Nesochthamalus
{{Short description|Genus of barnacles}}
{{Speciesbox
| genus = Nesochthamalus
| species = intertextus
| image_alt = Nesochthamalus intertextus
| image_caption = Nesochthamalus intertextus
| image = Nesochthamalus_intertextus_(10.3897-zookeys.945.39044)_Figure_9.jpg
| parent_authority = {{nowrap|Foster & Newman, 1987}}{{rp|326}}
| authority = (Darwin, 1854){{cite book|last=Darwin|first=Charles|title=A monograph on the sub-class Cirripedia with figures of all species. The Balanidae, the Verrucidae, etc.|url=https://archive.org/details/monographonsubcl02darw|year=1854|publisher=Ray Society|location=London|pages=[https://archive.org/details/monographonsubcl02darw/page/1 1]–684}}{{rp|[https://www.biodiversitylibrary.org/page/2006876 467]}}
| synonyms = Chthamalus intertextus Darwin, 1854
}}
The barnacle genus Nesochthamalus was erected by Foster & Newman, 1987,{{cite journal |author1=Brian A. Foster |author2=William A. Newman |name-list-style=amp |year=1987 |title=Chthamalid Barnacles of Easter Island. Peripheral Pacific Isolation of Notochthamalinae new-subfamily and Hembeli group of Euraphiinae (Cirripedia: Chthamaloidea). |journal=Bulletin of Marine Science |volume=41 |issue=2 |pages=322–336 |url=http://www.ingentaconnect.com/content/umrsmas/bullmar/1987/00000041/00000002/art00020}} to include sole species Chthamalus intertextus originally named by Darwin in 1854. It is widespread on islands in Western Pacific Ocean, including Hawaii, and presents combinations of unusual features which make easily recognizable for field workers. These include dirty white shell exterior with deep purple colored interior, operculars colored purple. Opercular plates on each side calcify together in all but youngest individuals, and cannot be separated or easily distinguished from each other. This feature is shared only by Rehderella belyaevi, but in latter species, scutum and tergum can be distinguished by raised ridge replacing old articular margin. Unique feature of Nesochthamalus is its basis. In young individuals, it is entirely membraneous, and with age, becomes secondarily calcareous progressively inwards, leaving only the center membraneous. As the basis calcifies, it rises off the substrate forming a saucer shape when viewed from the side. In addition, interior of shell is secondarily calcified.{{rp|75}}{{rp|[https://www.biodiversitylibrary.org/page/34961111 31]}}
Diagnosis and Discussion
Nesochthamalus intertextus is notable and distinctive in its outer shell and basis structure. General shape is flattened, with relatively large oval aperture. Crowded colonies do not become columnar, as in Chamaesipho. Unworn specimens are dirty white, and show vertical ribbing, prominent growth ridges, and wavy or chevroned sutures between shell plates. Because of its intertidal habitat, most specimens are eroded, particularly in their upper, older portions. As wear progresses, plate sutures become straighter. Interior of shell is colored deep violet, more intensely at plate sutures. This color only shows on exterior on unworn sections between ribs. Interior surface of shell plates is covered in small pits.{{rp|[https://www.biodiversitylibrary.org/page/34961111 31]}}{{rp|75}}
In young specimens, basis is entirely membraneous. As the animal grows, basis calcifies progressively inwards, leaving only central portion membraneous. As this happens, layers of secondary calcareous material form on inside of shell. This secondary calcification and basis are also colored purple, and serve to halt further enlargement of shell size. As calcification proceeds, outer parts of the animal are raised off substrate, showing convex, saucer-shaped side view.{{rp|[https://www.biodiversitylibrary.org/page/34961111 31]}}
Opercular plates are unusual in being completely fused on each side, with no trace left of scutal/tergal articulation, except in youngest individuals. Where scutum and tergum are separable, they are equal in size, and colored deep violet, with tergum darker than scutum.{{rp|75}} Tergum shows clear spur, and 4 tergal depressor muscle crests. Otherwise, these plates are nearly featureless, except for numerous pits on inside surface.{{rp|[https://www.biodiversitylibrary.org/page/34961113 33]}}
= Synonymy and Nomenclature =
== ''Nesochthamalus'' ==
- Nesochthamalus {{aut|Foster & Newman}}, 1987: 326, (Original Description): {{aut|Poltarukha}}, 2006: 75,{{cite book|last=Poltarukha|first=O. P.|title=Identification Atlas of Superfamily Chthamaloidea (Cirripedia Thoracica) barnacles in World Ocean|year=2006|publisher=KMK Scientific Press, Ltd.|location=Moscow|isbn=978-5-87317-278-8|pages=1–198 [In Russian]}} (discussion).
