Ohyamashimo Formation

{{short description|Geologic formation in Japan}}

{{Infobox rockunit

| name = Ohyamashimo Formation

| image =

| caption =

| type = Geological formation

| age = Albian
~{{fossil range|112.1|106.4}}

| period = Albian

| prilithology = Conglomerate, Sandstone, Mudstone

| otherlithology = Tuff

| namedfor = Ohyamashimo, Tamba-Sasayama

| namedby = Hayashi et al., 2017{{Cite journal |last1=Hayashi |first1=Keiichi |last2=Fujita |first2=Saki |last3=Koarai |first3=Kazuto |last4=Matsukawa |first4=Masaki |date=2017-09-15 |title=Stratigraphy and paleoenvironment of the Cretaceous Sasayama Group in the Sasayama area, Hyogo Prefecture, western Japan |journal=The Journal of the Geological Society of Japan |language=en |volume=123 |issue=9 |pages=747–764 |doi=10.5575/geosoc.2017.0016 |issn=0016-7630|doi-access=free }}

| region = Hyōgo Prefecture

| country = Japan

| coordinates = {{coord|35.1|N|135.1|E|display=inline,title}}

| paleocoordinates = {{coord|47.9|N|132.8|E|display=inline}}

| unitof = Sasayama Group

| subunits =

| underlies = Sawada Formation

| overlies = Basement (Unconformity)

| thickness =1300 metres

| extent =

| area =

| map = {{Location map+ | Japan

| relief = 1

| width = 250

| float = center

| places =

{{Location map~ | Japan

| lat_deg = 35.1

| lon_deg = 135.1

| mark = Green pog.svg

| marksize = 12

}}

}}

| map_caption =

}}

The Ohyamashimo Formation(大山下層) is an Early Cretaceous (Albian) geologic formation in Japan. It has been dated to the early-mid Albian, between 112.1 ± 0.4 Ma and 106.4 ± 0.4 Ma. Dinosaur remains have been discovered from this formation, including the sauropod Tambatitanis, the theropod Hypnovenator and the ceratopsian Sasayamagnomus. Other representative fossils from the formation include the monstersaurian lizard Morohasaurus and eutherian mammal Sasayamamylos. The depositional environment represents a fluvial system with a subhumid to semi-arid climate. The Ohyamashimo Formation was called the 'Lower Formation' of the Sasayama Group before it was named in 2017.

Fossil content

{{Paleobiota-key-compact}}

= Dinosaurs =

== Ankylosaurs ==

class="wikitable" align="center"
colspan="7" align="center" | Ankylosaurs of the Ohyamashimo Formation
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Ankylosauria Indet.

| Indeterminate

| Tamba-Sasayama

| Albian

| Dental remains.

|

|

== Ceratopsians ==

class="wikitable" align="center"
colspan="7" align="center" | Ceratopsians of the Ohyamashimo Formation
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Sasayamagnomus{{cite journal |last1=Tanaka |first1=Tomonori |last2=Chiba |first2=Kentaro |last3=Ikeda |first3=Tadahiro |last4=Ryan |first4=Michael J. |title=A new neoceratopsian (Ornithischia, Ceratopsia) from the Lower Cretaceous Ohyamashimo Formation (Albian), southwestern Japan |journal=Papers in Palaeontology |date=September 2024 |volume=10 |issue=5 |doi=10.1002/spp2.1587}}

| S. saegusai

| Tamba-Sasayama

| Albian

| Skull elements and partial postcranial remains.

| A neoceratopcian; closely related to Aquilops americanus.

| File:Sasayamagnomus_saegusai.png

== Ornithopods ==

class="wikitable" align="center"
colspan="7" align="center" | Ornithopods of the Ohyamashimo Formation
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Iguanodontia Indet.

| Indeterminate

| Tamba-Sasayama

| Albian

| Dental remains.

|

|

== Sauropods ==

class="wikitable" align="center"
colspan="7" align="center" | Sauropods of the Ohyamashimo Formation
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Sauropoda Indet.

