Thorneochloa

{{Short description|Genus of flowering plants}}

{{Speciesbox

| image = Thorneochloa diegoensis 111988936.jpg

| image_caption =

| status = G4

| status_system = TNC

| status_ref = {{Cite web|date=2022-05-30 |url= https://explorer.natureserve.org/Taxon/ELEMENT_GLOBAL.2.160322|access-date=30 May 2022 |website=NatureServe Explorer Achnatherum diegoense|title = NatureServe Explorer - Achnatherum diegoense |publisher=NatureServe}}

| taxon = Thorneochloa diegoensis

| parent_authority = Romasch., P.M.Peterson & Soreng

| authority = (Swallen) Romasch.

| display_parents = 3

| synonyms = * Achnatherum diegoense {{small|(Swallen) Barkworth}}

  • Stipa diegoensis {{small|Swallen}}

| synonyms_ref = [https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:77199108-1 Thorneochloa diegoensis (Swallen) Romasch.] Plants of the World Online. Retrieved 22 May 2023.

}}

Thorneochloa is a genus of flowering plants belonging to the family Poaceae. It contains a single species, Thorneochloa diegoensis, commonly known as San Diego needlegrass.{{cite web |author1=J. Travis Columbus, James P. Smith, Jr. & Douglas H. Goldman |title=Stipa diegoensis |url=https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=45664 |website=Jepson eFlora |publisher=Jepson Flora Project |access-date=25 October 2024}} It is a perennial grass native to southern California and northern Baja California.{{cite web |title=Thorneochloa Romasch., P.M.Peterson & Soreng {{!}} Plants of the World Online {{!}} Kew Science |url=https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:77199107-1 |website=Plants of the World Online |access-date=17 March 2021 |language=en}}

Description

{{see also|Glossary of leaf morphology|Glossary of plant morphology|Glossary of botanical terms}}

Thorneochloa diegoensis is a bunching perennial grass with culms (stems) between {{cvt|70|-|140|cm|in}} tall in an erect or ascending habit. The culms are {{cvt|2|-|4|mm|in}} thick, and characteristically {{cvt|3|-|9|mm|in}} below the lower nodes the internodes have a dense pubescence of retrorse (pointing downward) hairs. The leaf sheath may be glabrous or pubescent, and the collar with a tuft of hairs that measure {{cvt|1.5|-|2|mm|in}} long. The leaf blades are {{cvt|15|-|40|cm|in}} long by {{cvt|1|-|3.5|mm|in}} wide.{{Cite journal |last1=Peterson |first1=Paul M. |last2=Romaschenko |first2=Konstantin |last3=Soreng |first3=Robert J. |last4=Valdés Reyna |first4=Jesus |date=2019-07-16 |title=A key to the North American genera of Stipeae (Poaceae, Pooideae) with descriptions and taxonomic names for species of Eriocoma, Neotrinia, Oloptum, and five new genera: Barkworthia, ×Eriosella, Pseudoeriocoma, Ptilagrostiella, and Thorneochloa |journal=PhytoKeys |issue=126 |pages=89–125 |doi=10.3897/phytokeys.126.34096 |doi-access=free |issn=1314-2003 |pmc=6650443 |pmid=31360096|bibcode=2019PhytK.126...89P }}

The inflorescence, a narrow, densely-flowered panicle, is up to {{cvt|30|cm|in}} long, with ascending, appressed branches. The pedicels are usually shorter than the spikelets. The spikelet is lanceolate with a single fertile floret. The glumes are {{cvt|8|-|11.5|mm|in}} long. The floret and lemma are {{cvt|5.5|-|7.5|mm|in}} long, the lemma evenly hairy and its margins enveloping most of the palea. The lemma is 1.3 to 2 × longer than the palea, and has a long scabrous awn {{cvt|20|-|50|mm|in}} long that is bent twice. Flowering is typically from March to June.

Taxonomy

Thorneochloa diegoensis was first described as Stipa diegoensis by Jason Richard Swallen in 1940 based on specimens collected by Frank F. Gander in Proctor Valley.{{cite journal |last1=Swallen |first1=Jason R. |title=Miscellaneous new American grasses |journal=Journal of the Washington Academy of Sciences |date=1940 |volume=30 |page=212 |url=https://www.biodiversitylibrary.org/page/39700244#page/242 |access-date=5 June 2025 |publisher=Washington Academy of Sciences |location=Washington, D.C.}} It was later transferred into the genus Achnatherum by Mary Barkworth in 1993, along with many grasses mostly segregated from Stipa as part of a new delineation of the North American Stipeae.{{cite journal |last1=Barkworth |first1=Mary E. |title=North American Stipeae (Gramineae): Taxonomic changes and other comments |journal=Phytologia |date=January 1993 |volume=74 |issue=1 |pages=1-25 |url=https://www.biodiversitylibrary.org/page/13068903#page/3 |access-date=5 June 2025}}

A reorganization of the Stipeae tribe based on molecular phylogenetics by Peterson et al. in 2019 resulted in the new monotypic genus Thorneochloa and a new combination for this species, Thorneochloa diegoensis. Thorneochloa is part of a strongly-supported clade with Achnatherum that is characterized by a maize-like epidermal pattern on the lemma. The genus is named in honor of botanist Robert Folger Thorne.

Distribution and habitat

Thorneochloa diegoensis is native to southern California and northern Baja California. It occurs on all of the northern Channel Islands and San Nicolas Island, and on the mainland in San Diego County in the Peninsular Ranges, continuing south across the international border along the coast of Baja California to San Quintín and Isla Todos Santos.{{Cite journal |last1=Rebman |first1=Jon P. |last2=Gibson |first2=Judy |last3=Rich |first3=Karen |date=15 November 2016 |title=Annotated Checklist of the Vascular Plants of Baja California, Mexico |url=http://sdplantatlas.org/pdffiles/BajaChecklist2016.pdf |journal=Proceedings of the San Diego Society of Natural History |publisher=San Diego Natural History Museum |volume=45 |pages=289 |via=San Diego Plant Atlas}} It is a resident of chaparral and coastal sage scrub ecosystems below 350 feet in elevation, especially near streams.[https://web.archive.org/web/20110611141946/http://herbarium.usu.edu/webmanual/info2.asp?name=Achnatherum_diegoense&type=treatment Grass Manual Treatment]

References

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