australidelphia
{{Short description|Superorder of marsupials}}
{{Automatic taxobox
| fossil_range = {{fossilrange|61.6|0|earliest=Paleocene}} Early Paleocene to present{{Cite web|title=PBDB|url=https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=99550&is_real_user=1|access-date=2021-07-13|website=paleobiodb.org}}
| image = Sumpfwallabycele4.jpg
| image_caption = A swamp wallaby
| taxon = Australidelphia
| authority = Szalay 1982
| subdivision_ranks = Orders
| subdivision = * Microbiotheria
- †Polydolopimorphia?
- Eomarsupialia
{{Small|Archer, 1984}}{{cite journal |last1=Beck |first1=R. M. D. |last2=Travouillon |first2=K. J. |last3=Aplin |first3=K. P. |last4=Godthelp |first4=H. |last5=Archer |first5=M. |title=The Osteology and Systematics of the Enigmatic Australian Oligo-Miocene Metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia; ?Australidelphia; Marsupialia) |journal=Journal of Mammalian Evolution |date=2014 |volume=21 |issue=2 |pages=127–172 |doi=10.1007/s10914-013-9236-3|s2cid=18490996 |url=https://usir.salford.ac.uk/id/eprint/33350/1/Beck_et_al_%282014%29_Yalkaparidon.pdf }} - Diprotodontia
- Agreodontia
{{Small|Beck et al., 2014}} - Dasyuromorphia
- Notoryctemorphia
- Peramelemorphia
}}
Australidelphia is a superorder of marsupials encompassing about three-quarters of all living marsupial species, including all those native to Australasia and one South American species, the monito del monte. Unlike other American marsupials, which belong to the Ameridelphia, Australidelphia’s lineage emerged in South America, with genetic evidence (retrotransposon insertion sites) showing the monito del monte as its most ancient branch.{{Cite web |last=Schiewe |first=Jessie |date=2010-07-28 |title=Australia's marsupials originated in what is now South America, study says |url=http://www.latimes.com/news/science/la-sci-marsupial-20100728,0,5549873.story |url-status=live |archive-url=https://web.archive.org/web/20100801001045/http://www.latimes.com/news/science/la-sci-marsupial-20100728%2C0%2C5549873.story |archive-date=1 August 2010 |access-date=2010-08-01 |work=Los Angeles Times}}{{Cite journal |last1=Nilsson |first1=M. A. |last2=Churakov |first2=G. |last3=Sommer |first3=M. |last4=Van Tran |first4=N. |last5=Zemann |first5=A. |last6=Brosius |first6=J. |last7=Schmitz |first7=J. |date=2010-07-27 |editor1-last=Penny |editor1-first=David |title=Tracking Marsupial Evolution Using Archaic Genomic Retroposon Insertions |journal=PLOS Biology |volume=8 |issue=7 |pages=e1000436 |doi=10.1371/journal.pbio.1000436 |pmc=2910653 |pmid=20668664 |doi-access=free}}
Within this superorder, the Australian members form a distinct group (clade) named Eomarsupialia, though their internal relationships (branching order) is yet to be determined. Studies suggest Australidelphia originated in South America alongside other major marsupial groups, likely dispersing to Australia via Antarctica in a single event after the monito’s lineage (Microbiotheria) diverged, leaving other South American orders (Didelphimorphia and Paucituberculata) as more basal.
