Azumamorula mutica

{{Short description|Species of gastropod}}

{{Italic title}}

{{Use dmy dates|date=March 2020}}

{{Taxobox

| name = Azumamorula mutica

| image =

| image_caption =

| regnum = Animalia

| phylum = Mollusca

| classis = Gastropoda

| unranked_superfamilia = clade Caenogastropoda
clade Hypsogastropoda
clade Neogastropoda

| superfamilia = Muricoidea

| familia = Muricidae

| subfamilia = Ergalataxinae

| genus = Azumamorula

| species = A. mutica

| binomial = Azumamorula mutica

| binomial_authority = (Lamarck, 1816)

| synonyms_ref =

| synonyms =

  • Drupa mutica (Lamarck, 1816)
  • Morula (Azumamorula) mutica (Lamarck, 1816)
  • Ricinula mutica Lamarck, 1816 (basionym)
  • Ricinula pisolina Lamarck, 1822

}}

Azumamorula mutica, common name the smooth ricinula, is a species of sea snail, a marine gastropod mollusk in the family Muricidae, the murex snails or rock snails.{{WRMS species|217139|Morula (Azumamorula) mutica (Lamarck, 1816)||2 February 2020}} It is the only species in genus Azumamorula.

Description

The shell size varies between 15 mm and 20 mm

{{Expand section|date=April 2010}}

Distribution

This species is distributed in the Indian Ocean along Madagascar, the Mascarene basin, Réunion, and along Eastern Australia.

Taxonomy

This species was originally named Ricinula mutica by Lamarck in 1816,{{Cite web |title=WoRMS - World Register of Marine Species - Azumamorula mutica (Lamarck, 1816) |url=https://www.marinespecies.org/aphia.php?p=taxdetails&id=1335434 |access-date=2023-11-19 |website=www.marinespecies.org}} grouping it with species that are now mostly in Drupa and Morula.{{efn|R. horrida (D. morum Röding 1798), R. arachnoides (Murex ricinus Linnaeus, now D. ricinus), R. aspera (Mor. aspera), R. clathrata (D. clathrata), R. nodus (D. uva Röding 1798, Mor. uva), and R. digitata (Drupa grossularia Röding 1798, Drupina grossularia)}} In 1823, Dall defined two new genera, Drupina and Morulina, to hold species of Drupa and Morula having simple non-plaited columnellae. He chose R. mutica as the type species for Morulina.{{efn|Along with Morulina mutica, Dall lists triangulata, fiscellum (Murichorda fiscellum, Muricodrupa fiscellum), ocellata (Ergalatax contracta), ambusta (Morula spinosa), musiva (Tenguella musiva), ochrostoma (Pascula ochrostoma), polyphrostoma, dumosa (Purpura dumosa), fuscoimbricata, fenestrata (Muricodrupa fenestrata), and elata (Drupella cornus)}}{{cite journal |last1=Dall |first1=W. H. |title=Notes on Drupa and Morula |journal=Proceedings of the Academy of Natural Sciences of Philadelphia |date=1923 |volume=75 |pages=303–306 |jstor=4063885 }}

Thiele in 1931 and Wenz in 1941 had placed Morulina into Drupa. In 1968, Emerson found that the teeth on the radula of D. mutica had an unusual shape. As a result, he desired to split it out to its own genus, but the name Morulina was already taken as a genus of arthropod. Thus he introduced a new name Azumamorula, recognising Masao Azuma who had extracted the radula specimen.{{cite journal |last1=Emerson |title=Azumamorula, new name for Morulina Dall, 1923, not Boerner, 1906 (Gastropoda: Muricacea) |journal=The Nautilus |date=1968 |volume=81 |issue=4 |pages=125–126 |url=https://www.biodiversitylibrary.org/page/8528150 |accessdate=1 February 2020}}

In 2004, Houart placed Azumamorula and Oppomorus as subgenera of Morula, and moved Morula from subfamily Rapaninae to Ergalataxinae.{{cite journal |last1=Houart |first1=Roland |title=Review of the Recent species of Morula (Oppomorus), M. (Azumamorula) and M. (Habromorula) (Gastropoda: Muricidae: Ergalataxinae) |journal=Novapex |date=2004-11-10 |volume=5 |issue=4 |pages=91–130 |url=https://www.biodiversitylibrary.org/page/42323468#page/261/}}Molluscabase lists Drivas & Jay (1987) as the basis for its record on M. (A.) mutica. MolluscaBase eds. (2020). MolluscaBase. Morula (Azumamorula) mutica (Lamarck, 1816). Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=217139 on 2020-02-01 However, in 2013, a phylogenetic study of the sequences for four genes indicated that Morula (Morula) sensu stricto was polyphyletic, and Morula (Oppomorus) was a separate clade.{{Cite journal|last=Claremont|first=Martine |last2=Houart|first2=Roland |last3=Williams|first3=Suzanne T. |last4=Reid|first4=David G. |date=2013-02-01 |title=A molecular phylogenetic framework for the Ergalataxinae (Neogastropoda: Muricidae) |journal=Journal of Molluscan Studies |language=en |volume=79|issue=1|pages=19–29 |doi=10.1093/mollus/eys028|issn=0260-1230|doi-access=free}} The relevant genes from A. mutica haven't been sequenced, but it's unlikely to be part of a more-restricted Morula–Habromorula group. As of 2020, Houart lists Azumamorula as a separate genus{{Cite journal|last=Houart|first=Roland |last2=Zuccon|first2=Dario |last3=Puillandre|first3=Nicolas |date=2019-04-09 |title=Description of new genera and new species of Ergalataxinae (Gastropoda: Muricidae) |url=https://www.researchgate.net/publication/332301456}} and Molluscabase classes the combination Azumamorula mutica as "accepted".MolluscaBase (2019). MolluscaBase. Azumamorula mutica (Lamarck, 1816). Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=1335434 on 2020-02-01

Notes

{{Notelist}}

References

{{Reflist}}

  • Lamarck, J.B.P.A. de M. 1816. Liste des objets représentés dans les planches de cette livraison. pp. 1–16 in Lamarck, J.B.P.A. de M. Tableau encyclopédique et méthodique des trois règnes de la nature. Vers, coquilles, mollusques et polypiers. Paris : Agasse Part 23 pp. 1–16, pls 391–488.
  • Reeve, L.A. 1846. Monograph of the genus Ricinula. pls 1–6 in Reeve, L.A. (ed). Conchologia Iconica. London : L. Reeve & Co. Vol. 3.
  • Dautzenberg, Ph. (1929). Mollusques testacés marins de Madagascar. Faune des Colonies Francaises, Tome III
  • Drivas, J. & M. Jay (1988). Coquillages de La Réunion et de l'île Maurice
  • Wilson, B. 1994. Australian Marine Shells. Prosobranch Gastropods. Kallaroo, WA : Odyssey Publishing Vol. 2 370 pp.