Lulworth Formation

{{Infobox rockunit

| name = Lulworth Formation

| image = Lulworth_cove.JPG

| caption = Lulworth Cove, an area where the Lulworth Formation is exposed

| type = Formation

| age = Tithonian - Berriasian {{Fossil range|152|139}}

| period = Berriasian

| prilithology = Calcarenite, Micrite, Mudstone, Marl

| otherlithology = Gypsum

| namedfor = Lulworth

| namedby =

| region = England

| country = United Kingdom

| coordinates =

| unitof = Purbeck Group

| subunits = * Worbarrow Tout Member

Ridgway Member
Mupe Member

| underlies = Durlston Formation

| overlies = Portland Stone Formation

| thickness = In Dorset 27-63 metres, In Weald 86 m Vale of Wardour up to 15m

| extent =

| area =

| map =

| map_caption =

| location_ts = West side of Worbarrow Tout

}}

The Lulworth Formation is a geologic formation in England. It dates from the late Tithonian to the mid Berriasian. It is a subunit of the Purbeck Group.{{cite web |url= http://www.bgs.ac.uk/lexicon/lexicon.cfm?pub=LULW|title= Lulworth Formation|publisher= British Geological Survey|work = BGS Lexicon of Named Rock Units|access-date= 28 August 2018}} In Dorset, it consists of three members, which are in ascending order, the Mupe Member, the Ridgway Member, and the Warbarrow Tout Member. The Mupe Member is typically 11 to 16 m thick and largely consists of marls and micrites with interbeds of calcareous mudstone.{{cite web |url= http://www.bgs.ac.uk/lexicon/lexicon.cfm?pub=MUP|title= Mupe Member|publisher= British Geological Survey|work = BGS Lexicon of Named Rock Units|access-date= 28 August 2018}} The Ridgeway Member is about 3 to 7 m thick and consists of in its western portion carbonaceous muds, marls and micrites, in the east the muds are replaced by micritic limestone.{{cite web |url= http://www.bgs.ac.uk/lexicon/lexicon.cfm?pub=RID|title= Ridgeway Member|publisher= British Geological Survey|work = BGS Lexicon of Named Rock Units|access-date= 28 August 2018}} The Warbarrow Tout Member is 17 to 39 m thick and consists of limestone at the base and micrite and mudstone for the rest of the sequence,{{cite web |url= http://www.bgs.ac.uk/lexicon/lexicon.cfm?pub=WOT|title= Warbarrow Tout Member|publisher= British Geological Survey|work = BGS Lexicon of Named Rock Units|access-date= 28 August 2018}} this member is the primary source of the vertebrate fossils within the formation.{{cite web |url= https://paleobiodb.org/classic?a=basicCollectionSearch&collection_no=28401|title= Sunnydown Farm Quarry (clay: upper horizon; equivalent to DB 102/103 )|work = Paleobiology Database|access-date= 28 August 2018}}{{cite web |url= https://paleobiodb.org/classic?a=basicCollectionSearch&collection_no=72142|title= Mammal Bed, Durlston Bay (DB83)|work = Paleobiology Database|access-date= 28 August 2018}} Elsewhere the unit is undifferentiated.

Vertebrate paleobiota

= Amphibians =

class="wikitable" align="center" width="100%" ! colspan="7" align="center" \|Amphibians reported from the Lulworth Formation{{Cite journal\|last1=Evans\|first1=Susan E.\|last2=McGowan\|first2=Gerard J.\|date=2002\|title=Lissamphibian remains from the Purbeck Limestone Group, southern England\|journal=Special Papers in Palaeontology\|pages=104–119}}
Genus !Species !Location !Stratigraphic position !Material !Notes !Images
Apricosiren \|A. ensomi \| \|Worbarrow Tout Member \| \|Salamander \|
Celtedens \|C. megacephalus \| \|Worbarrow Tout Member \| \|Albanerpetontid \|
Sunnybatrachus \|S. purbeckensis \| \|Worbarrow Tout Member \| \|Frog, affinites to Discoglossidae \|
?Batrachosauroididae \|Indeterminate \| \|Worbarrow Tout Member \| \|Salamander \|

class="wikitable" align="center" width="100%"

