Manis

{{Automatic taxobox

| name = Asiatic pangolin

| fossil_range = {{fossil_range|12.9|0}} Middle Miocene - presentPhilippe Gaubert, Agostinho Antunes, Hao Meng, Lin Miao, Stéphane Peigné, Fabienne Justy, Flobert Njiokou, Sylvain Dufour, Emmanuel Danquah, Jayanthi Alahakoon, Erik Verheyen, William T Stanley, Stephen J O’Brien, Warren E Johnson, Shu-Jin Luo (2018) [https://academic.oup.com/jhered/article/109/4/347/4622594 "The Complete Phylogeny of Pangolins: Scaling Up Resources for the Molecular Tracing of the Most Trafficked Mammals on Earth"] Journal of Heredity, Volume 109, Issue 4, Pages 347–359

| image = Manis.jpg

| image_caption = Living pangolins from genus Manis

| taxon = Manis

| parent_authority = Gray, 1821

| authority = Linnaeus, 1758

| type_species = Manis pentadactyla

| type_species_authority = Linnaeus, 1758

| subdivision_ranks = Species

| subdivision =

| range_map = Manis ranges.png

| range_map_caption = {{leftlegend|#7C1797|Manis crassicaudata|outline=black}}

| synonyms = {{collapsible list |bullets=true |title=synonyms of subfamily:

|Mania

|Manidae {{small|(Gray, 1821)}}

|Manina {{small|(Gray, 1825)}}J. E. Gray. (1825.) "An outline of an attempt at the disposition of Mammalia into Tribes and Families, with a list of genera apparently appertaining to each Tribe." Annals of Philosophy, new series 10:337-344

|Pholidotina {{small|(Gray, 1873)}}

}}

{{collapsible list |bullets=true |title=synonyms of genus:

|Pangolin {{small|(Gray, 1873)}}

|Pangolinus {{small|(Rafinesque, 1821)}}

|Paramanis {{small|(Pocock, 1924)}}

|Phatages {{small|(Sundevall, 1843)}}

|Phatagenus {{small|(Sundevall, 1843)}}

|Pholidotus {{small|(Brisson, 1762)}}

|Quaggelo {{small|(Frisch, 1775)}}

}}

| synonyms_ref = {{Cite book |url=https://archive.org/details/indexgenerummam00palmgoog |page=[https://archive.org/details/indexgenerummam00palmgoog/page/n827 822] |title=Index Generum Mammalium: A List of the Genera and Families of Mammals |last=Palmer |first=Theodore Sherman |date=1904 |publisher=U. S. Government Printing Office |language=en}}{{Cite web |url=https://www.mv.helsinki.fi/home/mhaaramo/metazoa/deuterostoma/chordata/synapsida/eutheria/leptictida/history_Manis.html |title=Taxonomic history of the genus Manis |website=Mikko's Phylogeny Archive |access-date=2019-09-18}}

}}

Manis ("spirit") is a genus of South Asian and East Asian pangolins, the Asiatic pangolins, from subfamily Maninae, within family Manidae.{{Catalogue of Life|d36852dde74a91d7c965591b175947ec}}{{MSW3 Pholidota |id=13900003 |pages=530-531}}

Taxonomy

Subfamily: Maninae (Asiatic pangolins)
Genus: Manis (Asiatic pangolins)
Manis crassicaudata (Indian pangolin)
Manis pentadactyla (Chinese pangolin)
Manis aurita{{cite journal|vauthors=Zijlstra JS|title=Manis aurita Hodgson 1837 as a valid species of pangolin: a comment to Wangmo et al. (2025)|journal=Mammalian Biology|date=25 April 2025|doi=10.1007/s42991-025-00495-x}} (= Manis indoburmanica)Wangmo, L. K., Ghosh, A., Dolker, S., Joshi, B. D., Sharma, L. K., & Thakur, M. (2025). [https://link.springer.com/article/10.1007/s42991-024-00475-7 Indo-Burmese pangolin (Manis indoburmanica): a novel phylogenetic species of pangolin evolved in Asia.] Mammalian Biology, 1-8.
Manis mysteria (Cryptic pangolin)https://www.sci.news/biology/manis-mysteria-12316.html
†Manis hungarica
†Manis lydekkeri
Subgenus: Paramanis
Manis culionensis (Philippine pangolin)
Manis javanica (Sunda pangolin)
†Manis palaeojavanica (Giant asian pangolin)

Phylogeny

Phylogenetic position of genus Manis within family Manidae.{{cite journal |last=Gaudin |first=Timothy |year=2009 |title=The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis |journal=Journal of Mammalian Evolution |publisher=Springer Science+Business Media |location=Heidelberg, Germany |volume=16 |issue=4 |pages=235–305 |doi=10.1007/s10914-009-9119-9 |s2cid=1773698 |url=http://web2.utc.edu/~gvv824/Gaudin%20et%20al%202009.pdf |access-date=2020-11-19 |archive-date=2015-09-25 |archive-url=https://web.archive.org/web/20150925134257/http://web2.utc.edu/~gvv824/Gaudin%20et%20al%202009.pdf |url-status=dead }}{{cite journal |last1=Kondrashov |first1=Peter |last2=Agadjanian |first2=Alexandre K. |title=A nearly complete skeleton of Ernanodon (Mammalia, Palaeanodonta) from Mongolia: morphofunctional analysis |journal=Journal of Vertebrate Paleontology |volume=32 |issue=5 |year=2012 |issn=0272-4634 |doi=10.1080/02724634.2012.694319 |pages=983–1001|bibcode=2012JVPal..32..983K |s2cid=86059673 }}

{{clade |style=font-size:85%;line-height:100%;

|label1=Pholidotamorpha|sublabel1={{small|(Pholidota sensu lato)}}

|1={{clade

|1=†Palaeanodonta 60px

|label2=Pholidota|sublabel2=sensu stricto

|2={{clade

|thickness2=2

|1=†Euromanis

|label2=?|state2=dotted

|2=†Pholidota sp. (BC 16’08)

|3={{clade

|1=†Eurotamanduidae