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Pacullidae

{{Short description|Family of spiders}}

{{Automatic taxobox

| oldest_fossil = Cenomanian

| image =

| image_caption =

| taxon = Pacullidae

| authority = Simon, 1894

| range_map =

| subdivision_ranks =

| subdivision =

| diversity = 4 genera, 39 species

}}

Pacullidae is a family of araneomorph spiders first described by Eugène Simon in 1894.{{cite book| last=Simon| first=E.| year=1894| title=Histoire naturelle des araignées}} It was merged into Tetrablemmidae in 1958,{{cite journal| last=Bonnet| first=P.| year=1958| title=Bibliographia araneorum| journal=Douladoure, Toulouse| volume=2| issue=4| page=3297}}{{cite journal| last=Brignoli| first=P. M.| year=1973| title=Ragni della Melanesia, I. Un nuovo Tetrablemma di Guadalcanal (Isole Salomone) (Araneae Tetrablemmidae)| journal=Memorie della Società Entomologica Italiana, Genova| volume=72| page=86}} then raised back to family status after a large phylogenetic study in 2017.{{Citation |last1=Wheeler |first1=Ward C.| year=2016 |title=The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling |journal=Cladistics |volume=33|issue=6|pages=574–616|doi=10.1111/cla.12182 |pmid=34724759 |s2cid=35535038|display-authors=etal|doi-access=free}}

Description

The family Pacullidae contains three-clawed spiders with six eyes, lacking a cribellum. They resemble spiders from the family Tetrablemmidae in some respects but are much larger, always exceeding 5 mm long, have a very wrinkled (rugose) cuticle, and females do not have large membranous receptacles.

Phylogeny

Pacullidae falls within the Synspermiata clade, a clade of former haplogyne spiders with "synsperm" – encapsulated groups of 2–4 fused sperm cells. Within this clade, it groups with four other families, including Tetrablemmidae, but is distinct from the latter, being most closely related to Diguetidae. Together with Pholcidae, these four families are placed in the "lost trachea clade", a group of families that have lost their posterior respiratory system.

{{clade|style=line-height:100%

|label1="Lost trachea clade"

|1={{clade

|1={{clade

|1=Tetrablemmidae

|2={{clade

|1=Plectreuridae

|2={{clade

|1=Diguetidae

|2=Pacullidae

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|2=Pholcidae

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Genera and species

{{as of|2019|04}}, the World Spider Catalog accepts the following genera and species:{{cite web| title=Family: Pacullidae Simon, 1894| website=World Spider Catalog| access-date=2019-04-22| publisher=Natural History Museum Bern| url=http://www.wsc.nmbe.ch/family/121}}

References

{{Reflist}}

{{Araneae}}

{{Taxonbar|from=Q33134682}}

{{Authority control}}

Category:Araneomorphae families