! colspan="7" align="center" |Amphibians reported from the Lulworth Formation{{Cite journal|last1=Evans|first1=Susan E.|last2=McGowan|first2=Gerard J.|date=2002|title=Lissamphibian remains from the Purbeck Limestone Group, southern England|journal=Special Papers in Palaeontology|pages=104–119}}

Genus

!Species

!Location

!Stratigraphic position

!Material

!Notes

!Images

Apricosiren

|A. ensomi

|Worbarrow Tout Member

|Salamander

Celtedens

|C. megacephalus

|Worbarrow Tout Member

|Albanerpetontid

Sunnybatrachus

|S. purbeckensis

|Worbarrow Tout Member

|Frog, affinites to Discoglossidae

?Batrachosauroididae

|Indeterminate

|Worbarrow Tout Member

|Salamander

= Turtles =

class="wikitable" align="center" width="100%" ! colspan="7" align="center" \|Turtles reported from the Lulworth Formation{{Cite journal\|last=Milner\|first=Andrew R.\|date=November 2004\|title=The turtles of the Purbeck Limestone Group of Dorset, southern England\|journal=Palaeontology\|volume=47\|issue=6\|pages=1441–1467\|doi=10.1111/j.0031-0239.2004.00418.x\|bibcode=2004Palgy..47.1441M \|issn=0031-0239\|url=https://eprints.bbk.ac.uk/55/1/Purbeck_turtles.pdf}}
Genus !Species !Location !Stratigraphic position !Material !Notes !Images
Dorsetochelys \|D. typocardium, D. delairi \| \| \| \|Both species probably represent the same taxon{{Cite journal\|last=Pérez-García\|first=A.\|date=May 2014\|title=Revision of the poorly known Dorsetochelys typocardium, a relatively abundant pleurosternid turtle (Paracryptodira) in the Early Cretaceous of Europe\|journal=Cretaceous Research\|volume=49\|pages=152–162\|doi=10.1016/j.cretres.2014.02.015\|bibcode=2014CrRes..49..152P \|issn=0195-6671}} \|
Helochelydra \|H. anglica \| \| \| \| \|
Hylaeochelys \|H. latiscutata \| \| \| \| \|
Pleurosternon \|P. bullocki \| \| \| \| \|

class="wikitable" align="center" width="100%"

! colspan="7" align="center" |Turtles reported from the Lulworth Formation{{Cite journal|last=Milner|first=Andrew R.|date=November 2004|title=The turtles of the Purbeck Limestone Group of Dorset, southern England|journal=Palaeontology|volume=47|issue=6|pages=1441–1467|doi=10.1111/j.0031-0239.2004.00418.x|bibcode=2004Palgy..47.1441M |issn=0031-0239|url=https://eprints.bbk.ac.uk/55/1/Purbeck_turtles.pdf}}

Genus

!Species

!Location

!Stratigraphic position

!Material

!Notes

!Images

Dorsetochelys

|D. typocardium, D. delairi

|Both species probably represent the same taxon{{Cite journal|last=Pérez-García|first=A.|date=May 2014|title=Revision of the poorly known Dorsetochelys typocardium, a relatively abundant pleurosternid turtle (Paracryptodira) in the Early Cretaceous of Europe|journal=Cretaceous Research|volume=49|pages=152–162|doi=10.1016/j.cretres.2014.02.015|bibcode=2014CrRes..49..152P |issn=0195-6671}}