| Indeterminate

| Tamba-Sasayama

| Albia

|

|

|

Tambatitanis{{cite journal |last1=Saegusa |first1=Haruo |last2=Ikeda |first2=Tadahiro |title=A new titanosauriform sauropod (Dinosauria: Saurischia) from the Lower Cretaceous of Hyogo, Japan |journal=Zootaxa |date=12 August 2014 |volume=3848 |issue=1 |doi=10.11646/zootaxa.3848.1.1}}

| T. amicitiae

| Tamba-Sasayama

| Albian

| A single partial skeleton.

| A titanosauriform sauropod

| File:Tambatitanis Palaeotaku 24.png

== Theropods ==

class="wikitable" align="center"
colspan="7" align="center" | Theropods of the Ohyamashimo Formation
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Dromaeosauridae Indet.

| Indeterminate

| Tamba-Sasayama

| Albian

| Dental remains.

|

|

Hypnovenator{{cite journal |last1=Kubota |first1=Katsuhiro |last2=Kobayashi |first2=Yoshitsugu |last3=Ikeda |first3=Tadahiro |title=Early Cretaceous troodontine troodontid (Dinosauria: Theropoda) from the Ohyamashimo Formation of Japan reveals the early evolution of Troodontinae |journal=Scientific Reports |date=25 July 2024 |volume=14 |issue=1 |doi=10.1038/s41598-024-66815-2|pmc=11272788 }}

| H. matsubaraetoheorum

| Tamba-Sasayama

| Albian

| A single partial skeleton.

| A troodontid theropod; closely related to Gobivenator mongoliensis.

| File:Hypnovenator matsubaraetoheorum.png

Therizinosauria Indet.

| Indeterminate

| Tamba-Sasayama

| Albian

| Dental remains.

|

|

Tyrannosauroidea Indet.

| Indeterminate

| Tamba-Sasayama

| Albian

| Dental remains.

|

|

= Squamates =

class="wikitable" align="center"
colspan="7" align="center" | Squamtes of the Ohyamashimo Formation
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Morohasaurus{{cite journal |last1=Ikeda |first1=Tadahiro |last2=Ota |first2=Hidetoshi |last3=Tanaka |first3=Tomonori |last4=Ikuno |first4=Kenji |last5=Kubota |first5=Katsuhiro |last6=Tanaka |first6=Kohei |last7=Saegusa |first7=Haruo |title=A fossil Monstersauria (Squamata: Anguimorpha) from the Lower Cretaceous Ohyamashimo Formation of the Sasayama Group in Tamba City, Hyogo Prefecture, Japan |journal=Cretaceous Research |date=February 2022 |volume=130 |pages=105063 |doi=10.1016/j.cretres.2021.105063|doi-access=free }}

| M. kamitakiensis

| Tamba-Sasayama

| Albian

| A nearly complete left dentary.

| A monstersaurian lizard

|

Pachygenys{{cite journal |last1=Ikeda |first1=Tadahiro |last2=Ota |first2=Hidetoshi |last3=Saegusa |first3=Haruo |title=A new fossil lizard from the Lower Cretaceous Sasayama Group of Hyogo Prefecture, western Honshu, Japan |journal=Journal of Vertebrate Paleontology |date=2 January 2015 |volume=35 |issue=1 |pages=e885032 |doi=10.1080/02724634.2014.885032}}

| P. adachii

| Tamba-Sasayama

| Albian

| A single, partially broken right mandible

| A scincomorphan lizard

|

= Amphibians =

class="wikitable" align="center"
colspan="7" align="center" | Amphibian of the Ohyamashimo Formation{{cite journal |last1=Ikeda |first1=Tadahiro |last2=Ota |first2=Hidetoshi |last3=Matsui |first3=Masafumi |title=New fossil anurans from the Lower Cretaceous Sasayama Group of Hyogo Prefecture, Western Honshu, Japan |journal=Cretaceous Research |date=June 2016 |volume=61 |pages=108–123 |doi=10.1016/j.cretres.2015.12.024}}
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Hyogobatrachus

| H. wadai

| Tamba-Sasayama

| Albian

| A nearly complete skeleton.

