Yezo Group

{{distinguish|Yeso Group}}

{{Infobox rockunit

| name = Yezo Group

| image =

| caption =

| type = Stratigraphic group

| age = {{Fossilrange|Aptian|Paleocene}}

| period = Late Cretaceous

| prilithology = Mudstone, sandstone

| otherlithology = Tuffite, conglomerate

| namedfor =

| namedby =

| region = Hokkaido
Sakhalin

| country = Japan, Russia

| coordinates =

| unitof =

| subunits = Shuparogawa Formation, Maruyama Formation, Hikagenosawa Formation, Saku Formation, Mikasa Formation, Kashima Formation, Haborogawa Formation, Osoushinai Formation, Hakobuchi Formation

| underlies = Ishikari Group, Poronai Group, Neogene deposits

| overlies = Sorachi Group

| thickness = {{Convert|10000|m|ft|-1}}

| extent =

| area =

| map =

| map_caption =

}}

The Yezo Group is a stratigraphic group in Hokkaido, Japan and Sakhalin, Russia which is primarily Late Cretaceous in age (Aptian to Earliest Paleocene). It is exposed as roughly north–south trending belt extending 1,500 kilometres through central Hokkaido from Urakawa to Cape Sōya and Sakhalin from the south coast to Alexandrovsk-Sakhalinsky District.{{Cite journal|last1=Shigeta|first1=Yasunari|last2=Maeda|first2=Haruyoshi|date=2005|title=Yezo Group Research in Sakhalin—A Historical Review|url=https://www.kahaku.go.jp/english/research/researcher/papers/18151.pdf|journal=The Cretaceous System in the Makarov Area, Southern Sakhalin, Russian Far East|publisher=National Science Museum Monographs|volume=31|pages=1–24}} It consists of marine forearc basin sediments, typically turbiditic and bioturbated mudstones and sandstones with subordinate conglomerate primarily deposited on the continental shelf and slope of the ancient Yezo subduction margin. It forms a continuous depositional sequence with the Sorachi Group, which overlies the Horokanai Ophiolite. The sequence gradually shallows upwards with the terminal Hakobuchi Formation representing a fluvial-inner shelf environment.{{Cite journal|last1=Takashima|first1=Reishi|last2=Kawabe|first2=Fumihisa|last3=Nishi|first3=Hiroshi|last4=Moriya|first4=Kazuyoshi|last5=Wani|first5=Ryoji|last6=Ando|first6=Hisao|date=June 2004|title=Geology and stratigraphy of forearc basin sediments in Hokkaido, Japan: Cretaceous environmental events on the north-west Pacific margin|journal=Cretaceous Research|volume=25|issue=3|pages=365–390|doi=10.1016/j.cretres.2004.02.004|bibcode=2004CrRes..25..365T |hdl=2115/17182|hdl-access=free}}

