.uk/programmes/b098j5l3|website=bbc.co.uk|publisher=BBC|title=Tim Birkhead on bird promiscuity|first=Jim|last=Al-Khalili|author-link=Jim Al-Khalili|year=2017|access-date=2018-01-08}} The Infinite Monkey Cage in 2018.{{citation needed|date=February 2018}} His book ‘The Most Perfect Thing’ provided the basis for the TV documentary ‘Attenborough’s Eggs’ introduced by David Attenborough (2018).{{citation needed|date=February 2018}} Birkhead has been honorary curator of the Alfred Denny Museum in the University of Sheffield between 1980-2018.{{Cite web|url=http://www.shef.ac.uk/alfred-denny-museum|title=Alfred Denny Museum - Alfred Denny Museum - The University of Sheffield|last=Sheffield|first=University of|website=www.shef.ac.uk|language=en-GB|access-date=2018-01-08}}{{Citation|last=The University of Sheffield|title=Aisha's Letter|date=2017-02-10|url=https://www.youtube.com/watch?v=Ca9NCtXgN-c|accessdate=2018-01-08}}

He has given numerous public lectures, including at Café Scientique, the Cheltenham Science Festival and numerous literary festivals including Ways with Words (Sheffield) and Hay on Wye. His TED (conference) lecture on the history of ornithology has been viewed over 100,000 times.{{Citation|last=Birkhead|first=Tim|title=The early birdwatchers|date=26 September 2010 |url=https://www.ted.com/talks/tim_birkhead_the_wisdom_of_birds|language=en|accessdate=2018-01-08}}

Over the course of his career, Birkhead has received a number of awards:

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• McColvin Medal for best reference book: Cambridge Encyclopaedia of Ornithology in 1991
• President, International Society for Behavioural Ecology (1996-1998)
• Brockington Visitorship, Queens University, Canada in 2003{{citation needed|date=February 2018}}
• Consul Cremer Prize for The Red Canary in 2003
• Elected a Fellow of the Royal Society (FRS) in 2004
• ISIHighlyCited.com - Designated Highly Cited Researcher Plant & Animal Science, 2004{{citation needed|date=February 2018}}
• Senate Award for Sustained Excellence in Teaching, University of Sheffield, 2007.
• Animal & Plant Sciences ‘Teacher of the Year’, 2009.{{citation needed|date=February 2018}}
• Winner: Bird Book of the Year Award, for The Wisdom of Birds, from the British Trust for Ornithology and British Birds, 2009.
• Elected Honorary Member of the American Ornithologists Union, 2010.{{citation needed|date=February 2018}}
• Elected Honorary Member of the Linnaean Society of New York, 2011
• Elliot Coues Medal, American Ornithologists Union in 2011{{citation needed|date=February 2018}}{{cite journal | title=Elliott Coues Award, 2011 | journal=The Auk | publisher=Oxford University Press (OUP) | volume=129 | issue=1 | year=2012 | issn=0004-8038 | doi=10.1525/auk.2012.129.1.187 | pages=187–188| doi-access=free }}
• ASAB (Association for the Study of Animal Behaviour) Medal in 2012{{citation needed|date=February 2018}}
• President of the Association for the Study of Animal Behaviour 2013-16
• Vice-president of the British Trust for Ornithology in 2012{{citation needed|date=February 2018}}
• Winner of the Society for Biology, Bioscience Teacher of the Year, 2013.{{Cite web|url=http://shef.ac.uk/news/nr/tim-birkhead-%20bioscience-teacher-%20society-%20biology-1.269973|title=Professor named top of the class - Latest - News - The University of Sheffield|last=Sheffield|first=University of|website=shef.ac.uk|language=en-GB|access-date=2018-01-08}}
• Zoological Society of London, Silver Medal, 2014.
• Spallazani Medal, Biology of Spermatozoa community, 2015.
• Eisenmann Medal, the Linnaean Society of New York, 2016.{{citation needed|date=February 2018}}
• Godman-Salvin Medal from the British Ornithologists' Union, 2016.
• Founders’ Medal of the Society for the Study of the History of Natural History (SHNH), 2016.
• Winner of the Zoological Society of London’s Award for Communicating Zoology to a general audience for The Most Perfect Thing, 2017.
• Stephen Jay Gould Prize for increasing public understanding of evolutionary biology, Evolution Society, 2017.{{citation needed|date=February 2018}}
• Awarded a National Teaching Fellowship by the Higher Education Academy (HEA) in 2017.
• Elected Honorary Member of the (Deutschen Ornitholgen-Gesellschaft [DO-G] German Ornithological Society), 2017{{citation needed|date=February 2018}}

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Category:British zoologists

Category:Academics of the University of Sheffield

Category:Alumni of the University of Oxford

Category:Fellows of the Royal Society

Category:Alumni of Newcastle University

Category:Living people

Category:1950 births

Category:Place of birth missing (living people)

Category:British science writers

Category:People educated at Leeds Grammar School