Lupulella

{{Short description|Genus of jackals native to Africa}}

{{Use dmy dates|date=January 2024}}

{{Automatic taxobox

| fossil_range = Pliocene - recent

| image = Lupulella.jpg

| image_caption = Side-striped and black-backed jackals

| taxon = Lupulella

| display_parents = 4

| authority = Hilzheimer, 1906

| type_species = Canis mesomelas{{cite journal |last1= Geraads |first1= D. |date= March 2011 |title= A revision of the fossil Canidae (Mammalia) of north-western Africa |journal= Palaeontology |volume= 54 |issue= 2 |pages= 429–446 |doi= 10.1111/j.1475-4983.2011.01039.x}}

| type_species_authority = Schreber, 1775

| subdivision_ranks = Species

| subdivision = *Lupulella adusta

| range_map = Lupulella range.png

| range_map_caption = Lupulella range
{{color box|#222AB6}} Lupulella adusta
{{color box|#EC2525}} Lupulella mesomelas

}}

Lupulella is a genus of canine found in Africa. This genus consists of only two extant species, the black-backed jackal (Lupulella mesomelas) and the side-striped jackal (Lupulella adusta).

Taxonomy

The two species had previously been considered members of the genus Canis. In 2017, a taxonomic review recommended that these two species be recognised as genus Lupulella. In response to this review, the American Society of Mammalogists recognised the new genus.

In 2019, a workshop hosted by the IUCN/SSC Canid Specialist Group recommends that because DNA evidence shows the side-striped jackal (Canis adustus) and black-backed jackal (Canis mesomelas) to form a monophyletic lineage that sits outside of the Canis/Cuon/Lycaon clade, that they should be placed in a distinct genus, Lupulella (Hilzheimer, 1906) with the names Lupulella adusta and Lupulella mesomelas.

{{Species table |genus=Lupulella |authority-name= Schreber |authority-year=1775 |species-count=two|no-note=y|narrow-percent=75}}

{{Species table/row

|name= Side-striped jackal |binomial=Lupulella adusta

|image=File:CANIS ADUSTUS dans le park zoologique de kinshasa (cropped).jpg |image-size=180px |image-alt=

|authority-name=Sundevall |authority-year=1847 |authority-not-original=yes

|range= central and southern Africa

|range-image=File:Side-striped Jackal area.png

|range-image-size=180px

|size=

|habitat=

|hunting=

|iucn-status= LC

|population=

|direction=

|subspecies={{Collapsible list |expand=yes |title=Seven subspecies |bullets=on

|L. a. adusta

|L. a. bweha

|L. a. centralis

|L. a. grayi

|L. a. kaffensis

|L. a. lateralis

|L. a. notatus

}}

}}

{{Species table/row

|name= Black-backed jackal |binomial=Lupulella mesomelas

|image=File:Black-backed Jackal (Lupulella mesomelas) (51995687394), crop.jpg |image-size=180px |image-alt=

|authority-name=Schreber |authority-year=1775 |authority-not-original=yes

|range= eastern and southern Africa

|range-image=File:Black-backed Jackal area.png

|range-image-size=180px

|size=

|habitat=

|hunting=

|iucn-status= LC

|population=

|direction=

|subspecies={{Collapsible list |expand=yes |title=Two subspecies |bullets=on

|Cape black-backed jackal, L. m. mesomelas

|East African black-backed jackal, L. m. schmidti

}}

}}

{{Species table/end}}

Phylogeny

Cladogram based on genomic data:K. Lindblad-Toh, C. M. Wade, T. S. Mikkelsen, E. K. Karlsson, D. B. Jaffe, M. Kamal, M. Clamp, J. L. Chang, E. J. Kulbokas, M. C. Zody, E. Mauceli, X. Xie, M. Breen, R. K. Wayne, E. A. Ostrander, C. P. Ponting, F. Galibert, D. R. Smith, P. J. Dejong, E. Kirkness, P. Alvarez, T. Biagi, W. Brockman, J. Butler, C. W. Chin, A. Cook, J. Cuff, M. J. Daly, D. Decaprio e S. Gnerre, Genome sequence, comparative analysis and haplotype structure of the domestic dog, in Nature, vol. 438, nº 7069, 2005, pp. 803–819, Bibcode:2005Natur.438..803L, DOI:10.1038/nature04338, {{PMID|16341006}}.

{{clade| style=font-size:100%;line-height:85%

|label1=True dogs

|1={{clade

|label1=

|1=Cerdocyonina50px

|label2=Wolf-like canids

|2={{clade

|label1=

|1=Canis/Cuon/Lycaon50px

|label2=Lupulella

|2={{clade

|label1=

|1=Side-striped jackal50px

|label2=

|2=Black-backed jackal50px

}} }} }} }}

References

{{Reflist|refs=

{{cite web |first1=Francisco|last1=Alvares|first2=Wieslaw|last2=Bogdanowicz|first3=Liz A.D.|last3=Campbell|first4=Rachel|last4=Godinho|first5=Jennifer|last5=Hatlauf|first6=Yadvendradev V.|last6=Jhala|first7=Andrew C.|last7=Kitchener|first8=Klaus-Peter|last8=Koepfli|first9=Miha|last9=Krofel|first10=Patricia D.|last10=Moehlman|first11=Helen|last11=Senn |first12=Claudio|last12=Sillero-Zubiri|first13=Suvi|last13=Viranta|first14=Geraldine|last14=Werhahn|year=2019|website=IUCN/SSC Canid Specialist Group|url=https://www.canids.org/CBC/Old_World_Canis_Taxonomy_Workshop.pdf|title=Old World Canis spp. with taxonomic ambiguity: Workshop conclusions and recommendations. CIBIO. Vairão, Portugal, 28th - 30th May 2019|access-date=6 March 2020}}

{{cite book|last1= Castelló|first1=José R.|title=Canids of the World: Wolves, Wild Dogs, Foxes, Jackals, Coyotes, and Their Relatives (Princeton Field Guides)|publisher=Princeton University Press|year=2018|chapter=Ch2-Wolf-like Canids|pages=160–165|isbn=978-0691176857|url={{Google books|plainurl=yes|id=islgDwAAQBAJ|page=160}}}}

{{cite web|title=Mammal Diversity Database|publisher=American Society of Mammalogists|url=https://www.mammaldiversity.org/explore.html|access-date=28 December 2020}}

{{cite iucn |author=Hoffmann, M. |date=2014 |title=Canis adustus |volume=2014 |page=e.T3753A46254734 |doi=10.2305/IUCN.UK.2014-1.RLTS.T3753A46254734.en |access-date=12 November 2021}}

{{cite journal|doi=10.1186/s40850-017-0015-0|title=Rediscovering a forgotten canid species|journal=BMC Zoology|volume=2|year=2017|last1=Viranta|first1=Suvi|last2=Atickem|first2=Anagaw|last3=Werdelin|first3=Lars|last4=Stenseth|first4=Nils Chr.|url=http://nrm.diva-portal.org/smash/get/diva2:1089580/FULLTEXT01|doi-access=free|hdl=10852/55258|hdl-access=free}}

}}