Numerous fossils are known from the unit, mostly ammonites and bivalves, but also marine vertebrates such as mosasaurs, plesiosaurs{{Cite journal|last1=Kaim|first1=Andrzej|last2=Kobayashi|first2=Yoshitsugu|last3=Echizenya|first3=Hiroki|last4=Jenkins|first4=Robert G.|last5=Tanabe|first5=Kazushige|title=Chemosynthesis-Based Associations on Cretaceous Plesiosaurid Carcasses|url=https://www.academia.edu/1427886|journal=Acta Palaeontologica Polonica|year=2008 |volume=53|issue=1|pages=97–104|doi=10.4202/app.2008.0106 |s2cid=128406537 |issn=0567-7920|doi-access=free}} and marine turtles. Dinosaur remains are among the fossils that have been recovered from the group. These include a partial cervical vertebra and right hand from the therizinosaurid Paralitherizinosaurus, from Early Campanian Osoushinai Formation from northern Hokkaido.{{Cite journal |last1=Kobayashi |first1=Yoshitsugu |last2=Takasaki |first2=Ryuji |last3=Fiorillo |first3=Anthony R. |last4=Chinzorig |first4=Tsogtbaatar |last5=Hikida |first5=Yoshinori |date=2022-05-03 |title=New therizinosaurid dinosaur from the marine Osoushinai Formation (Upper Cretaceous, Japan) provides insight for function and evolution of therizinosaur claws |journal=Scientific Reports |language=en |volume=12 |issue=1 |pages=7207 |doi=10.1038/s41598-022-11063-5| pmid=35504901 |pmc=9065154 |bibcode=2022NatSR..12.7207K |issn=2045-2322}}{{Cite journal|last1=Murakami|first1=Mizuki|last2=Hirayama|first2=Ren|last3=Hikida|first3=Yoshinori|last4=Hirano|first4=Hiromichi|date=December 2008|title=A theropod dinosaur (Saurischia: Maniraptora) from the Upper Cretaceous Yezo Group of Hokkaido, Northern Japan|journal=Paleontological Research|volume=12|issue=4|pages=421–425|doi=10.2517/prpsj.12.421|s2cid=128744799|issn=1342-8144}} Nipponosaurus is known from an unnamed unit of the group from Southern Sakhalin, probably late Santonian or early Campanian in age.Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Cretaceous, Asia)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 593-600. {{ISBN|0-520-24209-2}} Kamuysaurus, which is known from the early Maastrichtian Hakobuchi Formation of southern Hokkaido, was also discovered in layers of this group.{{Cite journal|last1=Sakurai|first1=Kazuhiko|last2=Sato|first2=Tamaki|last3=Chinzorig|first3=Tsogtbaatar|last4=Tanaka|first4=Kohei|last5=Fiorillo|first5=Anthony R.|last6=Chiba|first6=Kentaro|last7=Takasaki|first7=Ryuji|last8=Nishimura|first8=Tomohiro|last9=Kobayashi|first9=Yoshitsugu|date=September 5, 2019|title=A New Hadrosaurine (Dinosauria: Hadrosauridae) from the Marine Deposits of the Late Cretaceous Hakobuchi Formation, Yezo Group, Japan|journal=Scientific Reports|volume=9|issue=1|page=12389|doi=10.1038/s41598-019-48607-1|pmid=31488887|pmc=6728324|bibcode=2019NatSR...912389K|doi-access=free}}

Fossil content

=Reptiles=

class="wikitable" align="center"
colspan="6" align="center" | Reptiles reported from the Yezo Group
GenusSpeciesPresence

!Material

NotesImages
Anomalochelys

|A. angulata

|Hobetsu-cho, Hokkaido. (Cenomanian){{cite journal|author1=Ren Hirayama |author2=Kazuhiko Sakurai |author3=Tsutomu Chitoku |author4=Gentaro Kawakami |author5=Norio Kito |year=2001 |title=Anomalochelys angulata, an unusual land turtle of family Nanhsiungchelyidae (Superfamily Treionychoidea; Order Testudines) from the Upper Cretaceous of Hokkaido, North Japan |journal=Russian Journal of Herpetology |volume=8 |issue=2 |pages=127–138 |url=http://rjh.folium.ru/contents/2001/2001-02.htm |url-status=dead |archiveurl=https://web.archive.org/web/20110826064544/http://rjh.folium.ru/contents/2001/2001-02.htm |archivedate=2011-08-26 }}

|

|A nanhsiungchelyid.

|

Chupkaornis

|C. keraorum

|Kashima Formation (Coniacian to Santonian).{{Cite journal |last1=Tanaka |first1=Tomonori |last2=Kobayashi |first2=Yoshitsugu |last3=Kurihara |first3=Ken'ichi |last4=Fiorillo |first4=Anthony R. |last5=Kano |first5=Manabu |date=2018-06-21 |title=The oldest Asian hesperornithiform from the Upper Cretaceous of Japan, and the phylogenetic reassessment of Hesperornithiformes |url=https://doi.org/10.1080/14772019.2017.1341960 |journal=Journal of Systematic Palaeontology |volume=16 |issue=8 |pages=689–709 |doi=10.1080/14772019.2017.1341960 |bibcode=2018JSPal..16..689T |s2cid=134930629 |issn=1477-2019|url-access=subscription }}

|Cervical vertebrae, torso vertebrae, femur, and fibula.

|A hesperornithiform.

|File:Chupkaornis fossils.jpg

Kamuysaurus

|K. japonicus

|Hakobuchi Formation.

|"A nearly complete skeleton with the skull and mandible (HMG-1219)".

|A hadrosaur.

