:Candeleros Formation

{{Short description|Geologic formation in Argentina}}

{{Infobox rockunit

| name = Candeleros Formation

| image = Fm Candeleros cerro Vagon.jpg

| caption = Candeleros Formation near Cerro El Vagon, Neuquen, Argentina

| type = Geological formation

| period = Cenomanian

| age = early Cenomanian
~{{fossil range|99|97}}

| prilithology = Eolian sandstone

| otherlithology = Conglomerate, siltstone, paleosol

| namedfor = Candeleros Hill

| namedby = Wichmann

| year_ts = 1929

| region = Mendoza, Neuquén & Río Negro Provinces

| country = Argentina

| coordinates = {{coord|39.4|S|69.2|W|display=inline,title}}

| paleocoordinates = {{coord|46.5|S|45.5|W|display=inline}}

| unitof = Neuquén Group
 Río Limay Subgroup

| subunits =

| underlies = Huincul Formation

| overlies = Lohan Cura Formation

| thickness = {{convert|300|m|ft|abbr=on}}

| extent = Neuquén Basin

| area =

| map = {{Location map+ | Argentina

| relief = 1

| width = 250

| float = center

| places =

{{Location map~ | Argentina

| lat_deg = -39.4

| lon_deg = -69.2

| mark = Lightgreen pog.svg

| marksize = 12

}}

}}

| map_caption =

}}

The Candeleros Formation is a geologic formation that crops out in the Río Negro, Neuquén, and Mendoza provinces of northern Patagonia, Argentina. It is the oldest formation in the Neuquén Group and belongs to the Rio Limay Subgroup. Formerly that subgroup was treated as a formation, and the Candeleros Formation was known as the Candeleros Member.Sánchez et al., 2006

Description

The type locality of the Candeleros Formation is Candeleros Hill in Neuquén Province, after which the formation was named by Wichmann in 1929.Wichmann, 1929 This formation unconformably overlies the Lohan Cura Formation, and it is fused with the Huincul Formation, also a unit of the Neuquén Group. The sediments of the latter are of lighter greenish and yellow colors and the fused boundary between the Candeleros and Huincul formations is easily recognizable.Leanza et al., 2004

The Candeleros Formation is almost {{convert|300|m|ft}} thick in some sections. Overall, the formation represents a part of the ancient Kokorkom desert with braided river system, made up mostly of sandstones and conglomerates. There are also isolated sections that represent eolian (wind-blown) deposition, as well as siltstones deposited under swamp conditions. Paleosols (soil deposits) are common in some sections as well.

Fossil content

File:Hunting season in Argentina.jpg of Candeleros Formation]]

The Candeleros Formation has a very diverse fossil fauna, including fish, frogs, mammals, rhynchocephalians, snakes, turtles, notosuchians, pterosaurs, and abundant dinosaurs including rebbachisaurid and titanosaurian sauropods and diverse theropods, and the enigmatic armored Jakapil.

Ichnofossils found in the formation include those assigned to Sousaichnum monettae, Limayichnus major, Bressaniichnus patagonicus, Deferraiichnum mapuchense, and Picunichnus benedettoi.Leonardi, 1994, p. 79

{{paleobiota-key-compact}}

= Fish =

class="wikitable sortable" align="center" width="75%"
TaxaSpeciesLocalityMemberMaterialNotesImages
Ceratodus

| C. argentinus

|

|Upper

|

| A lungfish

|

= Amphibians =

class="wikitable sortable" align="center" width="75%"
TaxaSpeciesLocalityMemberMaterialNotesImages
AvitabatrachusBaez, Ana & Trueb, Linda & Calvo, Jorge. (2000). The earliest known pipoid frog from South America: A new genus from the Middle Cretaceous of Argentina. Journal of Vertebrate Paleontology. 20. 490-500. 10.1671/0272-4634(2000)020%5B0490:TEKPFF%5D2.0.CO;2.

