Trioceros

{{Short description|Genus of lizards}}

{{automatic taxobox

| image = Jackson's Chameleon444.jpg

| image_caption = Jackson's chameleon
(Trioceros jacksonii)

| taxon = Trioceros

| authority = Swainson, 1839

| type_species = Chamaeleo oweni

| type_species_authority = Gray, 1831{{NRDB species |genus=Trioceros |species=oweni |access-date=17 May 2022}}

}}

Trioceros is a genus of lizards in the family Chamaeleonidae, the chameleons, native to lowlands and highlands in the African mainland, ranging from Ethiopia south to Mozambique and west as far as Ghana. Trioceros was considered a subgenus of the genus Chamaeleo until 2009, when it was elevated to full genus level.{{aut|Tilbury, C.R., & Tolley, K.A.}} (2009). "A re-appraisal of the systematics of the African genus Chamaeleo (Reptilia: Chamaeleonidae)". Zootaxa 2079: 57–68.

Trioceros vary greatly in appearance and size. Many species in the genus have various ornaments, such as one to four horns on the head, crests on the nape or throat, or spines or sail-like structures on the top of the back or tail. They are primarily found in adult males, but generally reduced or even absent in females, and there are also many species in the genus where both sexes lack conspicuous ornaments. Although horn-like structures are found in certain other chameleon genera, Trioceros is the only where it can be cylindrical, annulated and bony.

Trioceros contains species that lay eggs (similar to most other chameleon genera) and species that give birth to live young (similar to only Bradypodion). It is likely that giving birth to live young is an adaption to temperature, as it in Trioceros generally is found in species from highlands. The relatively cold highland climate slows the development of eggs laid in the ground; when instead retained inside the body until birth, a female can actively sun bask to increase the temperature.{{cite journal|last1=Hughes|first1=D.F.|last2=Blackburn|first2=D.G.|year=2020|title=Evolutionary origins of viviparity in Chamaeleonidae|journal=Journal of Zoological Systematics and Evolutionary Research|volume=58|issue=1|pages=284–302|doi=10.1111/jzs.12328|doi-access=free}}

Species

The following species are recognized as being valid.{{NRDB genus |genus=Trioceros |access-date= }}

class="wikitable"
ImageNameCommon NameSubspeciesDistribution
150pxTrioceros affinis
{{small|(Rüppell, 1845)}}
beardless Ethiopian montane chameleon, Rüppell's desert chameleonEthiopia.
150pxTrioceros balebicornutus
{{small|(Tilbury, 1998)}}
Bale two-horned chameleonEthiopia.
150pxTrioceros bitaeniatus
{{small|(Fischer, 1884)}}
side-striped chameleon, two-lined chameleonEthiopia, southern Sudan, Somalia, Kenya, Tanzania, Uganda, and the northeastern Democratic Republic of the Congo.
150pxTrioceros camerunensis
{{small|(L. Müller, 1909)}}
Cameroon dwarf chameleonCameroon.
Trioceros chapini
{{small|(de Witte, 1964)}}
Chapin's chameleon,Beolens et al., p. 51. grey chameleonGabon and Democratic Republic of the Congo.
Trioceros conirostratus
{{small|(Tilbury, 1998)}}
South Sudanese unicorn chameleonSouth Sudan and Uganda.
150pxTrioceros cristatus
{{small|(Stutchbury, 1837)}}
crested chameleon, fringed chameleonBioko, Equatorial Guinea, Cameroon, Central African Republic, Democratic Republic of the Congo, Republic of the Congo, Gabon, Nigeria, Ghana, and Togo.
150pxTrioceros deremensis
{{small|(Matschie, 1892)}}
Usambara giant three-horned chameleon, wavy chameleonEast Usambara, Uluguru, Nguu and Nguru Mountains, and Udzungwa Mountains.
150pxTrioceros ellioti
{{small|(Günther, 1895)}}
Elliot's chameleon,Beolens et al., p. 82. montane side-striped chameleon, Elliot's groove-throated chameleonBurundi, Kenya, South Sudan, Rwanda, Tanzania, Uganda, and Democratic Republic of the Congo.
150pxTrioceros feae
{{small|(Boulenger, 1906)}}
Bioko hornless chameleon, Bioko montane chameleon, Fea's chameleonBioko.
Trioceros fuelleborni
{{small|(Tornier, 1900)}}
(named after Friedrich Fülleborn),Beolens et al., p.95. flapjack chameleon, Ngosi Volcano chameleon, Poroto three-horned chameleon, mountain three-horned chameleonTanzania
Trioceros goetzei
{{small|(Tornier, 1899)}}
Goetze's chameleon,Beolens et al., p. 103. Ilolo chameleon, Goetze's whistling chameleon*Trioceros goetzei goetzei {{small|(Tornier, 1899)}} – Goetze's whistling chameleon