|File:Kamuysaurus.jpg

rowspan=2|Mesodermochelys

|M. undulatus

|Hakobuchi Formation{{Cite journal |last1=Hirayama |first1=Ren |last2=Chitoku |first2=Tsutomu |date=1996 |title=1022 Family Dermochelyidae (Superfamily Chelonioidea) from the Upper Cretaceous of North Japan |url=https://www.jstage.jst.go.jp/article/prpsj1951/1996/184/1996_184_597/_article/-char/ja/ |journal=日本古生物学會報告・紀事 新編 |volume=1996 |issue=184 |pages=597–622 |doi=10.14825/prpsj1951.1996.184_597}} and upper part of the Osoushinai Formation.{{Cite journal |last1=Hirayama |first1=Ren |last2=Hikida |first2=Yoshinori |date=March 1998 |title=Mesodermochelys (Testudines; Chelonioidea; Dermochelyidae) from the Late Cretaceous of Nakagawa-cho, Hokkaido, North Japan |url=https://www.researchgate.net/publication/256198184 |journal=Bulletin of the Nakagawa Museum of Natural History |volume=1 |pages=69 to 76}}

| -NMV-3, a proximal half of right humerus (Osoushinai Formation).

|A dermochelyid turtle.

|File:Mesodermochelys undulatus at Hobetsu Museum in Mukawa, Hokkaido.jpg

M. sp.

|Osoushinai Formation.

|A carapace with boreholes likely made by bivalves, and associated with many mollusc remains.

|A dermochelyid turtle.

|

Mosasaurus

|M. hobetsuensis

|Hakobuchi Formation.{{Cite journal |last1=Konishi |first1=Takuya |last2=Caldwell |first2=Michael W. |last3=Nishimura |first3=Tomohiro |last4=Sakurai |first4=Kazuhiko |last5=Tanoue |first5=Kyo |date=2016-10-02 |title=A new halisaurine mosasaur (Squamata: Halisaurinae) from Japan: the first record in the western Pacific realm and the first documented insights into binocular vision in mosasaurs |url=https://doi.org/10.1080/14772019.2015.1113447 |journal=Journal of Systematic Palaeontology |volume=14 |issue=10 |pages=809–839 |doi=10.1080/14772019.2015.1113447 |bibcode=2016JSPal..14..809K |s2cid=130644927 |issn=1477-2019|url-access=subscription }}

|

|A mosasaurine mosasaur.

|File:Carcass of Kamuysaurus.jpg

Nipponosaurus

|N. sachalinensis

|Sakhalin.{{Cite web |title=HUSCAP journal |url=https://eprints.lib.hokudai.ac.jp/journals/item.php?item=34225&handle=2115_35768&jname=269&vname=3749 |access-date=2022-08-15 |website=eprints.lib.hokudai.ac.jp}}

|

|A hadrosaur.

|File:Nipponosaurus dinosaur.png

Nodosauridae

|Indeterminate

|Hikagenosawa Formation. (Cenomanian){{cite journal |last1=Hawakaya |first1=Hiroshi |last2=Manabe |first2=Makoto |last3=Carpenter |first3=Kenneth |title=Nodosaurid Ankylosaur from the Cenomanian of Japan |journal=Journal of Vertebrate Paleontology |date=11 March 2005 |volume=25 |issue=1 |pages=240–245 |doi=10.1671/0272-4634(2005)025[0240:NAFTCO]2.0.CO;2}}

|Left rear half of the skull (MCM A522)

|

|

Paralitherizinosaurus

|P. japonicus

| Osoushinai Formation.

|Partial vertebra and partial right hand.

| A therizinosaur.

|File:Paralitherizinosaurus Restoration.png

Phosphorosaurus

|P. ponpetelegans

|Hakobuchi Formation.

|A well-preserved skull with some associated vertebrae.

|A halisaurine mosasaur with proportionately large eyes and binocular vision.

|File:Phosphorosaurus_ponpetelegans.jpg

Pliosauridae gen. et. sp. indet.

|Indeterminate

|Cenomanian to Turonian localities in Hokkaido.{{Cite journal |last1=Sato |first1=Tamaki |last2=Nagai |first2=Katsuhisa |last3=Echizenya |first3=Hiroki |last4=Shinmura |first4=Tatsuya |last5=Hikida |first5=Yoshinori |last6=Soeda |first6=Yuji |date=2023-05-26 |title=Pliosaurid (Reptilia: Sauropterygia) remains from the Upper Cretaceous of Japan, and their biostratigraphic and paleogeographic significance |journal=Cretaceous Research |volume=150 |language=en |pages=105593 |doi=10.1016/j.cretres.2023.105593 |issn=0195-6671|doi-access=free |bibcode=2023CrRes.15005593S }}

|4 indeterminate remains.

