Coelopidae

{{For|terms|Morphology of Diptera}}

Coelopids are small to medium-sized ({{convert|2.5|–|9|mm|abbr=on|sigfig=2}}, usually {{convert|4|–|7|mm|abbr=on}}), robust flies, predominantly with a flat body and darkly coloured. Coelopidae species are usually densely bristly or hairy. Their eyes are small. The arista is bare to pubescent. Ocelli and ocellar bristles are present. The postvertical bristles are parallel or converge. The two pairs of frontal bristles curve outward and scattered interfrontal setulae are present. Vibrissae are absent, but strong bristles occur near the vibrissal angle. The mesonotum is flat and the prothorax is separated from the propleuron by a membrane. The legs bear strong bristles and soft, dense hairs and the tibiae have subapical bristles. The wing is unmarked. The costa is entire, without interruptions. The subcosta is complete, crossvein BM-Cu is present, and the anal cell (cell cup) is closed. Legs usually densely hairy.

Coelopids are found in the wrack zone of temperate seashores where the larvae feed on rotting seaweed. They are sometimes very abundant in this habitat. They go through several generations a year. The females lay their eggs in small batches into fresh alga banks. Three larval instars occur. Larvae feed in a bacteria-laden mass. Pupation is seldom in the algal substrate that soon collapses, but more frequently in the highest sand layers. Larvae are also found in winter wrack heaps as bacteria raise temperatures to 20-30 °C even if the heap is superficially frozen. Larvae and pupae have numerous predators, including birds and the staphylinid Aleochara and suites of parasites confined to algal banks.

{{As of|2011}}, following Mathis and McAlpine's taxonomy, Coelopidae consists of two subfamilies: Coelopinae, with three tribes, twelve genera, and twenty-eight species, and Lopinae, consisting of just one monospecific genus.{{cite book|last1=Mathis|first1=Wayne N.|last2=McAlpine|first2=David K.|chapter=A Catalog and Conspectus on the Family Coelopidae (Diptera: Schizophora)|series=Myia|date=2011|volume=12|pages=171–205|title=Contributions to the Systema Dipterorum (Insecta: Diptera)|isbn=978-954-642-599-7|editor1-last=Brake|editor1-first=Irina|editor2-last=Thompson|editor2-first=F. Christian|hdl-access=free|hdl=10088/18924}}

• Subfamily Coelopinae Hendel, 1910
• Tribe Coelopini Hendel, 1910
• Genus Coelopa Meigen, 1830
• Subgenus Coelopa Meigen, 1830
• Subgenus Fucomyia Haliday, 1837
• Subgenus Neocoelopa Malloch, 1933{{cite journal |last1=Malloch |first1=J. R. |title=The genus Coelopa Meigen (Diptera, Coelopidae) |journal=The Annals and Magazine of Natural History |date=1933 |volume=11 |issue=10 |pages=339–350|doi=10.1080/00222933308673666 }}
• Tribe Coelopellini McAlpine, 1991{{cite journal |last1=McAlpine |first1=David K. |title=Review of the Australian kelp flies (Diptera: Coelopidae) |journal=Systematic Entomology |date=1991 |volume=16 |pages=29–84|doi=10.1111/j.1365-3113.1991.tb00573.x |s2cid=83627067 }}
• Genus Amma McAlpine, 1991
• Genus Beaopterus Lamb, 1909{{cite journal|last1=Lamb|first1=C. G.|title=The Diptera of the subantarctic Islands of New Zealand. Pp. 124-145. In Chilton, C. (ed.), The subantarctic islands of New Zealand|date=1909|pages=xxxv + 388 pp| url = https://www.biodiversitylibrary.org/item/44371#page/170/mode/1up|publisher=Phil. Inst.|access-date=1 December 2019|location=Canterbury, Wellington}}
• Genus Coelopella Malloch, 1933
• Genus Icaridion Lamb, 1909 Halteres absent and the wings are reduced to strips. New Zealand.
• Genus Rhis McAlpine, 1991
• Genus This McAlpine, 1991
• Tribe Glumini McAlpine, 1991
• Genus Chaetocoelopa Malloch, 1933
• Genus Coelopina Malloch, 1933
• Genus Dasycoelopa Malloch, 1933
• Genus Gluma McAlpine, 1991
• Genus Malacomyia Haliday in Westwood, 1840 (sometimes placed in Dryomyzidae
• Subfamily Lopinae McAlpine, 1991
• Genus Lopa McAlpine, 1991

Coelopa frigida (Fabricius) has been reared in the laboratory and used for genetic studies.