  • Trioceros goetzei nyikae {{small|(Loveridge, 1953)}} – Nyika whistling chameleon
Tanzania and Malawi.
Trioceros hanangensis
{{small|Krause & {{Interlanguage link|Wolfgang Böhme (herpetologist)|de|Wolfgang Böhme (Zoologe)|lt=Böhme}}, 2010}}
Mount Hanang montane dwarf chameleon, Mount Hanang chameleonTanzania
Trioceros harennae
{{small|(Largen, 1995)}}
Harenna hornless chameleon*Trioceros harennae harennae {{small|(Largen, 1995)}} – Harenna hornless chameleon
  • Trioceros harennae fitchi {{small|(Nečas, 2004)}} – Fitch's Harenna hornless chameleonBeolens et al., p. 90.
  • Ethiopia
    150pxTrioceros hoehnelii
    {{small|(Steindachner, 1891)}}
    helmeted chameleon, high-casqued chameleon, von Höhnel's chameleonBeolens et al., p. 124.Kenya and Uganda
    Trioceros incornutus
    {{small|(Loveridge, 1932)}}
    Ukinga hornless chameleonTanzania
    Trioceros ituriensis
    {{small|(K.P. Schmidt, 1919)}}
    Ituri forest chameleonDemocratic Republic of the Congo and Kenya.
    150pxTrioceros jacksonii
    {{small|(Boulenger, 1896)}}
    Jackson's chameleonBeolens et al., p. 132.*Trioceros jacksonii jacksonii {{small|(Boulenger, 1896)}} – Jackson's three-horned chameleon
  • Trioceros jacksonii merumontanus {{small|(Rand, 1958)}} – dwarf Jackson's chameleon
  • Trioceros jacksonii xantholophus {{small|(Eason, Ferguson & Hebrard, 1988)}} – Mount Kenya three-horned chameleon, yellow-crested Jackson's chameleon
  • south-central Kenya and northern Tanzania.
    150pxTrioceros johnstoni
    {{small|(Boulenger, 1901)}}
    Johnston's chameleon,Beolens et al., p. 135. Johnston's three-horned chameleon, Ruwenzori three-horned chameleonDemocratic Republic of the Congo, Burundi, Rwanda, and Uganda
    Trioceros kinangopensis
    {{small|Stipala et al., 2012}}
    Aberdare Mountains dwarf chameleonKenya.
    Trioceros kinetensis
    {{small|(K.P. Schmidt, 1943)}}
    Mount Kineti montane dwarf chameleon, Mount Kineti chameleonSouth Sudan.
    Trioceros laterispinis
    {{small|(Loveridge, 1932)}}
    spiny-flanked chameleonTanzania.
    Trioceros marsabitensis
    {{small|(Tilbury, 1991)}}
    Marsabit one-horned chameleon, Mt. Marsabit chameleon, Tilbury's chameleonKenya.
    150pxTrioceros melleri
    {{small|(Gray, 1865)}}
    giant one-horned chameleon, Meller's chameleon,Beolens et al., p. 175. Meller's giant one-horned chameleonMalawi, northern Mozambique, and Tanzania
    150pxTrioceros montium
    {{small|(Buchholz, 1874)}}
    Cameroon sailfin chameleonCameroon.
    Trioceros narraioca
    {{small|(Nečas, Modrý & Šlapeta, 2003)}}
    Mount Kulal chameleon or Mount Kulal stump-nosed chameleonKenya
    Trioceros ntunte
    {{small|(Nečas, Modry & Slapeta, 2005)}}
    Mount Nyiru chameleon, Nyiru montane dwarf chameleonKenya
    Trioceros nyirit
    {{small|Stipala et al., 2011}}
    Mount Mtelo stump-nosed chameleon, Pokot chameleonKenya
    150pxTrioceros oweni
    {{small|(Gray, 1831)}}
    Owen's chameleonBeolens et al., p. 198.Nigeria in the north, to Angola in the south, and Burundi in the east