- Type species: Chthamalus intertextus {{aut|Darwin}}, 1854: 467, by original designation.
== ''Nesochthamalus intertextus'' ==
- Chthamalus intertextus {{aut|Darwin}}, 1854: [https://www.biodiversitylibrary.org/page/2006876 467] (original description); {{aut|Pope}}, 1965: [https://www.biodiversitylibrary.org/page/34961109 29];{{cite journal|last=Pope|first=E. C.|title=A review of Australian and some Indomalayan Chthamalidae (Crustacea, Cirripedia)|journal=Proc. Linn. Soc. N. S. W.|year=1965|volume=90|pages=10–76}} (extensive re-description, discussion).
- Nesochthamalus intertextus ({{aut|Darwin}}). {{aut|Foster & Newman}}, 1987: 326 (generic reassignment); {{Aut|Poltarukha}}, 1996: 993;{{cite journal|last=Poltarukha|first=O. P.|title=Composition, phylogeny and position in system of subfamily Notochthamalinae (Crustacea, Chthamalidae)|journal=Zoologicheskii Zhurnal|year=1996|volume=75|issue=7|pages=985–994|issn=0044-5134}} 2006: 76; 2008: 75;{{cite book|author1=Poltarukha, O |author2=A. Yu. Zvyagintsev |title=Barnacles (Cirripedia Thoracica) of Vietnam and their role in the fouling communities|year=2008|publisher=KMK Scientific Press, Ltd.|location=Moscow|isbn=978-5-87317-453-9|page=335 p. [In Russian]}} (discussions, supplementary descriptions, habitat and range data).
- Euraphia intertexta ({{aut|Darwin}}). {{aut|Newman & Ross}}, 1976: [https://www.biodiversitylibrary.org/page/4328584 41];{{cite journal|last=Newman|first=W. A.|author2=A. Ross|title=Revision of the Balanomorph Barnacles including a catalog of the species|journal=Memoirs of San Diego Society of Natural History|year=1976|volume=9|pages=1–108}} (complete reference list to 1976): {{aut|Zevina et al.}}, 1992: 79;{{cite book|author=Zevina, G. B.|author2=A. Yu. Zvyagintsev|author3=C. E. Negashev|name-list-style=amp|title=Barnacles of coast of Vietnam and their role in fouling|year=1992|publisher=DVO AN CCCP|location=Vladivostok|page=144 p. [in Russian]}} (junior synonym).
- Type locality: "Philippine Archipelago; Mus. Cuming", as stated by Darwin's description, and verified by Pope, 1965:[https://www.biodiversitylibrary.org/page/34961115 35] as type locality. In the 19th century, precise type localities were not given. Darwin did not give broad geographic ranges, and usually general locality given at beginning of species descriptions can be taken as type locality.
- Type specimens and repository: Pope, 1965{{rp|[https://www.biodiversitylibrary.org/page/34961114 34]}} reported a group of specimens in British Museum (Natural History) labeled "Philippines - ex Museum Cuming" as probably type material. No holotype or lectotype has been selected.
Habitat and Geographical Distribution
Nesochthamalus can be found in mid-littoral environments, and does not generally form densely packed colonies as are characteristic of Chamaesipho.
Nesochthamalus intertextus is widespread on islands in West and Central Pacific Ocean. Southernmost reported occurrence is Port Moresby, New Guinea, and is absent from Australia. Other reports include Indonesia, Philippine Archipelago (type area), Taiwan, Japan, Okinawa, Ryukyu Islands, Fiji, Tuamotu, and Hawaii. It may be reasonably expected to be found throughout Micronesia and Melanesia.[https://www.biodiversitylibrary.org/page/34961115 35] To this list can be added Vietnam.
References
{{Reflist|32em}}
External links
{{Wikispecies-inline}}
- [http://marinespecies.org/aphia.php?p=taxdetails&id=733082 WoRMS]
- [https://web.archive.org/web/20140413130627/http://barnacle.biota.biodiv.tw/pages/1366 Taiwan barnacles]
{{Taxonbar|from=Q15731709}}