Araneomorphae

{{short description|Infraorder of arachnids}}

{{Automatic taxobox

| fossil_range = {{Fossil range|Triassic|present}}

| name = Araneomorph spiders

| image = Nephila inaurata1.JPG

| image_caption = Nephila inaurata (Nephilidae)

| taxon = Araneomorphae

| authority =

| diversity_link = Spider families

| diversity = 95 families

| subdivision_ranks = Subdivisions

| subdivision_ref =

| subdivision = *Filistatidae

}}

The Araneomorphae (also called the Labidognatha or "true spiders"{{cite book |author=Coddington, J.A. |title=Spiders of North America: an identification manual |publisher=American Arachnological Society |year=2005 |isbn=978-0-9771439-0-0 |editor=Ubick, D. |pages=18–24 |chapter=Phylogeny and Classification of Spiders |access-date=12 October 2008 |editor2=Paquin, P. |editor3=Cushing, P.E. |editor4=Roth, V. |chapter-url=http://si-pddr.si.edu/dspace/bitstream/10088/4365/1/CoddingtonSNAPhylogeny05.pdf |archive-url=https://web.archive.org/web/20081209110443/http://si-pddr.si.edu/dspace/bitstream/10088/4365/1/CoddingtonSNAPhylogeny05.pdf |archive-date=9 December 2008 |url-status=dead}}) are an infraorder of spiders. They are distinguishable by chelicerae (fangs) that point diagonally forward and cross in a pinching action, in contrast to the Mygalomorphae (tarantulas and their close kin), where they point straight down. Araneomorphs comprise the vast majority (about 93%{{Cite web |last=Bern |first=Natural History Museum |title=NMBE - World Spider Catalog |url=http://research.amnh.org/entomology/spiders/catalog/INTRO1.html |access-date=2023-06-07 |website=research.amnh.org |language=en}}) of living spiders.

Distinguishing characteristics

Most spider species are Araneomorphae, which have fangs that face towards each other, increasing the orientations that they can employ during prey-capture. They have fewer book lungs (when present) – usually one pair – and the females typically live one year.

The Mygalomorphae have fangs that face towards the ground, and which are parallel to the long axis of the spider's body, thus they have only one orientation they can employ during prey capture. They have two pairs of book lungs, and the females often live many years.{{cite web |title=ABOUT SPIDERS |website=www.publish.csiro.au |url=http://www.publish.csiro.au/samples/spiders/html/SPIDERS.HTM |access-date=2022-01-07}}

Image:Cheiracanthium punctorium frei 1 17 Forst Jungfernhdeide Jg 46 070920.jpg| This Cheiracanthium punctorium shows the orientation of Araneomorphae fangs.

Image:Atrax robustus.jpg|This Atrax robustus shows the orientation of Mygalomorphae fangs.

Spiders included

Almost all of the familiar spiders are included in the Araneomorphae group, one major exception being the tarantulas. There are a few other Mygalomorphae species that live around homes or gardens, but they typically are relatively small and not easily noticed.

The Araneomorphae, to the contrary, include the weavers of spiral webs; the cobweb spiders that live in the corners of rooms, and between windows and screens; the crab spiders that lurk on the surfaces of flowers in gardens; the jumping spiders that are visible hunting on surfaces; the wolf spiders that carpet hunting sites in sunny spots; and the large huntsman spiders.

Systematics

In older schemes, the Araneomorphae were divided into two lineages, the Hypochilae (containing only the family Hypochilidae), and the Neocribellatae. The Neocribellatae were in turn divided into the Austrochiloidea, and the two series Haplogynae and Entelogynae, each containing several superfamilies. Molecular phylogenetic studies have shown that the haplogynes in particular are not a monophyletic group. A 2020 study suggested the relationships among the major groups were as shown in the following cladogram.{{cite journal |last1=Magalhaes |first1=Ivan L. F. |last2=Azevedo |first2=Guilherme H. F. |last3=Michalik |first3=Peter |last4=Ramírez |first4=Martín J. |date=February 2020 |title=The fossil record of spiders revisited: implications for calibrating trees and evidence for a major faunal turnover since the Mesozoic |language=en |journal=Biological Reviews |issn=1464-7931 |doi=10.1111/brv.12559 |pmid=31713947 |s2cid=207937170 |volume=95 |issue=1 |pages=184–217 |url=https://www.researchgate.net/publication/337190453 }}

{{clade |style=font-size:100%;line-height:80%

|grouplabel1={{clade labels |label1=Haplogynae |top1=30% |color1=blue}}

|label1=Araneomorphae

|1={{clade

|1={{clade

|1={{clade

|2=Filistatidae |bar2=blue

|1=Hypochilidae

}}

|2=Synspermiata |bar2=blue

}}

|2={{clade

|1={{clade

|1=Leptonetidae |bar1=blue

|2=Austrochiloidea

}}

|2={{clade

|1=Palpimanoidea

|2=Entelegynae

}}

}}

}}

}}

The blue bar to the right shows the former Haplogynae in the sense of Coddington (2005).

