:en:Amplectobeluidae

In 2014, Amplectobeluidae was defined as the most inclusive clade including Amplectobelua symbrachiata but not Anomalocaris canadensis, Tamisiocaris borealis, or Hurdia victoria.{{cite journal | vauthors = Vinther J, Stein M, Longrich NR, Harper DA | title = A suspension-feeding anomalocarid from the Early Cambrian | journal = Nature | volume = 507 | issue = 7493 | pages = 496–9 | date = March 2014 | pmid = 24670770 | doi = 10.1038/nature13010 | bibcode = 2014Natur.507..496V | s2cid = 205237459 | url = http://dro.dur.ac.uk/21270/1/21270.pdf | author-link4 = David Harper (palaeontologist) }}

Amplectobeluids could be recognized by frontal appendages with well-developed first distal endite, which forming a pincer-like structure that presumably better suited for a grasping function.{{cite journal |vauthors=Daley AC, Paterson JR, Edgecombe GD, García-Bellido DC, Jago JB |year=2013 |title=New anatomical information on Anomalocaris from the Cambrian Emu Bay Shale and a reassessment of its inferred predatory habits |journal=Palaeontology |volume=56 |issue=5 |pages=971–990 |doi=10.1111/pala.12029 |doi-access=free}} Complete body fossils of amplectobeluids are only known by Amplectobelua and Lyrarapax, both showing combination of characters resembling Anomalocaris (i.e. streamlined body; small head with ovoid sclerites; well-developed swimming flaps; a pair of caudal furcae).{{cite journal |vauthors=Chen JY, Ramsköld L, Zhou GQ |date=May 1994 |title=Evidence for monophyly and arthropod affinity of Cambrian giant predators |journal=Science |volume=264 |issue=5163 |pages=1304–8 |bibcode=1994Sci...264.1304C |doi=10.1126/science.264.5163.1304 |pmid=17780848 |s2cid=1913482}}{{Cite journal |vauthors=Liu J, Lerosey-Aubril R, Steiner M, Dunlop JA, Shu D, Paterson JR |date=2018-11-01 |title=Origin of raptorial feeding in juvenile euarthropods revealed by a Cambrian radiodontan |journal=National Science Review |volume=5 |issue=6 |pages=863–869 |doi=10.1093/nsr/nwy057 |issn=2095-5138 |doi-access=free}}{{cite journal |vauthors=Cong P, Daley AC, Edgecombe GD, Hou X |date=August 2017 |title=The functional head of the Cambrian radiodontan (stem-group Euarthropoda) Amplectobelua symbrachiata |journal=BMC Evolutionary Biology |volume=17 |issue=1 |pages=208 |doi=10.1186/s12862-017-1049-1 |pmc=5577670 |pmid=28854872 |doi-access=free }}{{cite journal |vauthors=Moysiuk J, Caron JB |date=August 2019 |title=A new hurdiid radiodont from the Burgess Shale evinces the exploitation of Cambrian infaunal food sources |journal=Proceedings. Biological Sciences |volume=286 |issue=1908 |pages=20191079 |doi=10.1098/rspb.2019.1079 |pmc=6710600 |pmid=31362637}} Another distinctive features only known in amplectobeluid genera were pairs of gnathobase-like structures (known by Amplectobelua and Ramskoeldia),{{Cite journal |vauthors=Cong PY, Edgecombe GD, Daley AC, Guo J, Pates S, Hou XG |date=2018 |title=New radiodonts with gnathobase-like structures from the Cambrian Chengjiang biota and implications for the systematics of Radiodonta |url=https://serval.unil.ch/resource/serval:BIB_A3714F0A56F0.P001/REF.pdf |journal=Papers in Palaeontology |language=en |volume=4 |issue=4 |pages=605–621 |doi=10.1002/spp2.1219 |s2cid=90258934 |issn=2056-2802}} or an oral cone with combination of tetraradial arrangement and scale-like nodes (known by Lyrarapax and Guanshancaris).{{Cite journal |vauthors=Zeng H, Zhao F, Yin Z, Zhu M |date=2018 |title=A new radiodontan oral cone with a unique combination of anatomical features from the early Cambrian Guanshan Lagerstätte, eastern Yunnan, South China |url=https://www.cambridge.org/core/journals/journal-of-paleontology/article/abs/new-radiodontan-oral-cone-with-a-unique-combination-of-anatomical-features-from-the-early-cambrian-guanshan-lagerstatte-eastern-yunnan-south-china/538DF297E686AA758BD4C4FB609D5770 |journal=Journal of Paleontology |language=en |volume=92 |issue=1 |pages=40–48 |doi=10.1017/jpa.2017.77 |bibcode=2018JPal...92...40Z |issn=0022-3360 |s2cid=134157062}}{{Cite journal |vauthors=Jiao DG, Pates S, Lerosey-Aubril R, Ortega-Hernández J, Yang J, Lan T, Zhang XG |date=2021 |title=The endemic radiodonts of the Cambrian Stage 4 Guanshan biota of South China |journal=Acta Palaeontologica Polonica |language=en |volume=66 |doi=10.4202/app.00870.2020 |issn=0567-7920 |doi-access=free}}

Early in 2014, "Anomalocaris" kunmingensis was tentatively assigned to Amplectobelua by Vinther et al. Later that year, however, the discoverers of Lyrarapax unguispinus ignored that assessment and created a genus within Amplectobelua sensu Vinther et al. An indeterminate frontal appendage assignable to this group is known from the Parker Slate of Vermont.{{Cite journal |last1=Pari |first1=Giovanni |last2=Briggs |first2=Derek E.G. |last3=Gaines |first3=Robert R. |date=2022-02-16 |title=The soft-bodied biota of the Cambrian Series 2 Parker Quarry Lagerstätte of northwestern Vermont, USA |journal=Journal of Paleontology |volume=96 |issue=4 |pages=770–790 |doi=10.1017/jpa.2021.125 |bibcode=2022JPal...96..770P |s2cid=246933116 |issn=0022-3360|doi-access=free }}

An a posteriori-weighted phylogenetic analysis in 2014 found the following relationships within the Amplectobeluidae:{{cite journal | vauthors = Cong P, Ma X, Hou X, Edgecombe GD, Strausfeld NJ | title = Brain structure resolves the segmental affinity of anomalocaridid appendages | journal = Nature | volume = 513 | issue = 7519 | pages = 538–42 | date = September 2014 | pmid = 25043032 | doi = 10.1038/nature13486 | bibcode = 2014Natur.513..538C | s2cid = 4451239 }}

{{clade

|1={{clade

|1=NIGP 154565

|2="Anomalocaris" saron

}}

|2={{clade

|1="Anomalocaris" kunmingensis

|2={{clade

|1=Lyrarapax unguispinus

|2={{clade

|1=Amplectobelua symbrachiata

|2=Amplectobelua stephenensis

}}

}}

}}

}}

{{Reflist|30em}}

{{Radiodonta}}

{{Taxonbar|from=Q17409560}}

Category:Radiodonta

Category:Prehistoric arthropod families