    Trioceros perreti
    {{small|(Klaver & Böhme, 1992)}}
    Perret's chameleon,Beolens et al., p. 203. Perret's montane chameleonCameroon.
    150pxTrioceros pfefferi
    {{small|(Tornier, 1900)}}
    Pfeffer's two-horned chameleon,Beolens et al., p. 205, Pfeffer's chameleon, Bakossi two-horned chameleonCameroon.
    150pxTrioceros quadricornis
    {{small|(Tornier, 1899)}}
    four-horned chameleon*Trioceros quadricornis quadricornis {{small|(Tornier, 1899)}} – southern four-horned chameleon
  • Trioceros quadricornis eisentrauti {{small|(Mertens, 1968)}} – Rumpi Hills chameleon
  • Trioceros quadricornis gracilior
    {{small|(Böhme & Klaver, 1981)}} – northern four-horned chameleon
  • western Cameroon and southeastern Nigeria.
    150pxTrioceros rudis
    {{small|(Boulenger, 1906)}}
    coarse chameleon, rough chameleon, Ruwenzori side-striped chameleon, Rwenzori bearded montane dwarf chameleonwestern Uganda, Rwanda, Burundi, and eastern DR Congo
    Trioceros schoutedeni
    {{small|(Laurent, 1952)}}
    Schouteden's montane dwarf chameleonBeolens et al., p. 237.Rwanda and Democratic Republic of the Congo.
    Trioceros schubotzi
    {{small|(Sternfeld, 1912)}}
    Mount Kenya montane dwarf chameleon, Mount Kenya side-striped chameleon, Schubotz's chameleonBeolens et al., p. 238.Kenya
    Trioceros serratus
    {{small|(Mertens, 1922)}}
    Cameroon
    Trioceros sternfeldi
    {{small|(Rand, 1963)}}
    (named after Richard Sternfeld),Beolens et al., 253. Crater Highlands side-striped chameleon, Tanzanian montane dwarf chameleonTanzania
    150pxTrioceros tempeli
    {{small|(Tornier, 1899)}}
    Tanzania mountain chameleon, Tempel's chameleon,Beolens et al., p. 263. Udzungwa double-bearded chameleonTanzania
    150pxTrioceros werneri
    {{small|(Tornier, 1899)}}
    Werner's chameleon,Beolens et al., p. 282. Wemer's chameleon, Wemer's three-horned chameleonTanzania
    150pxTrioceros wiedersheimi
    {{small|(Nieden, 1910)}}
    Mount Lefo chameleon, Wiedersheim's chameleonBeolens et al., p. 285.Cameroon and Nigeria
    150pxTrioceros wolfgangboehmei
    {{small|Koppetsch, Nečas & Wipfler, 2021}}
    Ethiopia.

    Nota bene: In the above list, a binomial authority or trinomial authority in parentheses indicates that the species or subspecies was originally described in a genus other than Trioceros.

    Footnotes

    {{Reflist|20em}}

    References

    • {{aut|Boelens, Bo; Watkins, Michael; Grayson, Michael}} (2011). The Eponym Dictionary of Reptiles. Baltimore: Johns Hopkins University Press. xiii + 296 pp. {{ISBN|978-1-4214-0135-5}}.
    • {{aut|Swainson, W.}} (1839). The Natural History of Fishes, Amphibians, & Reptiles, or Monocardian Animals. Vol. II. London: Longman, Orme, Brown, Green & Longmans; John Taylor. (A. Spottiswoode, printer). 452 pp. (Trioceros, new genus, p. 369).

    {{Taxonbar|from=Q1758255}}

    Category:Lizard genera

    Category:Taxa named by William Swainson