Table of families

class="wikitable"

|+Key

Generabgcolor="lightblue" | 1bgcolor="lightgreen" | ≥2bgcolor="#FFA" | ≥10bgcolor="pink" | ≥100
Speciesbgcolor="lightblue" | 1–9bgcolor="lightgreen" | ≥10bgcolor="#FFA" | ≥100bgcolor="pink" | ≥1000

class="wikitable"

|+Araneomorphae families{{refn|group=notes|name=Note1|Unless otherwise shown, currently accepted families and counts based on the World Spider Catalog version 25.5 {{as of|lc=yes|2025|February|7}}.{{Citation |mode=cs1 |title=Currently valid spider genera and species |work=World Spider Catalog |publisher=Natural History Museum Bern |url=https://www.wsc.nmbe.ch/statistics/ |access-date=2025-02-07}} Currently valid spider genera and species In the World Spider Catalog, "species" counts include subspecies. Assignment to sub- and infraorders based on {{Harvtxt|Coddington|2005|p=20}} (when given there).}}

Family||Genera||Species||Common name||Example
Agelenidaebgcolor="#FFA"|97bgcolor="pink"|1420araneomorph funnel-web spidersHobo spider (Eratigena agrestis)
Amaurobiidaebgcolor="#FFA"|26bgcolor="#FFA"|202tangled nest spidersCallobius claustrarius
Anapidaebgcolor="#FFA"|59bgcolor="#FFA"|233Holarchaea novaeseelandiae
Ancylometidaebgcolor="lightblue"|1bgcolor="lightgreen"|11
Anyphaenidaebgcolor="#FFA"|58bgcolor="#FFA"|649anyphaenid sac spidersHibana velox (yellow ghost spider)
Araneidaebgcolor="pink"|198bgcolor="pink"|3144orb-weaver spidersZygiella x-notata
Archaeidaebgcolor="lightgreen"|6bgcolor="lightgreen"|93pelican spidersMadagascarchaea gracilicollis
Archoleptonetidaebgcolor="lightgreen"|2bgcolor="lightblue"|8Archoleptoneta gertschi
Arkyidaebgcolor="lightgreen"|2bgcolor="lightgreen"|38
Austrochilidaebgcolor="lightgreen"|2bgcolor="lightblue"|9Tasmanian cave spider (Hickmania troglodytes)
Caponiidaebgcolor="#FFA"|21bgcolor="#FFA"|153Diploglena capensis
Cheiracanthiidaebgcolor="#FFA"|15bgcolor="#FFA"|376Cheiracanthium mildei
Cicurinidaebgcolor="lightgreen"|4bgcolor="#FFA"|172
Cithaeronidaebgcolor="lightgreen"|2bgcolor="lightblue"|9
Clubionidaebgcolor="#FFA"|18bgcolor="#FFA"|667sac spidersClubiona trivialis
Corinnidaebgcolor="#FFA"|76bgcolor="#FFA"|876dark sac spidersCastianeira sp.
Ctenidaebgcolor="#FFA"|48bgcolor="#FFA"|605wandering spidersPhoneutria fera
Cyatholipidaebgcolor="#FFA"|23bgcolor="lightgreen"|58
Cybaeidaebgcolor="#FFA"|23bgcolor="#FFA"|301Cryphoeca silvicola
Cycloctenidaebgcolor="lightgreen"|8bgcolor="lightgreen"|80
Deinopidaebgcolor="lightgreen"|3bgcolor="lightgreen"|68net-casting spidersAsianopis subrufa (rufous net-casting spider)
Desidaebgcolor="#FFA"|63bgcolor="#FFA"|323intertidal spidersPhryganoporus candidus
Dictynidaebgcolor="#FFA"|51bgcolor="#FFA"|460Nigma walckenaeri
Diguetidaebgcolor="lightgreen"|2bgcolor="lightgreen"|16coneweb spiders
Dolomedidaebgcolor="lightgreen"|7bgcolor="#FFA"|128
Drymusidaebgcolor="lightgreen"|2bgcolor="lightgreen"|19false violin spiders
Dysderidaebgcolor="#FFA"|24bgcolor="#FFA"|653woodlouse hunter spidersWoodlouse spider (Dysdera crocata)
Eresidaebgcolor="lightgreen"|9bgcolor="#FFA"|106velvet spidersEresus sandaliatus
