Anacamptis pyramidalis

File:Fertilisation of Orchids figure 4.jpg's book Fertilisation of Orchids included an illustration of the head of a moth with its proboscis laden with several pairs of pollinia from Orchis pyramidalis]]

This hardy plant reaches on average {{convert|10|-|25|cm|in}} of height, with a maximum of {{convert|60|cm|in}}. The stem is erect and unbranched. The basal leaves are linear-lanceolate with parallel venation, up to {{convert|25|cm}} long, the cauline ones are shorter and barely visible on the stem. The arrangement of hermaphroditic flowers in a compact pyramidal shape is very distinctive and gives the orchid its common name. The colour of the flower varies from pink to purple, or rarely white, and the scent is described as "foxy". The flowers have six tepals, being three small sepals and three petals. Two small petals are on the sides, while the third and lower (labellum) is large and trilobate. At the back of the flower there is a tubular spur of about {{convert|1.5|cm|in}} long, while the labellum bears two lateral small flaps. The flowering period extends from April through July.

Anacamptis pyramidalis requires a sunny spot on diverse soils: loamy or clay. It can even grow on very alkaline soil. It can be found on meadows, in grassland, sand dunes, maquis as well as dry and well exposed slopes, at an altitude of {{convert|0|-|1600|m|round=50||disp=x| or more (| approx.) }} above sea level.[http://www.pakbs.org/pjbot/PDFs/44(SI1)/20.pdf Pakistan Journal of Botany - Studies on the morphology, anatomy and ecology of Anacamptis pyramidalis (L.) in Turkey][https://pfaf.org/user/Plant.aspx?LatinName=Anacamptis+pyramidalis Plants for a Future - Anacamptis pyramidalis]

In the UK, Anacamptis pyramidalis is a very successful coloniser of disturbed soil, and can grow in a wide variety of locations, including road verges, reservoirs, quarries and airfields.{{Cite web |url=http://www.plantlife.org.uk/uk/discover-wild-plants-nature/plant-fungi-species/pyramidal-orchid |title=Plant Life - Pyramidal Orchid |access-date=2018-11-08 |archive-date=2018-06-12 |archive-url=https://web.archive.org/web/20180612010825/http://www.plantlife.org.uk/uk/discover-wild-plants-nature/plant-fungi-species/pyramidal-orchid |url-status=dead }}

This orchid is native to southwestern Eurasia, from western Europe through the Mediterranean region eastwards to Iran. In Germany, it is rare and was declared Orchid of the Year in 1990 to heighten awareness of this plant.

This orchid is especially common on the Isle of Wight in the South of England, and was designated the county plant in 2008. On the Isle of Wight, it favours growth in chalky or sandstone-rich soil,{{cite web |title=Pyramidal orchid |url=https://www.plantlife.org.uk/uk/discover-wild-plants-nature/plant-fungi-species/pyramidal-orchid#:~:text=Pyramidal%20orchid%20Anacamptis%20pyramidalis&text=The%20pyramidal%20orchid%20was%20chosen,soil%20in%20order%20to%20bloom. |website=Plantlife |access-date=17 June 2020 |language=en}} and thus can easily be found on the Downland and cliffs to the west and south of the island.{{cite web |title=Isle of Wight Downs IPA |url=https://www.plantlife.org.uk/uk/nature-reserves-important-plant-areas/important-plant-areas/isle-wight-downs |website=Plantlife |access-date=26 June 2020 |language=en}}{{cite web |url=https://www.iow.gov.uk/azservices/documents/2782-FE8-IW-Biodiversity-Action-Plan.pdf |title=Isle of Wight Biodiversity Action Plan, Maritime Cliffs and Slopes Habitat Action Plan |publisher=iow.gov.uk |access-date=26 June 2020 |date=December 2007}}

The flowers are pollinated by butterflies and moths. To ensure the fertilization, their morphology is well adapted to the proboscis of Lepidoptera, especially Euphydryas, Melanargia, Melitaea, Pieris and Zygaena species. The mechanism by which its pairs of pollinia attach themselves to an insect's proboscis was discovered by Charles Darwin and described in his book on the Fertilisation of Orchids.{{harvnb|Darwin|1862|pp=[http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=F800&pageseq=35 20–24], [http://darwin-online.org.uk/content/frameset?viewtype=side&itemID=F800&pageseq=52 37]}}

