Urocyclidae

The Urocyclidae are distributed in the Afrotropical realm, including Madagascar, the Comores and the islands of the Gulf of Guinea.{{cite book|last=Herbert|first=Dai|title=Field guide to the land snails and slugs of eastern south Africa|year=2004|publisher=Natal Museum|location=Pietermaritzburg :Natal Museum|isbn=0-620-32415-5|edition=1st|author2=Kilburn, Dick }}

The family Urocyclidae consists of 3 subfamilies:

• subfamily Rhysotininae Schileyko, 2002
• subfamily Sheldoniinae Connolly, 1925 (1912)Connolly (1925). Annals and Magazine of Natural History (9)15: 467. - synonyms: Peltatinae Godwin-Austen, 1912;Godwin-Austen H. H. (1912). "A review of South-African Land-Mollusca belonging to the Family Zonitidae". Annals and Magazine of Natural History (8)9: [https://archive.org/stream/ser8annalsmagazi09londuoft#page/122/mode/2up 122]-139. [https://archive.org/stream/ser8annalsmagazi09londuoft#page/1/mode/1up Plate 1]-7. [https://archive.org/stream/ser8annalsmagazi09londuoft#page/124/mode/2up p. 124]. Trochonanininae Connolly, 1912;Connolly (1912). Annals of the South African Museum 11(3): 101. Trochozonitinae Iredale, 1914;Iredale (1914). Proceedings of the Malacological Society of London 11(2): 122. Ledoulxiinae Pilsbry, 1919;Pilsbry H. A. (1919). Bulletin of the American Museum of Natural History 40: 245. Gymnarionidae Von Mol, 1970;Van Mol J. J. (1970). "Révision des Urocyclidae (Mollusca, Gastropoda, Pulmonata). Anatomie - systématique - zoogéographie". Annales Musée Royal de l’Afrique Centrale, Sciences Zoologiques 180: 1-231. Rhysotinidae Schileyko, 2002; Zonitarionini Schileyko, 2002; Acantharionini Schileyko, 2002
• subfamily Urocyclinae Simroth, 1889
• tribe Urocyclini Simroth, 1889 - synonyms: Atoxonini Schileyko, 2002; Buettneriini Schileyko, 2002
• tribe Dendrolimacini Van Goethem, 1977Van Goethem J. (1977). "Révision systématique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)". Musée Royal de l'Afrique Centrale , Annales, Sciences zoologiques 218: 355 pp., 4 plates.
• tribe Leptichnini Van Goethem, 1977
• tribe Upembellini Van Goethem, 1977

The following genera are recognised in the family Urocyclidae:

; unsorted

• † Koruella Pickford, 2009

;Subfamily Rhysotininae

• Rhysotina Ancey, 1887{{Bouchet 2005}}

;Subfamily Sheldoniinae

• Acantharion Binder & Tillier, 1985
• Africarion Godwin-Austen, 1883
• Amatarion Van Mol, 1970
• Angustivestis Pilsbry, 1919
• Aspidelus Morelet, 1884
• Aspidotomium Degner, 1932
• Belonarion Pilsbry, 1919
• Bloyetia Bourguignat, 1890
• Camerunarion Van Mol, 1970
• Carinazingis van Bruggen & de Winter, 1990
• Centrafricarion Van Mol, 1970
• Chlamydarion Van Mol, 1968
• Colparion Laidlaw, 1938
• Degneria Verdcourt, 1956
• Entagaricus Pilsbry, 1919
• Estria Poirier, 1887
• Falloonella Preston, 1914
• Fuellebornia Verdcourt, 1998
• Granularion Germain, 1912
• Gymnarion Pilsbry, 1919
• †Hamya Bourguignat, 1885
• Kerkophorus Godwin-Austen, 1912
• Lacrimarion Connolly, 1939
• Malagarion Tillier, 1979
• Mesafricarion Pilsbry, 1919
• Microkerkus Godwin-Austen, 1912
• Montanobloyetia Verdcourt, 1961
• Morrumbalia Verdcourt, 1998
• Paragranularion Van Mol, 1970
• Percivalia Preston, 1914
• Plicatonanina Verdcourt, 1961
• Principicochlea D. Holyoak & G. Holyoak, 2020
• Principitrochoidea D. Holyoak & G. Holyoak, 2020
• Ptilototheca Herbert, 2016
• Rhopalogonium Degner, 1932
• Selatodryas Herbert, 2017
• Senegalarion Van Mol, 1970
• Sheldonia Ancey, 1887 - the type genus of the subfamily SheldoniinaeHerbert D. G. (2004). [http://www.iucnredlist.org/search/details.php/44709/all Sheldonia puzeyi]. [http://www.iucnredlist.org 2006 IUCN Red List of Threatened Species.] Downloaded on 7 August 2007.
• Sjostedtina Verdcourt, 1961
• Sylvarion Van Mol, 1970
• Thapsia Albers, 1860
• Thielarion Van Mol, 1970
• Thomithapsia D. Holyoak & G. Holyoak, 2020
• Thomitrochoidea D. Holyoak & G. Holyoak, 2020
• Tresia Van Goethem, 1975
• Trochonanina Mousson, 1869
• Trochozonites Pfeffer, 1883
• Tropidocochlion Verdcourt, 1998
• Verdcourtia Van Mol, 1970
• Verrucarion Van Mol, 1970de Winter A. J. (1987). "Verrucarion demeryi, a new semi-slug from Sierra Leone (Mollusca: Pulmonata: Urocyclidae)". Basteria 51(4-6): 87-93. [http://cat.inist.fr/?aModele=afficheN&cpsidt=7598962 abstract].
• Zingis E. von Martens, 1878
• Zonitarion Pfeffer, 1883