| A. uliana

| El Gigante

|Lower

| Fragmentary skull and postcranial skeleton

| A pipimorph frog

|

= Dinosaurs =

== Ornithischians ==

class="wikitable sortable" align="center" width="75%"
TaxaSpeciesLocalityMemberMaterialNotesImages
style="background:#FEF6E4;"|Bonaparteichnium''{{cite journal | url=https://www.researchgate.net/publication/283994360 | title=Huellas de dinosaurios en la Formación Río Limay (Albiano-Cenomaniano?), Picún Leufú, Provincia del Neuquén, República Argentina (Ornithischia-Saurischia: Sauropoda-Theropoda) | author=Calvo, Jorge O. | journal=Ameghiniana | year=1991 | volume=28 | issue=3 | pages=241–258}}

|style="background:#FEF6E4;"| B. tali''

|style="background:#FEF6E4;"''|

|style="background:#FEF6E4;"''|

|style="background:#FEF6E4;"''| Footprints

|style="background:#FEF6E4;"''| Iguanodont ichnofossils

|style="background:#FEF6E4;"|

Jakapil''{{cite journal |vauthors=Riguetti FJ, Apesteguía S, Pereda-Suberbiola X |year=2022 |title=A new Cretaceous thyreophoran from Patagonia supports a South American lineage of armoured dinosaurs |journal=Scientific Reports |volume=12 |issue=1 |pages=Article number 11621 |doi=10.1038/s41598-022-15535-6 |pmid=35953515 |pmc=9372066 |doi-access=free|bibcode=2022NatSR..1211621R }}

| J. kaniukura

| Cerro Policía

| Upper

| A partial skeleton including several osteoderms and a complete lower jaw

| Possibly a basal thyreophoran{{cite journal |last1=Raven |first1=T. J. |last2=Barrett |first2=P. M. |last3=Joyce |first3=C. B. |last4=Maidment |first4=S. C. R. |year=2023 |title=The phylogenetic relationships and evolutionary history of the armoured dinosaurs (Ornithischia: Thyreophora) |journal=Journal of Systematic Palaeontology |volume=21 |issue=1 |at=2205433 |doi=10.1080/14772019.2023.2205433 |bibcode=2023JSPal..2105433R |url=https://cris.brighton.ac.uk/ws/files/40730447/Raven_et_al_Manuscript_PostReview_Final.pdf }}

| File:Jakapil UDL.png

== Saurischians ==

; Sauropods

class="wikitable sortable" align="center" width="75%"
TaxaSpeciesLocalityMemberMaterialNotesImages
Andesaurus

| A. delgadoi

|Near Villa El Chocon

|Lower

| A partial skeleton

| A basal titanosaur

| File:Andesaurus_LM.png

Campananeyen{{Cite journal |last1=Lerzo |first1=Lucas N. |last2=Fernández-Baldor |first2=Fidel Torcida |last3=Canale |first3=Juan I. |last4=Whitlock |first4=John A. |last5=Otero |first5=Alejandro |last6=Gallina |first6=Pablo A. |date=2024-08-13 |title=They all floated in the Cretaceous: new rebbachisaurid (Sauropoda, Diplodocoidea) with a highly pneumatized skeleton from the Upper Cretaceous (lower Cenomanian) of Patagonia, Argentina |journal=Historical Biology |language=en |pages=1–14 |doi=10.1080/08912963.2024.2383708 |issn=0891-2963}}

| C. fragilissimus

| Barda Atravesada de Las Campanas

|Lower

| A fragmentary skeleton

| A rebbachisaurid

| File:Campananeyen fragilissimus.png

Limaysaurus

| L. tessonei

|

|Upper

|

| A rebbachisaurid

|

Nopcsaspondylus

| N. alarconensis

|left bank of Limay River

|

|A single lost vertebra

| A rebbachisaurid

|

Rayososaurus

| R. agrioensis

| Agrio del Medio

|Upper

| A left scapula, an almost complete right scapular blade, and the distal three-quarters of a left femur.

| A rebbachisaurid

|

Titanosauria

| Indeterminate

|

|Upper

|

| A giant titanosaur, probably exceeding Patagotitan in size{{Cite journal |last1=Otero |first1=Alejandro |last2=Carballido |first2=José L. |last3=Salgado |first3=Leonardo |last4=Canudo |first4=José Ignacio |last5=Garrido |first5=Alberto C. |date=June 2021 |title=Report of a giant titanosaur sauropod from the Upper Cretaceous of Neuquén Province, Argentina |journal=Cretaceous Research |language=en |volume=122 |doi=10.1016/j.cretres.2021.104754 |issn=0195-6671}}

|

; Theropods

class="wikitable sortable" align="center" width="75%"
TaxaSpeciesLocalityMemberMaterialNotesImages
style="background:#FEF6E4;"''