Filistatidaebgcolor="#FFA"|18bgcolor="#FFA"|192crevice weaversSouthern house spider (Kukulcania hibernalis)
Fonteferreidaebgcolor="lightblue"|1bgcolor="lightblue"|1
Gallieniellidaebgcolor="lightgreen"|5bgcolor="lightgreen"|41
Gnaphosidaebgcolor="pink"|153bgcolor="pink"|2479flat-bellied ground spidersDrassodes cupreus
Gradungulidaebgcolor="lightgreen"|8bgcolor="lightgreen"|18large-clawed spidersProgradungula carraiensis (Carrai cave spider)
Hahniidaebgcolor="#FFA"|29bgcolor="#FFA"|240dwarf sheet spiders
Hersiliidaebgcolor="#FFA"|16bgcolor="#FFA"|187tree trunk spidersHersilia savignyi
Homalonychidaebgcolor="lightblue"|1bgcolor="lightblue"|2
Huttoniidaebgcolor="lightblue"|1bgcolor="lightblue"|1Huttonia palpimanoides
Hypochilidaebgcolor="lightgreen"|2bgcolor="lightgreen"|33lampshade spidersHypochilus thorelli
Lamponidaebgcolor="#FFA"|23bgcolor="#FFA"|192White-tailed spider (Lampona spp.)
Leptonetidaebgcolor="#FFA"|22bgcolor="#FFA"|397Tooth Cave spider (Tayshaneta myopica)
Linyphiidaebgcolor="pink"|640bgcolor="pink"|4940dwarf / money spidersLinyphia triangularis
Liocranidaebgcolor="#FFA"|35bgcolor="#FFA"|354liocranid sac spiders
Lycosidaebgcolor="pink"|135bgcolor="pink"|2490wolf spidersLycosa tarantula
Macrobunidaebgcolor="#FFA"|26bgcolor="lightgreen"|92
Malkaridaebgcolor="#FFA"|13bgcolor="lightgreen"|57shield spiders
Mecysmaucheniidaebgcolor="lightgreen"|7bgcolor="lightgreen"|25
Megadictynidaebgcolor="lightgreen"|2bgcolor="lightblue"|2
Mimetidaebgcolor="lightgreen"|8bgcolor="#FFA"|164pirate spidersOarces reticulatus
Miturgidaebgcolor="#FFA"|33bgcolor="#FFA"|191long-legged sac spiders
Myrmecicultoridaebgcolor="lightblue"|1bgcolor="lightblue"|1
Mysmenidaebgcolor="#FFA"|17bgcolor="#FFA"|188spurred orb-weavers
Nesticidaebgcolor="#FFA"|16bgcolor="#FFA"|292cave cobweb spidersNesticella marapu
Nicodamidaebgcolor="lightgreen"|7bgcolor="lightgreen"|27
Ochyroceratidaebgcolor="lightgreen"|9bgcolor="#FFA"|184midget ground weaversTheotima minutissima
Oecobiidaebgcolor="lightgreen"|7bgcolor="#FFA"|129disc web spidersOecobius navus
Oonopidaebgcolor="pink"|115bgcolor="pink"|1962dwarf hunting spidersOonops domesticus
Orsolobidaebgcolor="#FFA"|30bgcolor="#FFA"|189
Oxyopidaebgcolor="lightgreen"|9bgcolor="#FFA"|448lynx spidersPeucetia viridans (green lynx spider)
Pacullidaebgcolor="lightgreen"|4bgcolor="lightgreen"|38
Palpimanidaebgcolor="#FFA"|20bgcolor="#FFA"|182palp-footed spiders
Penestomidaebgcolor="lightblue"|1bgcolor="lightblue"|9
Periegopidaebgcolor="lightblue"|1bgcolor="lightblue"|3
Philodromidaebgcolor="#FFA"|30bgcolor="#FFA"|527philodromid crab spidersPhilodromus dispar
Pholcidaebgcolor="#FFA"|97bgcolor="pink"|2029daddy long-legs spidersPholcus phalangioides
Phrurolithidaebgcolor="#FFA"|25bgcolor="#FFA"|410
Physoglenidaebgcolor="#FFA"|13bgcolor="lightgreen"|72
Phyxelididaebgcolor="#FFA"|14bgcolor="lightgreen"|68
Pimoidaebgcolor="lightgreen"|2bgcolor="lightgreen"|87Pimoa cthulhu
Pisauridaebgcolor="#FFA"|45bgcolor="#FFA"|236nursery web spidersPisaura mirabilis
Plectreuridaebgcolor="lightgreen"|2bgcolor="lightgreen"|32