Phylogeny
The following cladogram is a phylogeny of Australidelphia based on the work of May-Collado, Kilpatrick & Agnarsson 2015,{{cite journal |author=May-Collado |year=2015 |title=Mammals from 'down under': a multi-gene species-level phylogeny of marsupial mammals (Mammalia, Metatheria). |journal=PeerJ |volume=3 |issue=e805 |pages=e805 |doi=10.7717/peerj.805 |pmid=25755933 |pmc=4349131|display-authors=etal |doi-access=free }} with extinct clades from Black et al. 2012.{{cite book |author=Black |title=Earth and Life |year=2012 |chapter=The Rise of Australian Marsupials: A Synopsis of Biostratigraphic, Phylogenetic, Palaeoecologic and Palaeobiogeographic Understanding |publisher=Springer Netherlands |isbn=9789048134274 |pages=983–1078 |doi=10.1007/978-90-481-3428-1_35|display-authors=etal}}
{{clade| style=font-size:90%;line-height:80%
|label1=Australidelphia
|1={{Clade
|label1=Microbiotheria
|1={{Clade
}}
|label2=Eomarsupialia
|2={{Clade
|label1=Agreodontia
|1={{Clade
|label1=Notoryctemorphia
|1=Notoryctidae
|2={{Clade
|label1=Peramelemorphia
|1={{Clade
|label1=†Yaraloidea
|1=†Yaralidae
|label2=Perameloidea
|2={{Clade
|2={{Clade
|1=Peramelidae
|2={{Clade
|1=†Chaeropodidae
|2=Peroryctidae
}}
}}
}}
}}
|label2=Dasyuromorphia
|2={{Clade
|1=†Thylacinidae
|2={{Clade
|2=Dasyuridae
}}
}}
}}
}}
|label2=Diprotodontia
|2={{Clade
|label1=Vombatiformes
|1={{Clade
|label1=Phascolarctimorphia
|label2=Vombatimorphia
|2={{Clade
|1=†Thylacoleonidae
|2={{Clade
|1=?†Wynyardiidae
|2=†Ilariidae
|3={{Clade
|1=†Maradidae
|2={{Clade
|1=Vombatidae
|2={{Clade
|1=†Palorchestidae
|2=†Diprotodontidae
}}
}}
}}
}}
}}
}}
|label2=Phalangerida
|2={{Clade
|label1=Phalangeriformes
|1={{Clade
|label1=Burramyoidea
|1=Burramyidae
|label2=Phalangeroidea
|2={{Clade
|2={{Clade
|1=?†Pilkipildridae
|2={{Clade
|1=†Ektopodontidae
|2=†Miralinidae
}}
}}
}}
}}
|label2=Macropodiformes
|2={{Clade
|label1=Petauroidea
|1={{Clade
|1=Acrobatidae
|2={{Clade
|1=Tarsipedidae
|2={{Clade
|1=Petauridae
}}
}}
}}
|label2=Macropodoidea
|2={{Clade
|1=?†Balbaridae
|2={{Clade
|2={{Clade
|1=Potoroidae
|2=Macropodidae
}}
}}
}}
}}
}}
}}
}}
}}
}}
Taxonomy
The orders within this group are listed below:
- Genus †Djarthia Godthelp, Wroe & Archer 1999
- Order Dasyuromorphia (71 species)
- Family Dasyuridae: antechinuses, quolls, dunnarts, Tasmanian devil, and allies
- Family Myrmecobiidae: numbat
- Family †Thylacinidae: thylacine
- Order Diprotodontia (117 species)
- Family Acrobatidae: feathertail glider and feather-tailed possum
- Family Burramyidae: pygmy possums
- Family Hypsiprymnodontidae: musky rat-kangaroo
- Family Macropodidae: kangaroos, wallabies, and allies
- Family Petauridae: striped possum, Leadbeater's possum, yellow-bellied glider, sugar glider, mahogany glider, squirrel glider
- Family Phalangeridae: brushtail possums and cuscuses
- Family Phascolarctidae: koala
- Family Potoridae: potoroos, rat kangaroos, bettongs
- Family Pseudocheiridae: ringtailed possums and allies
- Family Tarsipedidae: honey possum
- Family Vombatidae: wombats
- Family †Diprotodontidae: giant wombats
- Family †Palorchestidae: marsupial tapirs
- Family †Thylacoleonidae: marsupial lions
- Order Microbiotheria (1 species)
- Family Microbiotheriidae: monito del monte
- Order Notoryctemorphia (2 species)
- Family Notoryctidae: marsupial moles
- Order Peramelemorphia (21 species)
- Family Peramelidae: bandicoots and allies
- Family Thylacomyidae: bilbies
- Family †Chaeropodidae: pig-footed bandicoots
- Order †Yalkaparidontia Archer, Hand & Godthelp 1988
- Family †Yalkaparidontidae Archer, Hand & Godthelp 1988
References
{{Reflist}}
{{Metatheria|M.|state=autocollapse}}
{{Mammals}}
{{Agreodontia}}
{{Portal bar|Australia|Mammals}}
{{Taxonbar|from=Q631975}}
Category:Fauna of the Australasian realm
Category:Extant Paleocene first appearances
Category:Marsupials of Australia