Bothrocophias campbelli

{{Speciesbox

| image = Bothrocophias campbelli 253005085.jpg

| status = VU

| status_system = IUCN3.1

| status_ref = Caicedo JR, Calderón M, Ines Hladki A, Mendoza-Quijano F, Ramírez Pinilla M, Renjifo J, Urbina N (2019). "Bothrocophias campbelli ". The IUCN Red List of Threatened Species 2019: https://dx.doi.org/10.2305/IUCN.UK.2019-2.RLTS.T15203980A15203988.en. Accessed on 05 January 2022.

| genus = Bothrocophias

| species = campbelli

| authority = Freire-Lascano, 1991

| synonyms = *Bothrops campbelli
{{small|Freire-Lascano, 1991}}

Porthidium almawebi
{{small|Schätti & Kramer, 1993}}McDiarmid RW, Campbell JA, Touré TA (1999). Snake Species of the World: A Taxonomic and Geographic Reference, Volume 1. Washington, District of Columbia: Herpetologists' League. 511 pp. {{ISBN|1-893777-00-6}} (series). {{ISBN|1-893777-01-4}} (volume).
Bothrocophias campbelli
{{small|— Gutberlet & Campbell}}{{EMBL species|genus=Bothrocophias|species=campbelli}} www.reptile-database.org.

}}

Bothrocophias campbelli, commonly known as Campbell's toadheaded viper, the Ecuadorian toadheaded pitviper, and víbora boca de sapo in Spanish, is a species of venomous pitviper in the family Viperidae. The species is endemic to South America.Campbell JA, Lamar WW (2004). The Venomous Reptiles of the Western Hemisphere. 2 volumes. Ithaca and London: Comstock Publishing Associates. 870 pp. 1,500 plates. {{ISBN|0-8014-4141-2}}. No subspecies are currently recognized.{{ITIS |id=634857|taxon=Bothrocophias campbelli |accessdate=27 August 2008}}

Etymology

Bothrocophias campbelli was named in honor of American herpetologist Jonathan A. Campbell.Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). The Eponym Dictionary of Reptiles. Baltimore: Johns Hopkins University Press. xiii + 296 pp. {{ISBN|978-1-4214-0135-5}}. (Bothrocophias campbelli, p. 46).

Description

The total length (including tail) of the specimens of B. campbelli which are available ranges from the type specimen, an adult male {{convert|87.0|cm|in|frac=4|abbr=on}} (Freire-Lascano, 1991), to {{convert|105.7|cm|in|frac=8|abbr=on}} for a large gravid female (Freire-Lascano & Kuch, 2000), to {{convert|121.9|cm|in|abbr=on}} for a specimen from Imbabura, Ecuador, and another of {{convert|123.0|cm|in|frac=8|abbr=on}} from the type locality.

The scalation includes 21–25 (usually 23) rows of dorsal scales at midbody, all of which are keeled except for the first rows. The posterior dorsals have tubercular keels, even in small specimens, while in large specimens these keels are highly elevated. There are 152-167/159-177 ventral scales in males/females and 51-64/48-60 paired subcaudal scales in males/females. On the head there are 3–8 intersupraocular scales, 7–8 supralabial scales, the second of which is usually fused with the prelacunal to form a lacunolabial, and 9–10 (usually 9) sublabial scales.

Geographic range

Bothrocophias campbelli is found in the Pacific lowlands and slopes from west-central Colombia to Ecuador. The type locality given is "Recinto Huagal-Sacramento, cantón Pallatanga, Provincia de Chimborazo, Ecuador. 1500–2000 mts. de altura [4,900–6,600 feet]".

Habitat

The preferred natural habitat of B. campbelli is forest, at altitudes of {{convert|800|–|2,000|m|ft|abbr=on}}.

Diet

B. campbelli preys upon caecilians, lizards, snakes of the genus Atractus, and small rodents.

Reproduction

B. campbelli is viviparous.

Taxonomy

In previous accounts, Bothrocophias campbelli has often been included in Bothrops pulcher (W. Peters, 1862), e.g., Lachesis pulcher — Boulenger, 1896, Bothrops pulchra — Amaral, 1923, and Bothrops pulcher — J.Peters & Orejas-Miranda, 1970. Campbell and Lamar (1992) considered Bothrocophias campbelli to be a junior synonym of Bothrops pulcher, but their concept of the latter taxon was incorrect. Schätti and Kramer (1993) argued that Bothrops campbelli (Freire-Lascano, 1991) was a nomen invalidum and suggested a new name, Porthidium almawebi, as a replacement. However, it is clear that Freire-Lascano's (1991) scientific name is valid and has priority over the one proposed by Schätti and Kramer (1993).

Gutberlet and Campbell (2001) moved this taxon to a new genus: Bothrocophias (toadheaded pitvipers).