Stipeae
{{Short description|Tribe of grasses}}
{{Automatic taxobox
|image = Stipa gigantea 3.JPG
|image_caption = Celtica gigantea
|display_parents = 3
|taxon = Stipeae
|authority = Dumort.
|synonyms =
- supertribe Stipodae L. Liu
- subtribe Stipinae Griseb.
- Aciachninae Caro
- Ortachninae Caro
|subdivision_ranks = Genera
|subdivision = 28 genera, see text
}}
The Stipeae are a tribe of grasses within the subfamily Pooidae, with up to 600 described species.Barkworth ME, Arriaga MO, Smith JF, Jacobs SWL, Valdes-Reyna J, Bushman BS (2008) Molecules and Morphology in South American Stipeae (Poaceae). Systematic Botany 33(4), 719-731.
Description
The defining morphological features of the Stipeae include single-flowered spikelets lacking a rachilla extension, and the lemmas (the external bract) have either a sharp point or a terminal awn (long bristle).Cialdella AM, Giussani LM, Aagesen L, Zuloaga FO, Morrone O (2007) A phylogeny of Piptochaetium (Poaceae : Pooideae : Stipeae) and related genera based on a combined analysis including trnL-F, rp116, and morphology. Systematic Botany 32(3), 545-559.
Genera
The tribe includes 32 genera:{{cite journal|last1=Soreng|first1=Robert J.|last2=Peterson|first2=Paul M.|last3=Romaschenko|first3=Konstantin|last4=Davidse|first4=Gerrit|last5=Teisher|first5=Jordan K.|last6=Clark|first6=Lynn G.|last7=Barberá|first7=Patricia|last8=Gillespie|first8=Lynn J.|last9=Zuloaga|first9=Fernando O.|title=A worldwide phylogenetic classification of the Poaceae (Gramineae) II: An update and a comparison of two 2015 classifications|journal=Journal of Systematics and Evolution|volume=55|issue=4|year=2017|pages=259–290|issn=1674-4918|doi=10.1111/jse.12262|doi-access=free|hdl=10261/240149|hdl-access=free}} {{open access}}Peterson PM, Romaschenko K, Soreng RJ, Valdés Reyna J (2019) A key to the North American genera of Stipeae (Poaceae, Pooideae) with descriptions and taxonomic names for species of Eriocoma, Neotrinia, Oloptum, and five new genera: Barkworthia, × Eriosella, Pseudoeriocoma, Ptilagrostiella, and Thorneochloa. PhytoKeys 126: 89-125. https://doi.org/10.3897/phytokeys.126.34096{{cite web |title=Poaceae Barnhart |url=https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:30000032-2 |website=Plants of the World Online |publisher=Royal Botanic Gardens, Kew |access-date=27 October 2024}}
{{columns-list|colwidth=22em|
- Achnatherum {{small|P.Beauv.}} (syn. Aristella)
- Aciachne {{small|Benth.}}
- Amelichloa {{small|Arriaga & Barkworth}}
- Anemanthele {{small|Veldkamp}}
- Austrostipa {{small|S.W.L.Jacobs & J.Everett}}
- Barkworthia {{small|Romasch., P.M.Peterson & Soreng}}
- Celtica {{small|F.M.Vázquez & Barkworth}}
- Eriocoma {{small|Nutt.}}
- × Eriosella {{small|Romasch.}}
- Hesperostipa {{small|(Elias) Barkworth}}
- Jarava {{small|Ruiz & Pav.}}
- Lorenzochloa {{small|Reeder & C.Reeder}}
- Macrochloa {{small|Kunth}}
- Nassella {{small|(Trin.) É.Desv.}}
- Neotrinia {{small|(Tzvelev) M.Nobis, P.D.Gudkova & A.Nowak}}
- Oloptum {{small|Röser & Hamasha}}
- Ortachne {{small|Nees ex Steud.}}
- Orthoraphium {{small|Nees}}
- Oryzopsis {{small|Michx.}}
- Pappostipa {{small|(Speg.) Romasch., P.M.Peterson & Soreng}}
- Patis {{small|Ohwi}}
- Piptatheropsis {{small|Romasch., P.M.Peterson & Soreng}}
- Piptatherum {{small|P.Beauv.}}
- Piptochaetium {{small|J.Presl.}}
- Psammochloa {{small|Htchc.}}
- Pseudoeriocoma {{small|Romasch., P.M.Peterson & Soreng}}