Anacamptis pyramidalis has been suggested to form mycorrhizal relationships with Rhizoctonia, Fusarium and Papulaspora species.[https://www.researchgate.net/profile/Arzu_Cig/publication/327362280_In_vitro_symbiotic_germination_potentials_of_some_Anacamptis_Dactylorhiza_Orchis_and_Ophrys_terrestrial_orchid_species/links/5b8beee0299bf1d5a7391708/In-vitro-symbiotic-germination-potentials-of-some-Anacamptis-Dactylorhiza-Orchis-and-Ophrys-terrestrial-orchid-species.pdf Applied ecology and environmental research - In Vitro Symbiotic Germination Potentials of Some Anacamptis, Dactylorhiza, Orchis and Ophrys Terrestrial Orchid Species][http://journals.tubitak.gov.tr/botany/abstract.htm?id=10547 Turkish Journal of Botany - Diversity of endophytic fungi from various Aegean and Mediterranean orchids (saleps)]

There are some notable varieties, which are sometimes treated as subspecies – and as they seem to be limited to certain regions, this may be correct:{{citation needed|date=February 2015}}

• Anacamptis pyramidalis var. tanayensis (Chenevard) Soó in Keller – Tanay Pyramidal Orchid - Flowers darker and smaller. Fribourg and Valais cantons (Switzerland).
• Anacamptis pyramidalis var. urvilleana (Sommier & Caruana Gatto) Schlechter – Maltese Pyramidal Orchid, an endemic orchid from Malta with smaller and paler flowers flowering four–six weeks before Anacamptis pyramidalis.{{Cite journal|last=Mifsud|first=Stephen|year=2016|title=Taxonomic notes on Anacamptis pyramidalis var. urvilleana (Orchidaceae), a good endemic orchid from Malta.|journal=Journal Europäischer Orchideen|volume=48 |issue=1|pages=19–28}}
• Anacamptis pyramidalis var. sanguinea (Druce) Kreutz – Western Irish Pyramidal Orchid. -Inflorescence rounder, plant smaller overall. County Galway to County Kerry (Ireland)

The variety alba can be found anywhere in the Pyramidal Orchid's range; its flowers are white.

{{more citations needed|date=June 2020}}

The dried and ground tuber (from various species of Orchis and Anacamptis) can be made into a fine white powder, called salep. This is a very nutritious sweet starchlike substance. It is used in drinks, cereals and in making bread. In Turkey it is used in ice-creams.Eng Soon Teoh {{google books|iEGjDwAAQBAJ|Orchids as Aphrodisiac, Medicine or Food (2019)|page=46}} It was also used medicinally in diets for children and convalescents.

The pyramidal orchid was voted the County flower of the Isle of Wight in 2002 following a poll by the wild flora conservation charity Plantlife.Plantlife website [http://www.plantlife.org.uk/wild_plants/county_flowers/ County Flowers page] {{webarchive|url=https://web.archive.org/web/20150430170312/http://www.plantlife.org.uk/wild_plants/county_flowers/ |date=2015-04-30 }}

{{Reflist}}

• {{Citation

| last =Darwin

| first = Charles

| year =1862

| title =On the various contrivances by which British and foreign orchids are fertilised by insects, and on the good effects of intercrossing

| publication-place = London

| publisher = John Murray

| url =http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=F800&pageseq=1

| access-date =2009-07-30 }}

• {{Commons-inline}}
• {{Wikispecies-inline|Anacamptis pyramidalis|pyramidal orchid (Anacamptis pyramidalis)}}
• {{EOL}}
• [http://linnaeus.nrm.se/flora/mono/orchida/anaca/anacpyrv.jpg Den virtuella floran - Distribution]
• [http://www.maltawildplants.com/ORCH/Anacamptis_pyramidalis.php Comprehensive profile for Anacamptis pyramidalis from the website MaltaWildPlants.com]
• [https://www.biolib.cz/en/taxon/id42168/ Biolib]

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pyramidalis

Category:Orchids of Lebanon

Category:Orchids of Europe

Category:Flora of Germany

Category:Plants described in 1753

Category:Taxa named by Carl Linnaeus

Category:Flora of Malta