|Abelichnus

| A. astigerrae

|

|

| A footprint

|

|

Alnashetri

| A. cerropoliciensis

| La Buitrera

|Upper

| A fragmentary skeleton including leg and foot bones

| An alvarezsauroid

| File:Alnashetri LM.png

Bicentenaria

| B. argentina

| Ezequiel Ramos Mexia Reservoir

|Upper

| Much of a skeleton

| A coelurosaur

| File:Bicentenaria.jpg

Buitreraptor

| B. gonzalezorum

| La Buitrera

|Upper

| Several partial skeletons

| A dromaeosaurid

| File:BuitreraptorROM.JPG

Ekrixinatosaurus

| E. novasi

| Bajo del Añelo

| Upper

| A partial skeleton

| An abelisaurid

| File:Ekrixinatosaurus novasi by Henrique Paes.png

Giganotosaurus

| G. carolinii

| Los Candeleros and Villa El Chocón

| Upper/Lower

| A partial skull and a disarticulated postcranial skeleton

| A carcharodontosaurid

|File:Giganotos Db.jpg

= Pterosaurs =

class="wikitable sortable" align="center" width="75%"
TaxaSpeciesLocalityMemberMaterialNotesImages
Azhdarchidae{{Cite journal |last1=Agnolin |first1=Federico |last2=Rozadilla |first2=Sebastián |last3=Juárez-Valieri |first3=Rubén |last4=Meso |first4=Jorge |date=2023-10-25 |title=Oldest azhdarchid (Pterosauria) record from South America |url=http://revista.macn.gob.ar/ojs/index.php/RevMus/article/view/825 |journal=Revista del Museo Argentino de Ciencias Naturales |series=Nueva serie |language=es |volume=25 |issue=2 |pages=309–314 |issn=1853-0400}}

| Indeterminate

| Ezequiel Ramos Mexía Reservoir

| Upper

| A partial cervical vertebra

| An azhdarchid

|

= Other reptiles =

class="wikitable sortable" align="center" width="75%"
TaxaSpeciesLocalityMemberMaterialNotesImages
rowspan="3" |Araripesuchus

|A. buitreraensis

| La Buitrera, Cerro Policía and El Pueblito.

|Upper

| Multiple skull specimens.

| rowspan="3" | A small notosuchian crocodyliform

| rowspan = "3"| File:Araripesuchus patagonicus SEND.jpg

A manzanensis

|

|Upper

| A partial skull and lower jaw

A. patagonicus

| El Chocón.

|Lower

| An anterior half of the skull with an articulated mandible, lacking the most anterior portion of the rostrum and also preserves part of the postcranium

Najash

| N. rionegrina

|

|Upper

|

| A basal snake

| Image:Boli-ischi 100.jpg

Priosphenodon

| P. avelasi

|

|Upper

|

| An eilenodontine rhynchocephalian

| File:Priosphenodon skeleton.png

Tika

| T. giacchinoi

|

|Upper

|

| A sphenodontine rhynchocephalian

|

= Mammals =

class="wikitable sortable" align="center" width="75%"
TaxaSpeciesLocalityMemberMaterialNotesImages
Cronopio

| C. dentiacutus

| La Buitrera

| Upper

| Skull bones and teeth

| A small meridiolestidan mammal

| File:Cronopio NT.jpg

= Turtles =

class="wikitable sortable" align="center" width="75%"
TaxaSpeciesLocalityMemberMaterialNotesImages
Prochelidella{{Cite journal |last1=Maniel |first1=Ignacio J. |last2=de la Fuente |first2=Marcelo S. |last3=Apesteguía |first3=Sebastián |last4=Pérez Mayoral |first4=Joaquin |last5=Sanchez |first5=Maria L. |last6=Veiga |first6=Gonzalo D. |last7=Smales |first7=Ian |date=2020-02-24 |title=Cranial and postcranial remains of a new species of Prochelidella (Testudines: Pleurodira: Chelidae) from 'La Buitrera' (Cenomanian of Patagonia, Argentina), with comments on the monophyly of this extinct chelid genus from southern Gondwana |url=https://www.tandfonline.com/doi/full/10.1080/14772019.2020.1721579 |journal=Journal of Systematic Palaeontology |volume=18 |issue=12 |pages=1033–1055 |doi=10.1080/14772019.2020.1721579 |bibcode=2020JSPal..18.1033M |s2cid=214500365 |access-date=2020-07-11}}