|Pliosaur remains, some with skull sizes comparable to Megacephalosaurus.

|

Taniwhasaurus

|T. sp.

|Kashima Formation.{{Cite journal |last1=Matsui |first1=Kumiko |last2=Karasawa |first2=Tomoki |date=2020-10-18 |title=3D models related to the publication: Interacting with the inaccessible: utilization of multimedia-based visual contents of Japan's National Monument, the Taniwhasaurus mikasaensis (Mosasauridae) holotype for educational workshops at Mikasa City Museum |url=https://www.researchgate.net/publication/345340544 |journal=MorphoMuseuM |volume=6 |issue=5 |pages=e106 |doi=10.18563/journal.m3.106 |s2cid=241798505 |issn=2274-0422}}

|A fragmentary skull, not diagnostic to the species level.

|A tylosaurine mosasaur also known from New Zealand, Antarctica, and South Africa; originally informally named 'Yezosaurus mikasaensis'.

|File:Taniwhasaurus_mikasaensis.png

Pterodactyloidea

|Indeterminate

|

|Fragmentary femur, phalanx, and cervical vertebra

|Large sized form with an estimated wingspan of {{Convert|6.8|m|ft}}.{{Cite journal |last1=Kellner |first1=Alexander W.A. |last2=R. Costa |first2=Fabiana |last3=Wang |first3=Xiaolin |last4=Cheng |first4=Xin |date=2016-02-17 |title=Redescription of the first pterosaur remains from Japan: the largest flying reptile from Asia |url=http://www.tandfonline.com/doi/full/10.1080/08912963.2015.1028929 |journal=Historical Biology |language=en |volume=28 |issue=1–2 |pages=304–309 |doi=10.1080/08912963.2015.1028929 |bibcode=2016HBio...28..304K |issn=0891-2963|url-access=subscription }}

|

=Fish=

class="wikitable" align="center"
colspan="6" align="center" | Fish reported from the Yezo Group
GenusSpeciesPresence

!Material

NotesImages
Apsopelix

|A. miyazakii

| Nakagawa Town{{Cite journal |last1=Yabumoto |first1=Yoshitaka |last2=Hikida |first2=Yoshinori |last3=Nishino |first3=Takanobu |date=April 2012 |title=Apsopelix miyazakii, a New Species of Crossognathid Fish (Teleostei) from the Upper Cretaceous of Hokkaido, Japan |url=https://bioone.org/journals/paleontological-research/volume-16/issue-1/1342-8144-16.1.037/Apsopelix-miyazakii-a-New-Species-of-Crossognathid-Fish-Teleostei-from/10.2517/1342-8144-16.1.037.full |journal=Paleontological Research |volume=16 |issue=1 |pages=37–46 |doi=10.2517/1342-8144-16.1.037 |s2cid=129122259 |issn=1342-8144|url-access=subscription }} and Mikasa area,{{Cite journal |last1=Miyata |first1=Shinya |last2=Yabumoto |first2=Yoshitaka |last3=Nakajima |first3=Yasuhisa |last4=Ito |first4=Yasuhiro |last5=Sasaki |first5=Takenori |date=April 2022 |title=A Second Specimen of the Crossognathiform Fish Apsopelix miyazakii from the Cretaceous Yezo Group of Mikasa Area, Central Hokkaido, Japan |url=https://bioone.org/journals/paleontological-research/volume-26/issue-2/PR200024/A-Second-Specimen-of-the-Crossognathiform-Fish-Apsopelix-miyazakii-from/10.2517/PR200024.full |journal=Paleontological Research |volume=26 |issue=2 |pages=213–223 |doi=10.2517/PR200024 |s2cid=247958614 |issn=1342-8144|url-access=subscription }} Hokkaido (Turonian).

|2 specimens.

| A crossognathid.

|

=Echinoderms=

class="wikitable" align="center"
colspan="6" align="center" | Echinoderms reported from the Yezo Group
GenusSpeciesPresence

!Material

NotesImages
?Austinocrinus

|?A. sp.