- Ptilagrostiella {{small|Romasch., P.M.Peterson & Soreng}}
- Ptilagrostis {{small|Griseb.}}
- Stipa {{small|L.}}
- Stipellula {{small|Röser & Hamasha}} (syn. Stipella)
- Thorneochloa {{small|Romasch., P.M.Peterson & Soreng}}
- Timouria {{small|Roshev.}}
- Trikerala {{small|Bor}}
}}
Many species initially placed into Stipa have now been split off into new genera. Some recent papers have analysed relationships within and between the genera,{{cite journal|last1=Barber|first1=Janet C.|last2=Hames|first2=Kasey A.|last3=Cialdella|first3=Ana Maria|last4=Giussani|first4=Liliana M.|last5=Morrone|first5=Osvaldo|title=Phylogenetic relationships of Piptochaetium Presl (Poaceae: Stipeae) and related genera reconstructed from nuclear and chloroplast sequence datasets |journal=Taxon|volume=58|issue=2|year=2009|pages=375–380|issn=0040-0262|doi=10.1002/tax.582005}} [https://www.researchgate.net/profile/Liliana_Giussani/publication/233665373_Phylogenetic_relationships_of_Piptochaetium_Presl_Poaceae_Stipeae_and_related_genera_reconstructed_from_nuclear_and_chloroplast_sequence_datasets/links/552e76700cf22d43716e78ae/Phylogenetic-relationships-of-Piptochaetium-Presl-Poaceae-Stipeae-and-related-genera-reconstructed-from-nuclear-and-chloroplast-sequence-datasets.pdf pdf]Jacobs SWL, Bayer R, Everett J, Arriaga MO, Barkworth ME, Sabin-Badereau A, Torres MA, Vazquez FM, Bagnall N (2007) Systematics of the tribe Stipeae (Gramineae) using molecular data. Aliso 23, 349-361.Jacobs SWL, Everett J, Barkworth ME, Hsiao C (2000) Relationships within the Stipoid grasses (Gramineae). In 'Grass Systematics and Evolution.' (Eds SWL Jacobs and J Everett) pp. 75-82. (CSIRO: Melbourne) but a complete analysis has not yet been performed. Stipoid grasses use the C3 photosynthetic pathway and live in temperate areas worldwide.{{cite journal|author=Romaschenko K, Peterson PM, Soreng RJ, Garcia-Jacas N, Futorna O, Susanna A |date=2008| title=Molecular phylogenetic analysis of the American Stipeae (Poaceae) resolves Jarava sensu lato polyphyletic: evidence for a new genus, Pappostipa|journal= Journal of the Botanical Research Institute of Texas|volume= 2|issue=1|pages= 165–192|jstor=41971613}} [https://repository.si.edu/bitstream/handle/10088/6448/Romaschenko_etal_Pappostipa_highres.pdf?sequence=1&isAllowed=y pdf]
Known fossils date from the late Miocene.{{cite journal|last1=Thomasson|first1=Joseph R.|title=Berriochloa Gabeli and Berrichloa Huletti (Graminae: Stipeae), two new grass species from the late miocene Ash Hollow Formationof Nebraska and Kansas|journal=Journal of Paleontology|volume=79|issue=1|year=2005|pages=185–199|issn=0022-3360|doi=10.1666/0022-3360(2005)079<0185:BGABHG>2.0.CO;2|bibcode=2005JPal...79..185T }} [https://www.researchgate.net/profile/Joseph_Thomasson/publication/250070847_Berriochloa_gabeli_and_Berriochloa_huletti_Gramineae_Stipeae_two_new_grass_species_from_the_late_miocene_ash_hollow_formation_of_Nebraska_and_Kansas/links/56324ae608ae584878091c65/Berriochloa-gabeli-and-Berriochloa-huletti-Gramineae-Stipeae-two-new-grass-species-from-the-late-miocene-ash-hollow-formation-of-Nebraska-and-Kansas.pdf pdf]
References
{{Reflist}}
External links
- [https://www.delta-intkey.com/grass/www/stipa-sl.htm Stipa L. sensu lato] (sensu lato refers to the tribe) The grass genera of the world website by L. Watson, T.D. Macfarlane, and M.J. Dallwitz
- {{Commons category-inline|Stipeae}}
- {{Wikispecies-inline|Stipeae}}
{{Taxonbar|from=Q7617501}}