|P. buitreraensis

|

|Upper

|

|A small chelid turtle

|

Elkanemys{{Cite journal|last1=Maniel |first1=I.J. |last2=de la Fuente |first2=M. S. |last3=Zhuo |first3=J.I. |title=The first Cearachelyini (Pelomedusoides, Bothremydidae) turtle from the Upper Cretaceous of Patagonia, and an overview of the occurrence and diversity of Pelomedusoides in Patagonia |year=2021 |journal=Cretaceous Research |volume=125 |pages=Article 104869 |doi=10.1016/j.cretres.2021.104869 |bibcode=2021CrRes.12504869M }}

|E. pritchardi

|

|Upper

|

|A bothremydid turtle

|

See also

References

{{Reflist|3}}

{{commons category|Candeleros Formation}}

= Bibliography =

  • {{cite journal |last1=Otero |first1=A. |last2=Carballido |first2=J. L. |last3=Salgado |first3=L. |last4=Canudo |first4=J. I. |last5=Garrido |first5=A. C. |year=2021 |title=Report of a giant titanosaur sauropod from the Upper Cretaceous of Neuquén Province, Argentina |journal=Cretaceous Research |volume=122 |pages=Article 104754|doi=10.1016/j.cretres.2021.104754 |bibcode=2021CrRes.12204754O |s2cid=233582290 }} {{doi|10.1016/j.cretres.2021.104754}}
  • {{cite journal |last1=Canale |first1=Juan I. |last2=Cerda |first2=Ignacio |author3-link=Fernando Novas |last3=Novas |first3=Fernando E. |last4=Haluza |first4=Alejandro |year=2016 |title=Small-sized abelisaurid (Theropoda: Ceratosauria) remains from the Upper Cretaceous of northwest Patagonia, Argentina | journal=Cretaceous Research |volume=62 |pages=18–28|doi=10.1016/j.cretres.2016.02.001 |bibcode=2016CrRes..62...18C |hdl=11336/59930 |hdl-access=free }} {{doi|10.1016/j.cretres.2016.02.001}}
  • {{cite journal |last1=Sánchez |first1=María Lidia |last2=Heredia |first2=Susana |last3=Calvo |first3=Jorge O. |year=2006 |title=Paleoambientes sedimentarios del Cretácico Superior de la Formación Plottier (Grupo Neuquén), Departamento Confluencia, Neuquén (Sedimentary paleoenvironments in the Upper Cretaceous Plottier Formation (Neuquen Group), Confluencia, Neuquén) |url=https://www.researchgate.net/publication/262615949 |journal=Revista de la Asociación Geológica Argentina |volume=61 |pages=3–18 |accessdate=2019-02-16}}
  • {{cite journal |last1=Leanza |first1=H.A. |last2=Apesteguia |first2=S. |author3-link=Fernando Novas |last3=Novas |first3=F.E. |last4=De la Fuente |first4=M.S. |year=2004 |title=Cretaceous terrestrial beds from the Neuquén Basin (Argentina) and their tetrapod assemblages |url=https://www.researchgate.net/publication/257047651 |journal=Cretaceous Research |volume=25 |issue=1 |pages=61–87 |doi=10.1016/j.cretres.2003.10.005 |bibcode=2004CrRes..25...61L |accessdate=2019-02-16}}
  • {{cite web |last=Leonardi |first=Giuseppe |year=1994 |title=Annotated Atlas of South America Tetrapod Footprints (Devonian to Holocene) with an appendix on Mexico and Central America |url=https://www.researchgate.net/publication/309643073 |publisher=Ministerio de Minas e Energia - Companhia de Pesquisa de Recursos Minerais, Geological Service of Brazil |pages=1–248 |accessdate=2019-03-25}}
  • {{cite journal |last=Wichmann |first=R |year=1929 |title=Los Estratos con Dinosaurios y su techo en el este del Territorio del Neuquén ("The dinosaur-bearing strata and their upper limit in eastern Neuquén Territory") |journal=Dirección General de Geología, Minería e Hidrogeología Publicación |volume=32 |pages=1–9}}

Category:Geologic formations of Argentina

Category:Cretaceous Argentina

Category:Sandstone formations

Category:Aeolian deposits

Category:Fluvial deposits

Category:Paludal deposits