Helochelydra

|H. anglica

Hylaeochelys

|H. latiscutata

Pleurosternon

|P. bullocki

= Lepidosaurs =

class="wikitable" align="center" width="100%" ! colspan="7" align="center" \|Lepidosaurs reported from the Lulworth Formation
Genus !Species !Location !Stratigraphic position !Material !Notes !Images
Becklesius \|Indeterminate \| \| \| \| \|
Dorsetisaurus \|D. hebetidens, D. purbeckensis \| \| \| \| \|
Durotrigia \|D. triconidens \| \| \| \| \|
Homoeosaurus \|Indeterminate \| \| \| \|Rhynchocephalian \|
Opisthias \|Indeterminate \| \| \| \|Rhynchocephalian \|
Paramacellodus \|P. oweni \| \| \| \| \|
Parasaurillus \|P. pseudobtusus \| \| \| \| \|
Parviraptor \|P. estesi \| \| \| \|Stem-snake \|
Pseudosaurillus \|P. becklesi \| \| \| \| \|
Purbicella \|P. ragei \| \| \| \| \|
Saurillus \|S. robustidens \| \| \| \| \|

class="wikitable" align="center" width="100%"

! colspan="7" align="center" |Lepidosaurs reported from the Lulworth Formation

Genus

!Species

!Location

!Stratigraphic position

!Material

!Notes

!Images

Becklesius

|Indeterminate

Dorsetisaurus

|D. hebetidens, D. purbeckensis

Durotrigia

|D. triconidens

Homoeosaurus

|Indeterminate

|Rhynchocephalian

Opisthias

|Indeterminate

|Rhynchocephalian

Paramacellodus

|P. oweni

Parasaurillus

|P. pseudobtusus

Parviraptor

|P. estesi

|Stem-snake

Pseudosaurillus

|P. becklesi

Purbicella

|P. ragei

Saurillus

|S. robustidens

= Crocodyliformes =

class="wikitable" align="center" width="100%" ! colspan="7" align="center" \|Crocodyliformes reported from the Lulworth Formation
Genus !Species !Location !Stratigraphic position !Material !Notes !Images
cf. Bernissartia \|Indeterminate \| \| \|Teeth \| \|
Goniopholis \|Indeterminate \| \| \| \| \|
Nannosuchus \|N. gracilidens \| \| \| \| \|
Theriosuchus \|T. pusillus \| \| \| \| \|

class="wikitable" align="center" width="100%"

! colspan="7" align="center" |Crocodyliformes reported from the Lulworth Formation

Genus

!Species

!Location

!Stratigraphic position

!Material

!Notes

!Images

cf. Bernissartia

|Indeterminate

|Teeth

Goniopholis

|Indeterminate

Nannosuchus

|N. gracilidens

Theriosuchus

|T. pusillus

= Dinosaurs =

class="wikitable" align="center" width="100%" ! colspan="7" align="center" \|Dinosaurs reported from the Lulworth Formation
Genus !Species !Location !Stratigraphic position !Material !Notes !Images
Echinodon \|E. becklesii{{Cite web\|url=https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=58901\|title=Type specimen: BMNH 48209, 48210, a mandible. Its type locality is cliff face, Durlston Bay, which is in a Berriasian fluvial-lacustrine limestone in the Lulworth Formation of the United Kingdom.\|website=fossilworks.org}} \| Durlston Bay, Swanage \| \|Teeth and dentary \|A small heterodontosaurid \|
Owenodon \|O. hoggii{{Cite web\|url=https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=163545\|title=Type specimen: BMNH R2998. Its type locality is Durlston Bay, Swanage (DB105-107), which is in a Berriasian marine limestone in the Lulworth Formation of the United Kingdom.}} \|Durlston Bay, Swanage \| \|Partial dentary \|An iguanodontoid \|
Nuthetes \|N. destructor{{Cite web\|url=https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=66817\|title=Type specimen: DORCM G913, a mandible. Its type locality is Feather Quarry, Durlston Bay (DB102), which is in a Berriasian marine shale in the Lulworth Formation of the United Kingdom.}} \| Feather Quarry, Durlston Bay, Swanage \| \|Teeth, dentary fragment \|Small dromaeosaur \|

class="wikitable" align="center" width="100%"