; Subfamily Urocyclinae

Tribe Dendrolimacini

• Dendrolimax Heynemann, 1868 - the type genus of the tribe Dendrolimacini

Tribe Leptichnini

• Leptichnus Simroth, 1896 - the type genus of the tribe Leptichnini

Tribe Upembellini

• Leptichnoides Van Goethem, 1975
• Upembella Van Goethem, 1969 - the type genus of the tribe Upembellini

tribe Urocyclini

• Anisotoxon Van Goethem, 1975
• Atoxon Simroth, 1888
• Atoxonoides Van Goethem, 1973
• Atrichotoxon Simroth, 1910
• Buettneria Simroth, 1910Simroth H. (1910). In: Voeltzkow Reise O.-Afr., Wiss. Ergebn. 2(5): 611.
• Bukobia Simroth, 1896
• Elisolimax Cockerell, 1893
• Emphysetes Verdcourt, 2003
• Microcyclus Simroth, 1896
• Nupnus Van Goethem, 1975
• Pembatoxon Van Goethem, 1975
• Phaneroporus Simroth, 1888
• Polytoxon Simroth, 1897
• Pseudatoxon Van Goethem, 1975
• Tanzalimax Rowson, Paustian & Van Goethem, 2017
• Trichotoxon Simroth, 1888
• Udzungwalimax Rowson, Paustian & Van Goethem, 2017
• Urocyclus Gray, 1864 - the type genus of the family Urocyclidae

The following cladogram shows the phylogenic relationship of this family to other families in the limacoid clade:Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. {{doi|10.1046/j.1365-2699.2000.00403.x}}, [https://www.jstor.org/stable/2656267 JSTOR].

{{clade

|label1=limacoid clade

|1={{clade

|label1=Staffordioidea

|1=Staffordiidae

|2={{clade

|1={{clade

|label1=Dyakioidea

|1=Dyakiidae

|label2=Gastrodontoidea

|2={{clade

|1=Pristilomatidae

|2={{clade

|1=Chronidae

|2={{clade

|1=Euconulidae

|2=Trochomorphidae

}}

|3={{clade

|1=Gastrodontidae

|2=Oxychilidae

}}

}}

}}

}}

|2={{clade

|label1=Parmacelloidea

|1={{clade

|1=Trigonochlamydidae

|2=Parmacellidae

|3=Milacidae

}}

|2={{clade

|1={{clade

|label1=Zonitoidea

|1=Zonitidae

|label2=Helicarionoidea

|2={{clade

|1=Helicarionidae

|2={{clade

|1=Ariophantidae

|2=Urocyclidae

}}

}}

}}

|label2=Limacoidea

|2={{clade

|1=Vitrinidae

|2={{clade

|1=Boettgerillidae

|2={{clade

|1=Limacidae

|2=Agriolimacidae

}}

}}

}}

}}

}}

}}

}}

}}

{{Reflist}}

• {{Commons category-inline|Urocyclidae}}

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