| A frog

|

Tambabatrachus

| T. kawazu

| Tamba-Sasayama

| Albian

| An articulated skeleton lacking the anterior part of the skull, most of the pectoral girdle, manus, and feet.

| A frog

|

= Mammals =

class="wikitable" align="center"
colspan="7" align="center" | Mammals of the Ohyamashimo Formation
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Sasayamamylos{{cite journal |last1=Kusuhashi |first1=Nao |last2=Tsutsumi |first2=Yukiyasu |last3=Saegusa |first3=Haruo |last4=Horie |first4=Kenji |last5=Ikeda |first5=Tadahiro |last6=Yokoyama |first6=Kazumi |last7=Shiraishi |first7=Kazuyuki |title=A new Early Cretaceous eutherian mammal from the Sasayama Group, Hyogo, Japan |journal=Proceedings of the Royal Society B: Biological Sciences |date=22 May 2013 |volume=280 |issue=1759 |pages=20130142 |doi=10.1098/rspb.2013.0142|pmc=3619506 }}

| S. kawaii

| Tamba-Sasayama

| Albian

| Three dentaries.

| A asioryctitherian eutherian

|

= Crustaceans =

class="wikitable" align="center"
colspan="7" align="center" | Crustaceans of the Ohyamashimo Formation
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Mongolocypris

| M. sp.

| Tamba-Sasayama

| Albian

|

| A ostracod

|

Conchostraca Indet.

| Indeterminate

| Tamba-Sasayama

| Albian

|

| A clam shrimp

|

= Molluscs =

== Bivalves ==

class="wikitable" align="center"
colspan="7" align="center" | Bivalves of the Ohyamashimo Formation
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Sphaerium

| S. coreanicum

| Tamba-Sasayama

| Albian

|

| A sphaeriid bivalve

|

== Gastropods ==

class="wikitable" align="center"
colspan="7" align="center" | Gastropods of the Ohyamashimo Formation
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Viviparus

| V. cf. keishoensis

| Tamba-Sasayama

| Albian

|

| A viviparid gastropod

|

= Oofossils =

class="wikitable" align="center"
colspan="7" align="center" | Eggs of the Ohyamasimo Formation{{cite journal |last1=Tanaka |first1=Kohei |last2=Zelenitsky |first2=Darla K. |last3=Saegusa |first3=Haruo |last4=Ikeda |first4=Tadahiro |last5=DeBuhr |first5=Christopher L. |last6=Therrien |first6=François |title=Dinosaur eggshell assemblage from Japan reveals unknown diversity of small theropods |journal=Cretaceous Research |date=January 2016 |volume=57 |pages=350–363 |doi=10.1016/j.cretres.2015.06.002}}{{cite journal |last1=Tanaka |first1=Kohei |last2=Zelenitsky |first2=Darla K. |last3=Therrien |first3=François |last4=Ikeda |first4=Tadahiro |last5=Kubota |first5=Katsuhiro |last6=Saegusa |first6=Haruo |last7=Tanaka |first7=Tomonori |last8=Ikuno |first8=Kenji |title=Exceptionally small theropod eggs from the Lower Cretaceous Ohyamashimo Formation of Tamba, Hyogo Prefecture, Japan |journal=Cretaceous Research |date=October 2020 |volume=114 |pages=104519 |doi=10.1016/j.cretres.2020.104519}}
Genus

! Species

! Location

! Stratigraphic position

! Material

! Notes

! Images

Elongatoolithus

| E. sp.

|

|

|

|

|

Himeoolithus

| H. murakamii

|

|

|

|

|

Nipponoolithus

| N. ramosus

|

|

|

|

|

Prismatoolithus

| P. sp.

|

|

|

|

|

Subtiliolithus

| S. hyogoensis

|

|

|

|

|

References