! colspan="7" align="center" |Dinosaurs reported from the Lulworth Formation

Genus

!Species

!Location

!Stratigraphic position

!Material

!Notes

!Images

Echinodon

|E. becklesii{{Cite web|url=https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=58901|title=Type specimen: BMNH 48209, 48210, a mandible. Its type locality is cliff face, Durlston Bay, which is in a Berriasian fluvial-lacustrine limestone in the Lulworth Formation of the United Kingdom.|website=fossilworks.org}}

| Durlston Bay, Swanage

|Teeth and dentary

|A small heterodontosaurid

Owenodon

|O. hoggii{{Cite web|url=https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=163545|title=Type specimen: BMNH R2998. Its type locality is Durlston Bay, Swanage (DB105-107), which is in a Berriasian marine limestone in the Lulworth Formation of the United Kingdom.}}

|Durlston Bay, Swanage

|Partial dentary

|An iguanodontoid

Nuthetes

|N. destructor{{Cite web|url=https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=66817|title=Type specimen: DORCM G913, a mandible. Its type locality is Feather Quarry, Durlston Bay (DB102), which is in a Berriasian marine shale in the Lulworth Formation of the United Kingdom.}}

| Feather Quarry, Durlston Bay, Swanage

|Teeth, dentary fragment

|Small dromaeosaur

= Mammals =

class="wikitable" align="center" width="100%" ! colspan="7" align="center" \|Mammals reported from the Lulworth Formation
Genus !Species !Location !Stratigraphic position !Material !Notes !Images
Albionbaatar \|A. denisae \|Sunnydown Farm Quarry \| \| \|Albionbaataridae \|
Amblotherium \|A. pusillum, A. soricinum, A. nanum \|Mammal Bed \| \| \| \|
Bolodon \|B. minor, B. crassidens, B. osborni, B. elongatus, B. falconeri \|Mammal Bed \| \| \|Plagiaulacidae \|
Chunnelodon \|C. alopekodes \|Sunnydown Farm Quarry \| \| \| \|
Ctenacodon \|C. minor \|Mammal Bed \| \| \|Allodontidae \|
Dorsetodon \|D. haysomi \|Sunnydown Farm Quarry \| \| \|Paurodontidae \|
Durlstotherium \|D. newmani \|Mammal Bed \| \| \| \|
Durlstodon \|D. ensomi \|Mammal Bed \| \| \| \|
Gerhardodon \|G. purbeckensis \|Sunnydown Farm Quarry \| \| \|Pinheirodontidae \|
Kouriogenys \|K. minor \|Mammal Bed \| \| \|Peramuridae \|
Magnimus \|M. ensomi \|Sunnydown Farm Quarry \| \| \| \|
Peraiocynodon \|P. inexpectatus \|Mammal Bed \| \| \|Docodontidae \|
Peramuroides \|P. tenuiscus \|Mammal Bed \| \| \|Peramuridae \|
Peramus \|P. tenuirostris, P. dubius, P. minor \|Mammal Bed \| \| \|Peramuridae \|
Peraspalax \|P. talpoides \|Mammal Bed, Sunnydown Farm Quarry \| \| \|Dryolestidae \|
Phascolestes \|P. mustelula \|Mammal Bed, Sunnydown Farm Quarry \| \| \|Dryolestidae \|
Plagiaulax \|P. becklesii \|Mammal Bed \| \| \|Plagiaulacidae \|
Purbeckodon \|P. batei \|Sunnydown Farm Quarry \| \| \|Morganucodonta \|
Spalacotherium \|S. tricuspidens, S. evansae, S. hookeri \|Mammal Bed, Sunnydown Farm Quarry \| \| \|Spalacotheriidae \|
Sunnyodon \|S. notleyi \|Sunnydown Farm Quarry \| \| \|Paulchoffatiidae \|
Thereuodon \|T. taraktes \|Sunnydown Farm Quarry \| \| \| \|
Tinodon \|T. micron \|Sunnydown Farm Quarry \| \| \|Tinodontidae \|
Triconodon \|T. mordax \|Mammal Bed, Sunnydown Farm Quarry \| \| \| \|
Trioracodon \|T. bisulcus, T. major, T. oweni \|Mammal Bed, Feather Quarry \| \| \| \|

class="wikitable" align="center" width="100%"

! colspan="7" align="center" |Mammals reported from the Lulworth Formation

Genus

!Species

!Location

!Stratigraphic position

!Material

!Notes

!Images

Albionbaatar

|A. denisae

|Sunnydown Farm Quarry