shoot (botany)
{{short description|Young stem or branch}}
In botany, a plant shoot consists of any plant stem together with its appendages like leaves, lateral buds, flowering stems, and flower buds.{{cite book|last=Esau|first=K.|title=Plant Anatomy|date=1953|publisher=John Wiley & Sons Inc.|location=New York|page=411}}{{cite book|last=Cutter|first=E.G.|year=1971|title=Plant Anatomy, experiment and interpretation, Part 2 Organs|publisher=Edward Arnold|location=London|isbn=0-7131-2302-8|page=117}} The new growth from seed germination that grows upward is a shoot where leaves will develop. In the spring, perennial plant shoots are the new growth that grows from the ground in herbaceous plants or the new stem or flower growth that grows on woody plants.
In everyday speech, shoots are often synonymous with stems. Stems, which are an integral component of shoots, provide an axis for buds, fruits, and leaves.
Young shoots are often eaten by animals because the fibers in the new growth have not yet completed secondary cell wall development, making the young shoots softer and easier to chew and digest.
As shoots grow and age, the cells develop secondary cell walls that have a hard and tough structure.
Some plants (e.g. bracken) produce toxins that make their shoots inedible or less palatable.
File:Cucumber leaf.jpg|The shoot of a cucumber
File:GiantKnotweed048.jpg|Edible shoots of Sachaline
File:Sunflower seedlings.jpg|Sunflower seedlings germinate
File:Persea americana (Avocado) Sprout 08May2010.JPG|A young hass avocado shoot
Shoot types of woody plants
Many woody plants have distinct short shoots and long shoots. In some angiosperms, the short shoots, also called spur shoots or fruit spurs, produce the majority of flowers and fruit. A similar pattern occurs in some conifers and in Ginkgo, although the "short shoots" of some genera such as Picea are so small that they can be mistaken for part of the leaf that they have produced.{{citation |author1=Gifford, E.M. |author2=Foster, A.S. |year=1989 |title=Morphology, and evolution of vascular plants |publisher=W. H. Freeman and Company |location=New York}}
{{anchor|Heterophylly|Seasonal heterophylly|Anisophylly}}A related phenomenon is seasonal heterophylly, which involves visibly different leaves from spring growth and later lammas growth.{{citation |author=Eckenwalder, J.E. |year=1980 |title=Foliar Heteromorphism in Populus (Salicaceae), a Source of Confusion in the Taxonomy of Tertiary Leaf Remains |journal=Systematic Botany |volume=5 |issue=4 |pages=366–383 |jstor=2418518 |doi=10.2307/2418518}} Whereas spring growth mostly comes from buds formed the previous season, and often includes flowers, lammas growth often involves long shoots.{{cn|date=June 2024}}
File: Suckers on stump.jpg|Long shoot growth
File: Pyrus pyrifolia (Hosui) blossom.jpg|A mature fruiting spur on a Nashi pear tree, Pyrus pyrifolia
File: Cedrus deodara 02.JPG|On long shoots of Cedrus deodara individual leaves may have buds in the axils.
File:20130903Cedrus deodara2.jpg|Cedrus deodara forms short shoots (from buds) along the long shoots.
See also
- Bud
- Crown (botany)
- Heteroblasty, an abrupt change in the growth pattern of some plants as they mature
- Lateral shoot
- Phyllotaxis, the arrangement of leaves along a plant stem
- Seedling
- Sterigma, the "woody peg" below the leaf of some conifers
- Thorn (botany), true thorns, as distinct from spines or prickles, are short shoots