Siphula
{{Short description|Genus of lichens}}
{{Automatic taxobox
| image = Siphula ceratites.jpg
| image_caption = Siphula ceratites
| taxon = Siphula
| authority = Fr. (1831)
| type_species = Siphula ceratites
| type_species_authority = (Wahlenb.) Fr. (1831)
| subdivision_ranks = Species
| subdivision = 22
| synonyms = *Nylanderiella Hue (1914)
}}
Siphula is a genus of lichen-forming fungi in the family Icmadophilaceae. The widespread genus was circumscribed by the Swedish mycologist Elias Fries in 1831.
Description
The genus Siphula includes lichens with a shrubby, upright thallus (the lichen body) that is sparsely branched and has a whitish appearance. The outer surface, or {{lichengloss|cortex}}, is {{lichengloss|pseudoparenchymatous}}, meaning it consists of densely packed cells resembling plant parenchyma (common plant tissue). Siphula species partner with {{lichengloss|chlorococcoid}} algae, a type of green algae that supplies the lichen with energy through photosynthesis.
Inside the thallus, the medulla (inner tissue layer) is compact and has fungal filaments (hyphae) that are generally aligned in a longitudinal direction. No reproductive structures, such as apothecia (spore-producing bodies) or conidiomata (asexual spore-producing bodies), have been observed to occur in this genus.
Chemically, Siphula lichens contain various compounds including p- and m-depsides, dibenzofurans, and chromones, such as the compound siphulin.
Habitat and distribution
Siphula species are found growing on peaty soils and on acid-leached bark in temperate rainforests as well as in wet alpine environments. This genus has a cosmopolitan distribution, occurring across various regions worldwide.
Species
{{Asof|2024|October}}, Species Fungorum accepts 22 species of Siphula:{{Cite web |title=Species Fungorum - Search Page |url=https://www.speciesfungorum.org/Names/Names.asp |access-date=2024-10-25 |website=www.speciesfungorum.org}}
- Siphula australiensis {{au|Kantvilas (2004)}}
- Siphula capensis {{au|Kantvilas (2023)}}
- Siphula ceratites {{au|(Wahlenb.) Fr. (1831)}}
- Siphula chimantensis {{au|V.Marcano (2021)}}
- Siphula coriacea {{au|Taylor ex Nyl. (1860)}}
- Siphula crittendenii {{au|Kantvilas (2023)}}
- Siphula decumbens {{au|Nyl. (1888)}}
- Siphula dissoluta {{au|Nyl. (1888)}}
- Siphula fastigiata {{au|(Nyl.) Nyl. (1863)}}
- Siphula flavovirens {{au|Kantvilas, Zedda & Elix (2003)}}
- Siphula gracilis {{au|Kantvilas (1998)}}
- Siphula paramensis {{au|V.Marcano & L. Castillo (2021)}}
- Siphula parhamii {{au|Kantvilas (2004)}}
- Siphula pteruloides {{au|Nyl. (1859)}}
- Siphula subpteruloides {{au|V.Marcano (2021)}}
- Siphula subsimplex {{au|V.Marcano (2021)}}
- Siphula verrucigera {{au|(J.F.Gmel.) R.Sant. (1966)}}