| Unknown locality, possibly in south-central Hokkaido, thought to be Santonian-Campanian.{{Cite journal |last=HUNTER |first=AARON W. |date=2012-12-12 |title=Crinoid diversity in the Upper Cretaceous Yezo Supergroup, Hokkaido, northern Japan |url=https://www.researchgate.net/publication/286342199 |journal=Zoosymposia |volume=7 |issue=1 |pages=71–80 |doi=10.11646/zoosymposia.7.1.7 |hdl=20.500.11937/60598 |issn=1178-9913}}

|"One nodal and two pluricolumnals, associated on a bedding plane, so probably from a single individual".

| A crinoid.

|

?Balanocrinus

|?B. sp.

|Upper Coniacian-Campanian portions of the Haborogawa Formation).

|One pluricolumnal.

|A crinoid.

|

Isocrinus

|I. sp.

|Haborogawa Formation (upper Turonian-Coniacian).

|One pluricolumnal.

|A crinoid.

|

Isselicrinus

|I. sp.

|Campanian upper Yezo Group.

|One pluricolumnal and five very poorly preserved columnals.

|

|

=Molluscs=

class="wikitable" align="center"
colspan="6" align="center" | Molluscs reported from the Yezo Group
GenusSpeciesPresence

!Material

NotesImages
?Abyssochrysoidea

|

|Osoushinai Formation.{{Cite journal |last1=Jenkins |first1=Robert |last2=Kaim |first2=Andrzej |last3=Sato |first3=Kei |last4=Moriya |first4=Kazuhiro |last5=Hikida |first5=Yoshinori |last6=Hirayama |first6=Ren |date=2017 |title=Discovery of chemosynthesis-based association on the Cretaceous basal leatherback sea turtle from Japan |url=https://www.researchgate.net/publication/319472309 |journal=Acta Palaeontologica Polonica |volume=62 |doi=10.4202/app.00405.2017 |s2cid=76651553 |issn=0567-7920|doi-access=free }}

|A single specimen found associated with the remains of a Mesodermochelys sp.

|A gastropod.

|

Bivalvia

|Unidentified.

|Osoushinai Formation.

|A single specimen found associated with the remains of a Mesodermochelys sp.

|

|

?Cephalaspidea

|

|Osoushinai Formation.

|A single specimen found associated with the remains of a Mesodermochelys sp.

|A gastropod.

|

Cheloniceras

|C. sp.

|Tsukenai Formation (Aptian).{{Cite journal |last=Futakami |first=Masao |date=2018-09-01 |title=Aptian ammonite fauna from the Yezo Group of the Urakawa area in Hokkaido, Japan |url=https://www.sciencedirect.com/science/article/pii/S0195667117305141 |journal=Cretaceous Research |language=en |volume=89 |pages=224–234 |doi=10.1016/j.cretres.2018.03.025 |bibcode=2018CrRes..89..224F |s2cid=133625463 |issn=0195-6671|url-access=subscription }}

|

|An ammonite.

|

Colombiceras

|C. spathi

| Tsukenai Formation (Aptian).

|

| An ammonite.

|

Gaudryceras

|G. denseplicatum

|Satonosawa Creek, Hokkaido (middle Turonian).{{Cite journal |last1=Tanabe |first1=Kazushige |last2=Hikida |first2=Yoshinori |date=June 2010 |title=Jaws of a New Species ofNanaimoteuthis(Coleoidea: Vampyromorphida) from the Tuironian of Hokkaido, Japan |url=https://www.researchgate.net/publication/232690105 |journal=Paleontological Research |volume=14 |issue=2 |pages=145–150 |doi=10.2517/1342-8144-14.2.145 |s2cid=128429338 |issn=1342-8144}}

|

|An ammonoid.

|

?Gyrodes

|?G. sp.

|Osoushinai Formation.

|A single specimen found associated with the remains of a Mesodermochelys sp.

|

|

Haboroteuthis

|H. poseidon

|Upper Haborogawa Formation (Upper Santonian).{{Cite web |title=Late Cretaceous record of large soft-bodied coleoids based on lower jaw remains from Hokkaido, Japan - Acta Palaeontologica Polonica |url=https://www.app.pan.pl/article/item/app000572013.html |access-date=2022-08-18 |website=www.app.pan.pl |language=en}}

|"KMNH IvP 902,002, a lower jaw."

|A teuthid.

|

Hikidea

|H. cf. yasukawensis

|Osoushinai Formation.

|A single specimen found associated with the remains of a Mesodermochelys sp.