• Chaetocoelopa littoralis

{{Reflist}}

{{Refbegin}}

• {{cite journal

| last = McAlpine

| first = David K.

| year = 1991

| title = Review of the Australian Kelp Flies (Diptera: Coelopidae)

| journal = Systematic Entomology

| volume = 16

| pages = 29–84

| doi=10.1111/j.1365-3113.1991.tb00573.x

| s2cid = 83627067

}}

• {{cite book|last=McAlpine |first=David K. |date=1998 |chapter=Family Coelopidae |editor1-last=Papp |editor1-first=L. |editor2-last=Darvas |editor2-first=B. |title=Contributions to a Manual of Palaearctic Diptera |volume=3 |pages=335–340 |publisher=Science Herald |location=Budapest}}
• {{cite journal|last1=Meier|first1=Rudolf|last2=Wiegmann|first2=Brian M.|title=A phylogenetic analysis of Coelopidae (Diptera) based on morphological and DNA sequence data|journal=Molecular Phylogenetics and Evolution|date=2002|volume=25|issue=3|pages=393–407|doi=10.1016/S1055-7903(02)00276-2|pmid=12450746}}

{{Refend}}

{{Wikispecies|Coelopidae}}

{{Commons category}}

• [http://tolweb.org/Coelopidae/10652 Tree of Life Coelopidae]
• [http://eol.org/pages/478/media Family Coelopidae at EOL]
• [http://www.diptera.info/photogallery.php?album_id=100 Diptera.info] Images
• [http://bugguide.net/node/view/58825/bgimage BugGuide] images
• [http://delta-intkey.com/britin/dip/www/coelopid.htm]
• [https://web.archive.org/web/20060517034556/http://www.faunistik.net/DETINVERT/DIPTERA/COELOPIDAE/IMAGES/coelopa.frigida_pu01.gif Image of pupa]

• Hennig. 1937. Coelopidae.In: Lindner, E. (Ed.). Die Fliegen der Paläarktischen Region 5, 52, 1-38.Keys to Palaearctic species but now needs revision (in German).
• Malloch, J.R. 1933. The genus Coelopa Meigen (Diptera, Coelopidae). Ann. Mag. Nat. Hist. (10) 11: 339–50.
• McAlpine, David K. (1991). "Review of the Australian Kelp Flies (Diptera: Coelopidae)" (Print). Systematic Entomology 16: 29–84.
• Séguy, E. (1934) Diptères: Brachycères. II. Muscidae acalypterae, Scatophagidae. Paris: Éditions Faune de France 28. [https://web.archive.org/web/20131014031511/http://www.faunedefrance.org/BibliothequeVirtuelleNumerique%7CBiblioth%C3%A8que virtuelle numérique]
• Shtakel'berg, A.A. Family Coelopidae in Bei-Bienko, G. Ya, 1988 Keys to the insects of the European Part of the USSR Volume 5 (Diptera) Part 2 English edition.Keys to Palaearctic species but now needs revision .

• [https://web.archive.org/web/20051015233259/http://www.faunaeur.org/full_results.php?id=10903 West Palaearctic and Russia]
• [http://www.nearctica.com/nomina/main.htm Nearctic]
• [http://hbs.bishopmuseum.org/aocat/coelopidae.html Australasian/Oceanian]
• [https://web.archive.org/web/20190527005729/http://konchudb.agr.agr.kyushu-u.ac.jp/mokuroku/ Japan]

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Category:Brachycera families