Prodidomidaebgcolor="#FFA"|24bgcolor="#FFA"|195
Psechridaebgcolor="lightgreen"|2bgcolor="lightgreen"|62
Psilodercidaebgcolor="#FFA"|11bgcolor="#FFA"|224
Salticidaebgcolor="pink"|689bgcolor="pink"|6808jumping spidersZebra spider (Salticus scenicus)
Scytodidaebgcolor="lightgreen"|4bgcolor="#FFA"|253spitting spidersScytodes thoracica
Segestriidaebgcolor="lightgreen"|5bgcolor="#FFA"|181tubeweb spidersSegestria florentina
Selenopidaebgcolor="lightgreen"|9bgcolor="#FFA"|282wall spidersSelenops radiatus
Senoculidaebgcolor="lightblue"|1bgcolor="lightgreen"|31
Sicariidaebgcolor="lightgreen"|3bgcolor="#FFA"|176recluse spidersBrown recluse (Loxosceles reclusa)
Sparassidaebgcolor="#FFA"|97bgcolor="pink"|1519huntsman spidersDelena cancerides (Avondale spider)
Stenochilidaebgcolor="lightgreen"|2bgcolor="lightgreen"|13
Stiphidiidaebgcolor="#FFA"|20bgcolor="#FFA"|125Tartarus mullamullangensis
Symphytognathidaebgcolor="#FFA"|10bgcolor="#FFA"|104dwarf orb-weaversPatu digua
Synaphridaebgcolor="lightgreen"|3bgcolor="lightgreen"|13
Synotaxidaebgcolor="lightgreen"|5bgcolor="lightgreen"|38
Telemidaebgcolor="#FFA"|16bgcolor="#FFA"|107long-legged cave spiders
Tetrablemmidaebgcolor="#FFA"|27bgcolor="#FFA"|153armored spiders
Tetragnathidaebgcolor="#FFA"|45bgcolor="#FFA"|989long jawed orb-weaversLeucauge venusta (orchard spider)
Theridiidaebgcolor="pink"|131bgcolor="pink"|2583cobweb spidersRedback spider (Latrodectus hasselti)
Theridiosomatidaebgcolor="#FFA"|22bgcolor="#FFA"|150ray spidersTheridiosoma gemmosum
Thomisidaebgcolor="pink"|170bgcolor="pink"|2169crab spidersMisumena vatia (goldenrod crab spider)
Titanoecidaebgcolor="lightgreen"|5bgcolor="lightgreen"|67Goeldia obscura
Toxopidaebgcolor="#FFA"|14bgcolor="lightgreen"|82
Trachelidaebgcolor="#FFA"|29bgcolor="#FFA"|300
Trachycosmidaebgcolor="#FFA"|20bgcolor="#FFA"|148
Trechaleidaebgcolor="#FFA"|17bgcolor="#FFA"|136
Trochanteriidaebgcolor="lightgreen"|6bgcolor="lightgreen"|52
Trogloraptoridaebgcolor="lightblue"|1bgcolor="lightblue"|1Trogloraptor marchingtoni
Udubidaebgcolor="lightgreen"|6bgcolor="lightgreen"|57
Uloboridaebgcolor="#FFA"|19bgcolor="#FFA"|283hackled orb-weaversUloborus walckenaerius
Viridasiidaebgcolor="lightgreen"|3bgcolor="lightgreen"|14
Xenoctenidaebgcolor="lightgreen"|4bgcolor="lightgreen"|33
Zodariidaebgcolor="#FFA"|90bgcolor="pink"|1306Zodarion germanicum
Zoropsidaebgcolor="#FFA"|28bgcolor="#FFA"|186Zoropsis spinimana

= Extinct families =

Notes

{{reflist|group=notes}}

References

= Citations =

{{reflist|refs=

{{Citation |mode=cs1 |last=Coddington |first=Jonathan A. |editor1-last=Ubick |editor1-first=D. |editor2-last=Paquin |editor2-first=P. |editor3-last=Cushing |editor3-first=P.E. |editor4-last=Roth |editor4-first=V. |contribution=Phylogeny and classification of spiders |name-list-style=amp |year=2005 |title=Spiders of North America: an identification manual |publisher=American Arachnological Society |pages=18–24 |contribution-url=https://repository.si.edu/bitstream/handle/10088/4365/CoddingtonSNAPhylogeny05.pdf |access-date=2015-09-24}}

}}