|

|

rowspan="3" |Inoceramus

|I. (Sphenoceramus) orientalis

|Uppermost Haborogawa Formation (middle-upper part of the lower Campanian).{{Cite journal |last=Tsujino |first=Yasuyuki |date=2009 |title=The Upper Cretaceous Hakobuchi Formation of the Yezo Group in the Kotanbetsu area, Hokkaido, northeast Japan |url=https://www.jstage.jst.go.jp/article/geosoc/115/3/115_3_122/_article/-char/en |journal=The Journal of the Geological Society of Japan |volume=115 |issue=3 |pages=122–129 |doi=10.5575/geosoc.115.122|s2cid=130346354 |doi-access=free }}

|

|A bivalve.

|

I. (Sphenoceramus) schmidti

|Uppermost Haborogawa Formation (middle-upper part of the lower Campanian).

|

|A bivalve.

|

I. teshioensis

|Gakkonosawa Creek, Hokkaido (late Turonian).

|Disarticulated valves.

|A bivalve.

|

rowspan=3|Nanaimoteuthis

|N. hikidai

|Uppermost Haborogawa Formation (lower Campanian).

|"KMNH IvP 902,001, a lower jaw, the only specimen available."

|A vampyromorph.

|

N. jeletzkyi

|Hokkaido (Santonian to lower Campanian.){{Cite journal |last1=Tanabe |first1=Kazushige |last2=Misaki |first2=Akihiro |last3=Hikida |first3=Yoshinori |last4=Nishimura |first4=Tomohiro |date=February 2017 |title=New records of coleoid cephalopod jaws from the Upper Cretaceous of Hokkaido, Japan, and their paleobiogeographic and evolutionary implications |url=http://dx.doi.org/10.1016/j.cretres.2016.10.009 |journal=Cretaceous Research |volume=70 |pages=128–141 |doi=10.1016/j.cretres.2016.10.009 |bibcode=2017CrRes..70..128T |issn=0195-6671|url-access=subscription }}

|

|A vampyromorph.

|

N. yokotai

|Satonosawa Creek and Gakkonosawa Creek, Hokkaido (middle and late Turonian).

|Jaw fossils.

|A vampyromorph.

|

Paleocirroteuthis

|P. sp.

|Hokkaido (Santonian to lower Campanian).

|

|A cirroctopodid.

|

Provanna

|P. cf. nakagawensis

|Osoushinai Formation.

|3 specimens found associated with the remains of a Mesodermochelys sp.

|An abyssochrysoid gastropod.

|

Rhytidoplites

|R. adkinsi

|Tsukenai Formation (Aptian).

|

|An ammonite.

|

rowspan=2|Scalarites

|S. mihoensis

|Gakkonosawa Creek, Hokkaido (late Turonian).

|

|An ammonoid.

|

S. scalaris

|Satonosawa Creek, Hokkaido (middle Turonian).

|

|An ammonoid.

|

Tetragonites

|T. glabrus

|Satonosawa Creek, Hokkaido (middle Turonian).

|

|An ammonoid.

|

Thyasira

|T. tanabei

|Osoushinai Formation.

|2 articulated specimens found associated with the remains of a Mesodermochelys sp.

|A bivalve.

|

Yezoteuthis

|Y. giganteus

|Osoushinai Formation.{{Cite journal |last1=Tanabe |first1=Kazushige |last2=Hikida |first2=Yoshinori |last3=Iba |first3=Yasuhiro |date=2006 |title=Two coleoid jaws from the Upper Cretaceous of Hokkaido, Japan |url=https://www.cambridge.org/core/journals/journal-of-paleontology/article/abs/two-coleoid-jaws-from-the-upper-cretaceous-of-hokkaido-japan/E9AB99CDD77DBEF1736FF42771EB8E6B |journal=Journal of Paleontology |language=en |volume=80 |issue=1 |pages=138–145 |doi=10.1666/0022-3360(2006)080[0138:TCJFTU]2.0.CO;2 |issn=0022-3360|url-access=subscription }}

|Jaw fossils.

|A very large oegopsid.

|

=Paleoflora=

class="wikitable" align="center"
colspan="6" align="center" | Plants reported from the Yezo Group
GenusSpeciesPresence

!Material

NotesImages
Eydeia

|E. hokkaidoensis

|Haborogawa Formation (Coniacian-Santonian).{{Cite journal |last1=Stockey |first1=Ruth A. |last2=Nishida |first2=Harufumi |last3=Atkinson |first3=Brian A. |date=September 2016 |title=Anatomically preserved fossil cornalean fruits from the Upper Cretaceous of Hokkaido: Eydeia hokkaidoensis gen. et sp. nov. |journal=American Journal of Botany |language=en |volume=103 |issue=9 |pages=1642–1656 |doi=10.3732/ajb.1600151 |issn=0002-9122|doi-access=free |pmid=27589935 }}

|5 perimineralized fruits.

