Acalyptratae
{{Short description|Assemblage of flies}}
{{Automatic taxobox
| image = Marsh_fly01.jpg
| image_caption = Marsh fly (Sciomyzidae)
| display_parents = 5
| taxon = Acalyptratae
| subdivision_ranks = Superfamilies
| subdivision =
}}
The Acalyptratae or Acalyptrata are a subsection of the Schizophora, which are a section of the order Diptera, the "true flies". In various contexts the Acalyptratae also are referred to informally as the acalyptrate muscoids, or acalyptrates, as opposed to the Calyptratae. All forms of the name refer to the lack of calypters in the members of this subsection of flies. An alternative name, Acalypterae is current, though in minority usage. It was first used by Pierre-Justin-Marie Macquart in 1835 for a section of his tribe Muscides; he used it to refer to all acalyptrates plus scathophagids and phorids, but excluding Conopidae.
The confusing forms of the names stem from their first usage; Acalyptratae and Acalyptrata actually are adjectival forms in Neo-Latin. They were coined in the mid 19th century in contexts such as "Muscae Calyptratae and Acalyptratae" and "Diptera Acalyptrata", and the forms stuck.{{cite book|author=Carl Robert Osten-Sacken|title=Collected papers|url=https://books.google.com/books?id=geJMAAAAYAAJ&pg=RA1-PA127|year=1858|pages=1–}}
The Acalyptratae are a large assemblage, exhibiting very diverse habits, with one notable and perhaps surprising exception: no known acalyptrates are obligate blood-feeders (hematophagous), though blood feeding at various stages of the life history is common throughout other Dipteran sections.
Classification
The classification of the Acalyptratae has varied over time, and the below list is likely to change in future.
- Subsection Acalyptratae
- Superfamily Conopoidea
- Conopidae
- Superfamily Tephritoidea{{Cite journal |last1=Han |first1=Ho-Yeon |last2=Ro |first2=Kyung-Eui |year=2016 |title=Molecular phylogeny of the superfamily Tephritoidea (Insecta: Diptera) reanalysed based on expanded taxon sampling and sequence data |url=https://onlinelibrary.wiley.com/doi/10.1111/jzs.12139 |journal=Journal of Zoological Systematics and Evolutionary Research |language=en |volume=54 |issue=4 |pages=276–288 |doi=10.1111/jzs.12139|doi-access=free |url-access=subscription }}
- Ctenostylidae
- Eurygnathomyiidae
- Lonchaeidae
- Pallopteridae
- Piophilidae
- Platystomatidae
- Pyrgotidae
- Richardiidae
- Tephritidae (including Tachiniscidae)
- Ulidiidae
- Superfamily Nerioidea{{Cite journal |last=Lonsdale |first=Owen |date=2020-02-17 |title=Family groups of Diopsoidea and Nerioidea (Diptera: Schizophora)—Definition, history and relationships |url=https://www.biotaxa.org/Zootaxa/article/view/zootaxa.4735.1.1 |journal=Zootaxa |volume=4735 |issue=1 |pages=zootaxa.4735.1.1 |doi=10.11646/zootaxa.4735.1.1 |pmid=32230232 |s2cid=214311790 |issn=1175-5334|doi-access=free |url-access=subscription }}
- Cypselosomatidae
- Fergusoninidae
- Micropezidae
- Neriidae
- Pseudopomyzidae
- Strongylophthalmyiidae
- Tanypezidae
- Superfamily Diopsoidea
- Diopsidae
- Gobryidae
- Megamerinidae
- Nothybidae
- Psilidae
- Somatiidae
- Syringogastridae
- Superfamily Sciomyzoidea{{Cite web |title=Sciomyzoidea [ ] |url=https://diptera.myspecies.info/category/diptera-classification/sciomyzoidea |access-date=2022-11-01 |website=diptera.myspecies.info}}
- Coelopidae
- Dryomyzidae
- Helcomyzidae
- Helosciomyzidae
- Heterocheilidae
- Ropalomeridae
- Sepsidae
- Sciomyzidae (including Huttoninidae and Phaeomyiidae)
- Superfamily Sphaeroceroidea{{Cite web |title=Sphaeroceroidea [ ] |url=https://diptera.myspecies.info/category/diptera-classification/sphaeroceroidea |access-date=2022-11-01 |website=diptera.myspecies.info}}
- Chyromyidae
- Heleomyzidae
- Heteromyzidae (disputed)
- Nannodastiidae
- Sphaeroceridae
- Superfamily Lauxanioidea{{Cite journal |last1=Li |first1=Xuankun |last2=Li |first2=Wenliang |last3=Ding |first3=Shuangmei |last4=Cameron |first4=Stephen |last5=Mao |first5=Meng |last6=Shi |first6=Li |last7=Yang |first7=Ding |date=2017-04-14 |title=Mitochondrial Genomes Provide Insights into the Phylogeny of Lauxanioidea (Diptera: Cyclorrhapha) |journal=International Journal of Molecular Sciences |language=en |volume=18 |issue=4 |pages=773 |doi=10.3390/ijms18040773 |issn=1422-0067 |pmc=5412357 |pmid=28420076|doi-access=free }}
- Celyphidae
- Chamaemyiidae
- Lauxaniidae
- Superfamily Opomyzoidea
- Agromyzidae
- Anthomyzidae
- Asteiidae
- Aulacigastridae
- Clusiidae
- Marginidae
- Neminidae
- Neurochaetidae
- Odiniidae
- Opomyzidae
- Periscelididae
- Teratomyzidae
- Xenasteiidae
- Superfamily Ephydroidea{{Cite web |title=Ephydroidea [ ] |url=https://diptera.myspecies.info/category/diptera-classification/ephydroidea |access-date=2022-11-01 |website=diptera.myspecies.info}}
- Camillidae
- Curtonotidae
- Diastatidae
- Drosophilidae
- Ephydridae
- Mormotomyiidae
- Superfamily Carnoidea{{Cite web |title=Carnoidea [ ] |url=https://diptera.myspecies.info/category/diptera-classification/carnoidea |access-date=2022-11-01 |website=diptera.myspecies.info}}
- Acartophthalmidae
- Australimyzidae
- Braulidae
- Canacidae
- Carnidae
- Chloropidae
- Cryptochetidae
- Inbiomyiidae
- Milichiidae
- Acalyptratae incertae sedis
- Paraleucopidae
References
{{Reflist}}
External links
{{Wiktionary|Acalyptrata}}
- {{cite journal |last=McAlpine |first=David K. |year=1958 |title=A key to the Australian families of Acalptrate Diptera (Insecta) |journal=Records of the Australian Museum |volume=24 |issue=12 |pages=183–190 |url=http://australianmuseum.net.au/journal/McAlpine-1958-Rec-Aust-Mus-2412-183190/ |doi=10.3853/j.0067-1975.24.1958.650|doi-access=free }}
- {{cite web |title=Acalyptratae |work=Atlas of Living Australia |url=https://bie.ala.org.au/species/NZOR-4-102780 |access-date=2018-03-30 |archive-url=https://web.archive.org/web/20190113182356/https://bie.ala.org.au/species/NZOR-4-102780 |archive-date=2019-01-13 |url-status=dead }}
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