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Image:Homing pigeon.jpg. These have been selectively bred to home over long distances.]]

Homing is the inherent ability of an animal to navigate towards an original location through unfamiliar areas. This location can may be either a home territory, or a breeding spot.

History

Uses

Homing abilities can be used to find the way back to home in a migration. It is often used in reference to going back to a breeding spot seen years before, as in the case of Salmon. Homing abilities can also be used to go back to familiar territory when displaced over long distances, such as with the Red-bellied newt.

True navigation

Image:Caretta caretta 060417w2.jpgs home both using true navigation and magnetic orientation]] Some animals use true navigation for their homing. This means in familiar areas they will use landmarks such as roads, rivers or mountains when flying, or islands and other landmarks while swimming. However, this only works in familiar territory. Pigeons, for example, will often navigate using familiar landmarks, such as roads.[http://news.bbc.co.uk/2/hi/uk_news/3460977.stm BBC News: Pigeons reveal map-reading secret] Sea turtles will also use landmarks to orient themselves.Avens, L., (2003) "Homing Behavior, Navigation, and Orientation of Juvenile Sea Turtles" [http://www.sefsc.noaa.gov/PDFdocs/Avens_Dissertation_2003.pdf ]

Magnetic orientation

Many animals use magnetic orientation to find their way home. This is usually used together with other methods, such as a sun compass, as in the case of pigeons and turtles. This is also commonly used when no other methods are available. As in the case of lobsters,Lohmann, K., Pentcheff, N., Nevitt, G., Stetten, G., Zimmer-Faust, R., Jarrard H., and Boles, L., (1995) Magnetic orientation of spiny lobsters in the ocean: experiments with undersea coil systems The Journal of Experimental Biology 198(10); pg. 2041-2048 which live underwater, and mole rats,Kimchi, T., and Terkel, J., (2001) Magnetic compass orientation in the blind mole rat Spalax ehrenbergi The Journal of Experimental Biology 204(4); pg. 751-758 which live underground.

Celestial orientation

Celestial orientation, navigation using the stars, is commonly used for homing. Displaced Marbled Newts, for example, can only home when stars are visible.Diego-Rasilla, J., and Luengo, R., (2002) Journal of Ethology 20(1):137-141

Topographic memory

Topographic memory, memory of the contours surrounding the destination, is one common method for navigation. This is mainly used my animals with less intelligence, such as molluscs. Limpets use this to find their way back to the home scrape. Whether this is true homing, in the limpet's case, has been disputed.Villee, C., and Groody, T., (1940) American Midland Naturalist; 24(1):190-204

References

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