|A cornale similar to the extant Davidia involucrata.

|

Mikasapteris

|M. rothwellii

| Mikasa Locality.{{Cite journal |last1=Nishida |first1=Harufumi |last2=Stockey |first2=Ruth A. |last3=Takebe |first3=Yusuke |last4=Legrand |first4=Julien |last5=Yamada |first5=Toshihiro |date=2022-07-25 |title=Mikasapteris rothwellii gen. et sp. nov., a Permineralized Fertile Pinnule of a Probable Stem Polypod from the Late Cretaceous of Hokkaido, Japan |url=https://www.journals.uchicago.edu/doi/abs/10.1086/721262 |journal=International Journal of Plant Sciences |volume=183 |issue=7 |pages=576–586 |doi=10.1086/721262 |s2cid=251086117 |issn=1058-5893|url-access=subscription }}

|A permineralized fertile pinnule.

| A probable stem polypodioid leptosporangiate fern.

|

Mukawastrobus

|M. satoi

|Hakobuchi Formation.{{Cite journal |last1=Stockey |first1=Ruth A. |last2=Nishida |first2=Harufumi |last3=Rothwell |first3=Gar W. |date=2020 |title=Evolutionary diversification of taiwanioid conifers: evidence from a new Upper Cretaceous seed cone from Hokkaido, Japan |journal=Journal of Plant Research |volume=133 |issue=5 |pages=681–692|doi=10.1007/s10265-020-01214-y |pmid=32686035 |pmc=7429551 |bibcode=2020JPlR..133..681S }}

|"A single cylindrical seed cone 2 cm long, 1.1 cm wide."

|A cypress.

|

Nilssonia

|N. yezoensis

|Hakobuchi Formation.{{Cite book |first=Atsushi |last=Okubo |url=http://worldcat.org/oclc/249459457 |title=Nilssonia yezoensis, sp. nov., from the Upper Cretaceous Hakobuchi group, in Hokkaido, Japan |oclc=249459457}}

|

|A cycad.

|

Obirafructus

|O. kokubunii

|Haborogawa Formation (Coniacian to Santonian).{{Cite journal |last1=Kajita |first1=Yui |last2=Suzuki |first2=Mayumi Hanari |last3=Nishida |first3=Harufumi |date=May 2020 |title=A Multicarpellary Apocarpous Gynoecium from the Late Cretaceous (Coniacian–Santonian) of the Upper Yezo Group of Obira, Hokkaido, Japan: Obirafructus kokubunii gen. & sp. nov. |url=https://www.researchgate.net/publication/353759123 |journal=Acta Phytotaxonomica et Geobotanica |volume=72 |issue=1 |pages=1–21}}

|"A permineralized reproductive axis bearing at least 42 spirally arranged follicles."

|A saxifragale.

|

Stockeystrobus

|S. interdigitata

|Hoborogawa Formation (Coniacian to Santonian).{{Cite journal |last1=Rothwell |first1=Gar W. |last2=Ohana |first2=Tamiko |date=September 2016 |title=Stockeystrobusgen. nov. (Cupressaceae), and the evolutionary diversification of sequoioid conifer seed cones |url=https://www.researchgate.net/publication/308181978 |journal=Botany |volume=94 |issue=9 |pages=847–861 |doi=10.1139/cjb-2016-0025 |issn=1916-2790|doi-access=free }}

|A seed cone.

|A sequoioid.

|

Zamites

|Z. bayeri

|Kashima Formation (Coniacian).{{Cite journal |last=Yamada |first=Toshihiro |date=2009-10-01 |title=Structurally preserved Zamites bayeri Kvaček from the Coniacian Kashima Formation (Yezo Group) of Hokkaido, Japan |url=https://www.sciencedirect.com/science/article/pii/S0195667109000810 |journal=Cretaceous Research |language=en |volume=30 |issue=5 |pages=1301–1306 |doi=10.1016/j.cretres.2009.07.007 |bibcode=2009CrRes..30.1301Y |issn=0195-6671|url-access=subscription }}

|

|A bennettitale.

|

See also

References