trematosauridae
{{Short description|Extinct family of temnospondyls}}
{{Automatic taxobox
| name = Trematosaurids
| fossil_range = Triassic (Possible Jurassic record), {{Fossil range|251.9|220|latest=Jurassic}}
| image = Trematolestes hagdorni.JPG
| image_caption = Fossil of Trematolestes hagdorni in the State Museum of Natural History Stuttgart
| taxon = Trematosauridae
| authority = Watson, 1919
| subdivision_ranks = Subfamilies and genera
| subdivision =
}}
Trematosauridae is a family of large marine temnospondyls with several included genera.
Appearance and lifestyle
Trematosaurids are one of the most derived families of the Trematosauroidea superfamily in that they are the only family that have fully marine lifestyles. Long, slender snouts that are characteristic of the trematosaurids, with some members having rostra resembling those of modern-day gavials.
Traditionally, two subfamilies within Trematosauridae can be identified, the relatively short-nosed Trematosaurinae and the long-nosed Lonchorhynchinae.{{cite journal |last=Damani |first=Ross |year=2004 |title=Cranial anatomy and relationships of Microposaurus casei, a temnospondyl from the Middle Triassic of South Africa |journal= Journal of Vertebrate Paleontology|volume=24 |issue=3 |pages=533–541|doi=10.1671/0272-4634(2004)024[0533:CAAROM]2.0.CO;2 |issn=0272-4634 |s2cid=131215804 }} A third subfamily, Tertreminae, was named in 2000 and includes broad-snouted forms like Tirraturhinus.
Fossil record
Trematosaurids first appeared during the Induan{{cite journal |last1=Scheyer |first1=Torsten M. |last2=Romano |first2=Carlo |last3=Jenks |first3=Jim |last4=Bucher |first4=Hugo |date=19 March 2014 |title=Early Triassic Marine Biotic Recovery: The Predators' Perspective |journal=PLOS ONE |volume=9 |issue=3 |pages=e88987 |bibcode=2014PLoSO...988987S |doi=10.1371/journal.pone.0088987 |pmc=3960099 |pmid=24647136 |doi-access=free}} age (Wordie Creek Formation, Greenland) of the Early Triassic epoch. The family existed until around the Carnian age of the Late Triassic epoch,{{cite journal |last1=Schoch |first1=Rainer R. |last2=Milner |first2=Andrew R. |last3=Hellrung |first3=Hannah |year=2002 |title=The last trematosaurid amphibian Hyperokynodon keuperinus revisited |journal=Stuttgarter Beiträge zur Naturkunde, Serie B (Geologie und Paläontologie) |volume=321 |pages=1–9 |issn=0341-0153|url=http://www-alt.naturkundemuseum-bw.de/stuttgart/pdf/b_pdf/B321.pdf}} although by then they were very rare. By the Middle Triassic they had become widespread throughout Laurasia and Gondwana with fossils being found in Europe, Asia, Madagascar, and Australia. A possible trematosaurid has been found in the Toutunhe Formation in the Junggar Basin. If this analysis is accurate, it renders Trematosauridae one of the longest lived lineages of the Temnospondyli, having lasted as recently as the late Jurassic.(Maisch et al. 2004, p. 582)
In 2006, a new Middle Triassic genus Trematolestes from southern Germany has been reported. It was the sister taxon of the subfamily Lonchorhynchinae and its closest relative was Tertremoides.{{Cite journal |author=Rainer R. Schoch |year=2006 |title=A Complete Trematosaurid Amphibian From The Middle Triassic Of Germany|journal=Journal of Vertebrate Paleontology |volume=26 |issue=1 |pages=29–43 |doi=10.1671/0272-4634(2006)26[29:ACTAFT]2.0.CO;2 |s2cid=85829091 |url=http://www.bioone.org/doi/abs/10.1671/0272-4634(2006)26%5B29%3AACTAFT%5D2.0.CO%3B2|url-access=subscription }}
Phylogeny
Below is a cladogram from Steyer (2002) showing the phylogenetic relationships of trematosaurids:{{cite journal |last=Steyer |first=J. S. |year=2002 |title=The first articulated trematosaur 'amphibian' from the Lower Triassic of Madagascar: implications for the phylogeny of the group |journal=Palaeontology |volume=45 |issue=4 |pages=771–793 |doi=10.1111/1475-4983.00260|bibcode=2002Palgy..45..771S |s2cid=83515233 }}
{{clade| style=font-size:100%;line-height:85%
|label1=Trematosauridae
|1={{clade
|label1=Trematosaurinae
|1={{clade
|1={{clade
|1=Tertrema
|2={{clade
|2=Platystega}} }}
|2={{clade
|1=Luzocephalus
|2={{clade
|2=Trematosuchus}} }} }}
|label2=Lonchorhynchinae
|2={{clade
|1=Aphaneramma
|2={{clade
|2=Cosgriffius
|3={{clade
|2=Wantzosaurus}} }} }} }} }}
A cladogram after Novikov (2018) with only Early Triassic Eastern Europe taxa included:{{cite book|author=Novikov A.V.|year=2018|url=https://new.ras.ru/upload/iblock/7de/j85vtubof3mlf81izrqq9u9oh9eb5lrh.pdf|title=Early Triassic amphibians of Eastern Europe: evolution of dominant groups and peculiarities of changing communities|publisher=Moscow: RAS|page=138|lang=ru|isbn=978-5-906906-71-7}} {{cite web|url=https://new.ras.ru/upload/iblock/7de/j85vtubof3mlf81izrqq9u9oh9eb5lrh.pdf |title=Archive copy |date=December 8, 2023 |archive-url=https://web.archive.org/web/20231208092959/https://new.ras.ru/upload/iblock/7de/j85vtubof3mlf81izrqq9u9oh9eb5lrh.pdf |archive-date=2023-12-08 }}
{{clade
|label1=Trematosauridae
|1={{clade
|2={{clade
|1=Thoosuchus
|2={{clade
|2={{clade
|1=Angusaurus
|2={{clade
|2=Trematosaurus }} }} }} }} }} }}
References
{{Reflist}}
External links
- [https://web.archive.org/web/20090328190158/http://www.palaeos.com/Vertebrates/Units/160Temnospondyli/160.450.html Trematosauridae] at Palaeos
{{Stereospondyli|T.}}
{{Taxonbar|from=Q2342478}}
Category:Temnospondyl families
Category:Early Triassic first appearances
{{Temnospondyli-stub}}